TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan uttG 7tans topidSoc..lapan52(3):ISO-156,June2001 Two Diomeu Walker (LepidopteNroac,tuidae)from new species of Japan, Taiwan and Myanmar HiroshiYosmMoTo Tokyo High School, 39-1,Unoki 2-chome,Ota-ku,Tokyo, 146-O091Japan Abstract Diomea insulana sp. n. from Japan and Taiwan and Diomea .femostic tsap. n. from Myanmar are describe dand illustrate d.Cbrsa Walker, 1858 is sunk as a synonym of Diomea Walker, 1858, and Btoptina petrin aButler, 1879 ,so far placed under Coflsa i,s transferred into PaniUaMoore, 1885. Key words Diomea, Cbrsa, Panitki, Diomea Evida ,Diomea jankowski iC,onsa petrina ,new species, synonym, South East Asia, taxonomy. Diomea rotundota Waiker, 1858 and Cbrsa iZgnicolo Wraalker, 1858 are the type species by monotypy of Diomea Walker, !858 and Cbrsa Walker, I858, respectively. Kobes (1983) described Zigera suvarnadvipae from Sumatra, and Sugi (199 2t)ransferred it into Diomea in a check lis otf the Taiwanese Noctuidae . Sugi' s(199 2t)reatment seems to be correct, because the male genitali oaf suvarnaabipae have simple valvae with also simple harpes similar to those of D. rotunclata. As illustrat heedre ,recently I found that suvamadvipae is a clese relative of Cbrsa iignicolo r(aFi g9.), and I pointed out that Diomea and Cbrsa become synonymous with each other (Yoshimot o19,94), The generi cnames Diomea and Clorsa were publishe din Walker's same wotk at the same date ,and here as aii act ofthe fir srteviser I select Diomea as a valid name of thi sgenus, of which synonymy is summarized as fo11ows. Diomea Walker, 1858 ("lg57 "Li)s,t 5ipecimen tsepi dIlnseets Colin Bn Mus. 13: I079, 1 109. Type species: Diomea rotundata Walker, 18S8 (C`1857 "i)bi,d l13: 1110 ,by monotypy. Cbrsa Walker, 1858 (L`185 7L"is)t ,5Pecime nmspiO hLisect sthllv Bn Mus, 13: 1079 ,1101. Type species: Cbrsa iignicolor aWalker, 1858 ("1857 it'h)id ,13 :1101 ,by monotypy. Syn. nov. Zigera Walker, 1862, J Rroc .Linn. Soc. (Zool 6.: )182 . Type species: Zigera orbijZira Walker, 1862 ,thid 6: 182, by monotypy. Hbteroscot iBaryk, I949, Ark ZoeL 41 (A)1 ]138 . Type species: Hbteroscot isaCMgia Bryk, 1949, thid L41 (A)1 :138, pl. 5, fig .9, by original designation. This 20 inAsia Himalayas(Poole1,989), The genus contains about species east ofthe adults have generall ydull and dark forewin gsspeckled with black dots ,but are so rich in variety of wing pattern that the grouping of species seems to be yery diMcult . Among them the fbllowin gthree species, nj'ankowskii (Oberth U1r8,80) ,D. Ib,id dHampson, 1902 and D. insulana sp. n. have the same patter nofthe fbrewing maculation and the same basic structure of the male genitalia and apparently constitute a natural group within the genus . In the forewing pattern ,they have rather pale gray to brown and plain ground color with a large triangular costal black marking in the middle. This marking is comprised of a black reniform accompanying two costal blac kbar sabove i tand a dark shade between thes ebars, of which the inner one is oblique to reniform. The basal pait of the costa is stained with black and a short black bar is presen tbefbre the costal triangular black marking. The lineisthin black, the isirregularly dark postmedian and serrate, and subtermen shaded with grayis hbrown. Jn the male genitali ath,ese three species share the unique character that the NII-Electronic Library Service TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan Two New Specie sof Diomea from Japan, Taiwan and Myanmar ISI 2 4 5 6 8 9 10 Figs 1-10. Diomea spp. 1. D. jankowski (iOberthU rc)l,, Japan, 2. D, insulan asp. n., cP, holotype ,Japan, Okinawa I. 3, Ditto ,dn, paratype ,Taiwan, Nantou Hsien. 4. D, fivid Haarnpsen, dii ,Nepal. 5. Ditto, clZ, syntype, Simla ,BMNH. 6. D. sp. .ferrosticta n., cn, holotype ,Myanmar, Kachin, 7, D, eupsema Swinhoe, cii, Myanmar, Kachin. 8. Ditto ,6n ,syntype, Perak, BMNH. 9. D. iignicolo rWaalker, g, syntype, BMNH. 10. D. suvarnadvipae (Kobes) d,ii ,paratype ,Sumatra. harpes are asymmetric, the Ief tone being lenger . D. eupsema Swinhoe, 1902 has similar wing markings, but the fbrewing is broader and the male genitali aare diflere nitn having syrmnetric harpes . The early stages of these throe species are as yet unknewn, but the larva may be fungivoro ubse,cause in n cremata Butler ,l878 the larva is well known to feed on fungi of the Polyporaceae in Japan (Sug i1,987). Piomea jankowski(iOberthU(rF)ig1,) Capnocle sJ'ankowskit 'OberthUr ,lg80, Etud Elei t5: 87, pl .9, fig .1. Length of fbrewing 13-16 mm, expanse 26-31 mm. Male genital i(aFig 1.1). Uncus long and slender; tegumen narrow and high;valva with a NII-Electronic Library Service TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan IS2 HiroshiYosHnioTo small knob-lik eswelling near the middle ofthe costa; lef tharpe leng and it stip tapered and the right one gently curved; jux traoundish, widened below m{ddle, Aedeagus long and slender, vesica without cornutus. Material Japan:1dZ, Yamanashi Pref,Uenohara, 250m, 6.viii.I977,H. examined. Yoshjmoto leg. ;1 (1, Kanagawa Pref, Ohdarumi, 350m, 14 .viii. 1977, H. Yoshimoto leg,; l di, Tokyo, Okutama, Nippara, 700 ni ,10. vi, 1978 ,H, Yoshimoto leg. Distribution.Russia(thePrimoryeTerritory),Korea, Japan, Diomea insulana sp. noy. (Fig 2s, 3) Male. Length of fbrewing 13-14mm, expanse 23-27 mm. Very similar to junkowsk ibuit, fbrewing more tinged with gray ,with les serange tint; antemedian line obsolete; a black costal bar aboye antemedian line closer to the costal triangular marking; postmedian line not beyond injankowskii. so excurved cell as Male genitali (aFig I.2) , Uncus a litt ]sehorter than in junkowsk ivial;va with the costal swelling large r;lef tharpe much elongated ahd nearly equal to the tip ofthe valva and right harpe bent ;juxt ahjgher and narrower than in jonkowsk iiA.edeagus as in J'ankowskii. Holotype. Japan,Okinawa Pref,Okinawa I,,Kunigami,Okuma-rindo,26-27. 2000, cl, v, H. Yoshimoto leg , Paratypes ,1 cl, Taiwan, Nantou Hsien, Lushan Spa, 1,2oo m, 29. iv- 1. v. 1984 ,H.Yoshimoto leg. ;I dn, same localit y3,0, vii-1. viii .1984, K,Yazaki leg. Holotype will be deposite din Laboratory of Insec tSystematic sN,ational Institu toef Agro-Environmental SciencesT,sukuba. Distribution.Japan (theRyukyus),Taiwan. DiantealividaHampsen (Fig4s,5) Diomea tivid Htaxmpson, I902 ,Z Bombay nat Htst 14: 216. Length of fbrewing 12-13 mm, expanse 2I-25 mm, Male genitali (aFi g1,3) . Uncus long and stout; valva without costal kneb-lik eswelling; lef tharpe rounded at tip and right harpe spatular; juxt roaundish dorsall ya,nd deeply cleft at bottom, Aedeagus shorter than in the two preceding species. Materia lexamined. 1cl,syntype (Fig, 5S)i,mla, BMNH, not dissect edI.dFn,Nepal, Kathmandu, Godavari, 1,600 m, 1. v. 1990, M. S. Ljmbu leg. ;1 cii, Nepal, Janakpur ,Dolak- ha,Deolari2,8oo 25. 1994,M. S.Limbu leg, m, v-7. vi, Distribution. India,Nepal. Diomea empsema Swinhoe (Fig 7s, 8) Diomea eupsema Swinhoe, 1902, Ann, Mtig nat Htsa (7 )9: 85. Length of forewin g16 mm, expanse 29 mm. Male genitali a(Fi g1.4). Uncus slender, tapered at tip ;tegumen moderate]y high ;valva with it scostal margin a littl ceoncave before middle; harpe short and acute, angled at base; a densely setose ridge below base ofcosta; juxt haorseshoe-shape wdith it sbottom protruding NII-Electronic Library Service TThhee LLeepipdiopdteorpoltoegircaollSoocgleitycal Society oofJfap anJapan Two New Specie sof Diomea from Japan, Taiwdn dnd Myanmdr 153 ww/'l,I a/.mrw / //z atth.ttthtx. ' / /f//diny"tw ,l$il //I . -"l*ew F]ggli-16 Male genital iaof Diomea 1] DJankowsku (Oberth uJra)p,an 12 D insulana sp n, holotype, Japan, Okinawd I 13 D hvido Hdmpson, Nepal 14 D etcgxsema Swinhoe, Myanmar, 15 Dsuvarnadmpae (Kobes ),paratype, Sumatra 16 D forrost gipc tna holotype, Myanmar, Kachin caudally, sa¢cus wrde U-shaped Aedeagus slender Material exdmined 1[1, syntype (Fi g8), Goping, Perak, BMNH, not dissected I di1. Myanmar, Kachin, Putao, Machanbow, 500m, 22-23 vi 1998, K Yazaki leg DistribuUon Malaysia, Myanmar NII-Electronic Library Service TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan 154 Hiroshi YosHiMoTo Diomea fo,vvst sip.c nt. a(Fi g6). cin. Length of fbrewing 11 mm, expanse 21 mm, Forewing dark blackish brown, a pale bluis hgray shade beyond postmedian line ;orbicular represented by an ill-defin beldack bar. Hindwing with inner margin widely sutfused with ferreu sorange, it souter side define dby a line. whitish gray Male genitali (aFi g16,). Uncus long, a litt lsweollen befbre tip, which is pointed ;tegumen narrew; valva with both margins nearly parallel ,roundish at tip ;harpe very short; a small setose knob below harpe; juxt atriangular ;.saocus roundish. Aedeagus thick and short, weakly curved, Holotype. Myanmar, Kachin,Putao,Mt Nwe Zin,750m, 16-20. 1998,K. Yazaki cl, vi. leg . Holotype will be deposite din Laboratory of Insect Systematic sN,ational Institu tofe Agro-EnvironmentalSciencesT,sukuba. Distribution. Myanmar. The fo11owin gtwo are apparently sister species. The ground color ofboth wings is ligh tto dark brown with a reddish to purplis htinge, Forewing has a wide and black basal marking and a wide and black apical band, which is edged insid ewith pale yellowish brown and continues narrowing to tornus, and has two costal black bars. Hindwing has a similar terminal black band widened around apex. Diomea lignicolo r(aWalker c)o,mb. n. (Fig 9.) ConsaiignicoloWraalker,18S8"not 1857",Ltst SpecimenkspidiLnsectsCoiZhBn Mtis. 13: 110L Forewing brown, pale grayish with apex roundish. Materia lexamined, 1 9, syntype (Fi g9,) ,BMNH, not dissected. Distribution. SriLanka, China (Jjangx(iC)hen,1985,1999),Sikkim (Hampson,1895). Diomea (Kobes()Fig1.5) szavai:nadvipae Zigera suvarnadvipae Kobes, 1983 ,HOteroeer asumatn 2: 14 ,fig sIN, O, 8. Very similar to the preceding species, but the wing coloration is deeper reddish brown with a strong purplis htint , In the Sumatran specimens, the fbrewing is acuter at apex than in iignicolova. Materia lexamined. Sumatra: 1 6Z, paratype, Aek Tarum, Gunung Malayu, 9. x. 1983, Diehl leg. ;Nepal: 1 cl, Kathmandu, Godavari, 1,6o om, 22. vii. 1990 ;2 &, Mechi, Ilam, Godok, 400 m, 11-18. vi. 1993, M, S, Limbu leg . Taiwan: 1?, Sun Moon Lake, 2. viii. 1978,T.Niizatoleg. The Nepalese specimens befbre me are not so dark as the Sumatran ones, but enough darker than the preceding species and were determine das suvaznadyipae (se Hearuta, 1994) .But, the wing shape is more roundish as in Sri Lankan fignieolo raA. Taiwanese femal ehas intermediate forewing.The (Fig1.5) similarly coloration and a roundish rnale genitali a of the Nepalese and Sumatran specimens are completely identica lan,d at the present ,I consider that they represent suvamadvipae. But this determinatio nshould be confirrned in future through comparison of the male genital iwaith Sri Lankan specimens of iignicolora, NII-Electronic Library Service TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan Two New Species of Diomea from Japan, Taiwan and Myanmar 155 Distribution.Sumatra (Kobes1,983),India (Wang,1995),Nepal (Haruta1,994),Taiwan (Sugi1,992). Although the type species of Diomea and Cbrsa are congeneric, I think that al1 species now placed in Cbrsa should not be transferred into Diomea automatically. Among the six species of Cbrsa liste dby Poole (1989 B)n,zptin apeima Butler ,l879 has a strong affinity to PantiZ tdzispi l(aWalke r18,65), type species of Panido Moere, 1885 ,in both wing maeulation and the male genitalia ,and I here place it in PaniZla. Ptinilla (Butler), petn'na comb. rey. Bkptina petrin aButler, 1879, IUust mpical SPecimen sLepidL Hkiterocera CbiZ} Bin Mus 3: 64, pl. 56, fig. 3; Matsumura, 1905 ,Can Insec t.iap .1 :111. Panie2 kpetrina: Warren, 1913 ,in Seitz ,imcroiepia lPVid 3: 370 ;Matsumura, 1931 ,6000 nvus tinsects Japan-Enrpfre: 826, no.973 (fig .I)n;oue & Sugi ,1958 ,in Ineue ,Check List Lepid Japan 5: 585. Corsa petrina :Sugi, 1982, Moths Jopan 1: 888, 2: 401, pl.219 ,figs 20,21 ,pl.356 ,fig 5.; Poole, 1989, in Heppner, Lepia Cat (N. S. )118: 276. H)lposado otoensts Marumo, 1920, X Coza Agria 7bkyo lnzp .Uhiv .6: 262, pl.20 ,fig 2.; Inaba, 1933, Kontyfi 6: 208. Cbnsa otoensts: Inoue & Sugi, 1958, in lnoue, eneck List Lepid Jopan 5: 592; Ogata, 1958, Ibones Heterocerorum japonicotu mColon nath [2] :182 ,pl.116, fig 2.436. Acknowledgments In writing this paper I ewed much to Mr M. R. Honey of the Natural History Museum, London, for his kindnes sduring my stay at the museum. My sincere thanks are also due to Prof DrL.W, R, Kobes of Universit yof GOttingen ,G6ttingen ,fbr his kind gif tof type specimen, and to Messrs the lat eT. Haruta, Tokyo, Y. Kishida ,Tokyo, and K. Yazaki, fbr their kind otler of the valuable specimens, References Chen, Y.-X., 1985 , Lepidoptera :Noctuidae (4 ).Economic l)isee tthuna of China 32: xiv, 167 ,15 pls . Sience Press, Beijin. (I nChinese), , 1999 . Lepidopter aNoctuidae . Fkeuna Sinic a(Insect 1a6). Ixxiii ,1,596pp. ,68 pls .Science Press ,Beijing . (In Chinese). Hampson, G. F. ,189S . the Ftiuna of Britts hindia, inckidug Coj'lon and Burma (Moth s3). xxviii, 546 pp. Taylor and Francis, London. Haruta ,T. ,1994 . Noctuidae :Carocalina eand Ophiderinae. in Haruta, T. (Ed. )M,oths ofNepal, part 3. 7inea 14 (SuppL 1): 140-l53, pls 87-91. Kobes, L.W.R., 1983 . Weiter e6 neue Arten aus dem Regenwald (Lep .N,octuidae: Acronictinae, Sarrothripinae, Ophiderinae), Hleterocera sumatr. 2i 9-15. Poole, R.W,, 1989. Noctuidae. in Heppner, J.B.(Ed.)L,epid!Ctit.CN.S.)118:[i]-xi1i-1,314. Brill/Flo r&a Fauna Publ, ,Leiden. Sugi, S., 1987. A summary; general views on irnmature stages and life-histo roif1easrger moths in Japan. Iit Sugi, S, (Ed, )L,arvae of large rMbths in Jopan: 267-301. Kodansha Co. Ltd, Tekyo. , 1992 . Catocalina e.in Heppner, J.B. & H.Inoue (Eds )C,heeklist. Lepid 72iiwa n1 (2): 175-183. Wang, H. Y., l99S. Noctuid moths and it' asllied species from the neighboring counuies. Guidb Book to lnsects in TIiiwan 14: 1-216. Taipei. (In Chinese). Yoshimoto, H., 1994. Notes on Caduca athopunctata (Walke r()Noctuid aOpeh,iderinae )collected at Ishigakial' iI'. m,tahe Ryukyus. Japan Hleterocen' sXt s('l78 )3:3-34 (i nJapanese). NNIII-IE-leEcltreoncitcMrboranryiScer vLiciebrary Service TThhee　 LLeepipdiopdteorpoltoegiroaollogical　 SSoooiceityety 　ooff　 JJaapapnan 156 Hiroshi　YosHIMoTo 摘 要 日本 台湾 ミャ ン マ ー産の DiOmea の 2新種 とオオ トウアツバ の所属 （吉本　浩） ， ， Diomea　Walker 1858 と Corsa　Walker 1858の模式種が同属の可能性の高い こ とは以前に も触れたこ ， ， とがあるが （吉本 1994） 本報で は改めて これ らを互い にシノニ ム と認め，その属名には，同 じ日付 ， ， で公表され た 1）iomea と Corsa の 2つ の内　 Diemea を残す こ ととした．ただ し，現在 Corsaに 置か ， れる種は 模式種の lignieolo　raWalker 以外は1）iomea とは無縁で 別の 属に配置すべ きと考えられ ， ， る．特に，日本のオオ トウアツバ は，その斑紋や♂ 交尾器の形状か らもDiemea とは程遠く，む しろ 斑紋や♂交尾器の類似性などか ら，Panill　a Moore，1885に置 くの が穏当で ある．なお，本報で は Diomea の次の 2新種 も記載 した． 1）imea　insulana　sp ．　n．ミナ ミマ エ ヘ リモ ン クチバ （新称） D．　jankowsk（iObierthUr）マ エ ヘ リモ ン クチバ やイ ン ド，ネパ ール に産する D ．　 livid　Haampson に酷似 するが，♂交尾器が異なる．模式産地は沖縄本島北部の国頭村．台湾に も産する． Diomea　ferrest　ispc，　tna． 後翅内縁が幅広 く橙褐色 を呈する種で，属内に近似の もの はい な い．模式産地は ミャ ンマ ー北部の Kachin県 Putao． （Accepted　April　4，2001） Published　by　the　Lepidopterologic　aSlociet　yof　Japan， 5−20，Motoyokoyama 2，Hachioji，Tokyo，192−0063Japan 　 　 　 　 一 NNI工I工-EElleoetcrotniroonic　 LLiibrbarryary 　Service