TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan uettsk 7lrans,lqpidS.oc.Japan 59 (4)2:85-290, September2008 TWo new species and one new subspecies of Pyrginae (Lepidopt eHreasp,eriidae) fromSumatra Laos and Kiyoshi MARuyAMAi) and Jiro UEHARAi' `'9-7-106 Minami6sawa 5 chome, HachiQji-shi T,okyo, 192-0364 Japan; e-mail: hesperi@jn2,go-net,nc.jp 2'326, Iwai-cho, Hodogaya-ku, Yokohama-shi, Kanagawa, 240-O02 3Japan ;e-rmail:[email protected],jp Abstract Two new species and one new subspecies of the Pyrginae are described and illustrated: Capil asumatrana sp. nov, from Sumatra, Pintara prasobsuki tao subsp, nov, from Laos, and Cotadenia koiivaya isp. nov. from Laos. One taxonomic status is changed: Pintara prasobsttki (EK-Amnuay ,2006) ,sLat .nov,, which was originally describe das a subspecies of Capil apat{- rrpunetata. Key words Taxonomy, Capil asttmatrana sp. nov., Pintara parasobsuki lao sp. nov., Co]adenia koiwayai sp, noy., Pintara prasobsuki (EK-Arnnuay ,2006) ,stat. nov., Hesperiidae ,Sumatra, Laos. CZipil saumatrana Maruyama & Uehara, sp. noy (Fig 1s-4, 35-37) Description . g' .Palpus: second segment yellow; third segment thick and short. Antenna: lengt h12,8 mm; shaft black; club arcuate, black ;apiculus black .Forewing :length 31,O mm, upper side white, marginal and apical area black ,with black spots in spaces 2, 3 and in cell, cilia black ;underside same as upper side. Hindwing: lengt h23.3 mm, upper side white, marginal and tornal area black ,with black spots in spaces lb to 7, cilia black ;under- side same as upper side. Male genitali aU,ncus tip narrow. Valva diyided int othree parts ,the upper one (haip ep)ro- jecti ntghe, central one bent inwards, and the lower one (cuil lberro)ad and bent upwards. Aedeagus slender, 4. Palpus: second segment yellow; third segment thick and short, Antenna: lengt h12.9 mm; shaft black ;club arcuate, black ;apiculus black .Forewing :lengt h32.8 mm, upper side white, marginal and apical area black, with black spots in spaces lb, 2, 3 and in cell, spots in spaces 2, 3 and in cell Iarge c,ilia black ;underside same as upper side. Hindwing: length 25.6 mm, upper side white, marginal and torna larea black ,with black spots in spaces 1b to 7 and in cell, cilia black ;underside same as upper side. Materia lexamined. Holotype 8, Mt Dempo, E, Sumatra ,May 1997, now in K, Maruyama collection (Tbkyo )wi,ll be deposited in The Research Institu tofe Evolutionar yBielogy ([IbkyoP)a.ratypes.1r,Mt Dempo, E.Sumatra,April1985,inJ.Uehara collection (Kanagawa1)\;,SilarahiK,aloHills,Sumatra Utara, 17July1997,RickEffendyleg,,in K,Maruyama (Tokyo). collection Distribution.Sumatra. Remarks. The new species is similar to Capila pieridoid e(sFig 5s-7, 12-14) but differs fror nit in the wing pattern and structure of male genitali aM.ale forewin gof all subspecies belongin gto C. pieridoide hsave no black spots in spaces or cell, but underside of hindwing has a black spot in cell. Female forewin gand hindwing of all the known subspecies of C. pieridoides are brown. In the male genitali aup,per ponion of valya of new species is longe rthan in C.pieridoide san,d middle ponion of valva of new species is shorter than in NII-Electronic Library Service TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society oofJfap anJapan 2S6 Kiyoshi MARuyAMA and Jir oUEHARA (ldkhliage/'X"E,3 {1 xid,grv wa' 2v '"tw ge･.,･ .q N '" "" gf' ''su ?"'i t.'･!i.,･ge5' 8,･ x f 11 , 12 "f 1 14 tttt ' ･: . ･.s, ./ ,･t･･･ ･f ./ma 15 16 17 ･18 Figs 1-4. Capila s"matrana Maruyama & Uehara, sp. nov, 1. Hulotype di "u,ppersiclc. 2. Ditto, underside. 3, Paratyp eg, upperside. 4. Ditto ,underside. Figs 5-g. Copilapieri.doide ssojbe Evans, 1934, Malay Pen. 5. .pt. upperside. 6. g' ,underside. 7. 4,uppersid e8,. 4,underside, Figs 9-12. Pintara pvasobsuki lae Maruyama & Uehara, subsp. no-,. 9. Holotyp e5" ,uppcrsicle. 10. Ditto ,undersi(le, 11 .Paratype 9, upperside. 12. Ditto ,underside. Figs 13 ,14. Pintara prasobsuki prasobsuki (EK-Amnuay 2,006), N, Thailand, holotype (EK- Amnuay, 2006) g' .13. Upperside .14 .Undersidc. Figs 15-18. Capita pattrip"neatata tamdaoensis Dcvyatin, 1996 ,Tam Dao, N. Vietnam, 15. X, upperside. 16. g,undersid e1.7. 9,uppersid e1.8. g,underside. pieridoides, Pintaraprasobsuk ilao Maruyama & Uehara ,subsp. nov. (Fig s9-12, 19-22 ,4143) Descriptio n9.. Palpus :second segment slightly porrec twith white and black scales below, black above; thir dsegment thick and long .Antenna: lengt h13.2 mm; shaft btack above, whitish brown below; club arcuate, cream-colored below, black above; apiculus sharply NII-Electronic Library Service TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan 287 X,･in n A tettt ti l., tL e- Figs 19-22. Pa]pi of Pintara prasohsttk ilao Maruyama & Uehara ,subsp. nov. 19. Holotype 8', upperside. 20, Ditto ,underside, 21. Paratype 9,upperside2.2. Ditto u.ndcrside, Figs 23-26 . Palp iof Capii apauripuneatata tamdaoensis Devyatin, 1996, Tam Dao, N, Vietnam. 23. e',uppersi d2e4., r,underside. 25. ?,uppersid e2.6, ll, underside. pointed ,tip reddish brown below .Forewing :length 25.3 mm, with costal fold ,upper side brown with white hyalin espots in spaces lb to 8 and two in cell, with white hyaline streaks in spaces 2, 3, 4, 10, 11 and in cell, cilia brown; underside same as upper side. Hindwing: length 19.3 mm, upper side brown with black spots in spaces lb to 6 (als 7o in paratype) and in cell, cilia brown; underside same as upper side. 9. Palpus :second segment slightly porrec twith white and black scalesr below, black above; third segment thick and long .Antenna: length 13.5 mm; shaft black above, whitish brown below; club arcuate, cream-colored below, black above; apiculus sharply pointed ,tip reddish brown below. Forewing :lengt h26 mm, upper side brown with white hyalin espots in spaces 1b to 8 and in cell, with no white hyalin estreaks in spaces and in cell, cilia brewn; underside same as upper side. Hindwing: lengt h21 mm, upper side brown with black spots in spaces lb to 7 and in cell, cilia brewn; underside same as upper side, Ma]e genital i(aFig s41-43) ,Uncus same size as tegumen and curved, Gnathos short. Cuiller asymmetric, broad ,tip projectin gH.arpe simple and long. Aedeagus slender and curved, '/'"sc'""f'el'･wc. ･as ･27' Siiit/.., n -( . .)twt. 1 '' 31 32 33 34 Figs 27-30 .Coladeni akoiwa.v aMiaruyama & Uehara, sp. nov. 27. Holotype X, upperside. 28. Ditto ,underside. 29, PaTatyp e4, upperside. 30, Ditto ,underside. Figs 31, 32. Coladenia vitrea Leech, 1894, Siao-Lou (BMNH), o", 31, Upperside ,32, Underside. Figs 33, 34. Coladeni asheila Evans, 1939, Fukien (BMNH), o". 33. Upperside. 34. Underside. NII-Electronic Library Service TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan 288 Kiyoshi MARuyAMA and jiro UEHARA Material examined, Holotype 8, Nakhai to Laksao 20 km, C. Laos, alt 5.48 m, N17052.05.4, EI05002.32.9,4 May 2006,K, Maruyama leg,, inK.Maruyama (Tbkyo), now collection will be deposite din The Research Institut eof Evolutionar Byiology (rlbky oP)a.ratypes . 1 e'O,udomxay, N,Laos,2,May 1995,J.Uehara leg,;18'X,amneua, N. Laos,9,May 2004; 2 ?,Xamneua,N, Laos,May 2004, 8,May 2006,inJ.Uehara (Kanagawa). collection Distribution.Laos. Remarks. The new subspecies differ sfrom the nominotypical subspecies, P. prasobsuki (EK-Amnua y2,006), stat. nov., which was describe das (;ltpi lpaauripttneta ptvaasohsuki from N. Thailand ,as fo11ows .Discal spots in forewing cell are two but in the nominotypi- cal subspecies one. This species is quit esimilar to Capilapa"ripttnet aitn aappearance, but differ sfrom it as follows .The second segment of the palpus is slightly porrec tand thc third one long in P, prasohsuki; this character shows P. prasohsuki to be in the 7ingiades group of Evans C1949 )bu,t the second segment of the p2dpus is erect and the third one short in C. pauripunetata. This character shows C. pauripunetata to belong in the Cetaenorrhinus group of Evans (1949 )T.he underside of the second segment of the palpus has white and black scales in P, prasobsuk ibut pale yellow in C. pauripunetata. The male genital iofa the two species are quit edi fferent. Coladenia koiwayai Maruyama & Uehara, sp. nov. (Fjg 2s7-30, 47-49) Descriptio n.r, Palpus :second segment porrec tw,ith white scales below, with white and black scales above; third segment thic kand long .Antenna: lengt h7.8 mm; shaft black above, checked white and black below; club arcuate, black with white scales below ,black above; apiculus sharply pointed, black below and above. Forewjng: lengt h20.0 mm, upper side black with white hyalin espots in spaces 1b to 8 (some paratypes lack spots in spaces 4 and 5), 10 and in cell, disca land subapical spots coajoined, cilia black ;underside same as upper side but spaces la and lb with white scales. Hindwing: lengt h15.4 mm, upper side bagal area black ,marginal and tornal area white, disca ]area with hyalin espots, these spots coajoincd, cilia white; underside white, Male genital i(aFig 4s7-49) .Uncus shorter than tegumen in size. Cuille rbread ,tip round. Harpe long ,tip as spoon. Aedeagus slcnder and long. ?. Same as male but 1arge :Antcnna: lengt h9,6 mm. Forewing :length 22,8 mm. Hindwing: lengt h16.4 mm. Material examined. Holotype 8, Phu Pan,near Xamneua, N, Laos, alt. 1,858 m, N20"11.33.8,E104eOl.03.32,3March 2001, inK.Maruyama (rlbkyo), now collection will be deposite din The Research Institut eof Evolutionar Byiology (rlbky oP)ar,atypes , 3 e", same locali tays holotype ,25, 29 and 30 March 2001; 7 \, same localit yas holotype, 22, 23, 25, 30, 30, 30 and 30 March 2001 in K. Maruyama collectjon; 14 8, same locali tays holotypc ,14 March 1999, 2, 2 and 3 March 2001, 20 April 2002, 15 March 2004, 17, 21, 24 and 24 March, 18 and 22 April 2006, 13 and 19 March 2007; 1O \, same locali tays holo- type, 27 March, 7, 9 and 13 April 2001, 14 March and March 2002, 22 and 28 March 2005, 26,March, 26,April2006,inJ.Uehara (Kanagawa), collection DistributioLna.os. Remarks. The new species is similar to Coladenia vitrea Leech, 1894 (Fig 3s1, 32) and Cotadenia sheila Evans, 1939 (Fig s33, 34), but palpus second segment color is different. C. vitrea: gray ;C, sheila: brigh torange; C. koiwayai: whitish and hindwing costal area of C.koiwayaiis white. NII-Electronic Library Service TThheLee piLdeoppteirdoloopgitcaelrSooclieotygical Society ooff JJaapapnan 2S9 '' .pt.}e l m =a/wk.hgeva 37 ee,･be pt ."wwn ''x' it w as" .l K.pa tt- 38 39 40 ' /. a ntt.tt . "st s'iva e e./ltak} '.11i, 42 43 pt'kl:/l; ta./t/ wh. ,ll//･as, 'ecge{p l.v/ :'.･ va,,. ' . , - "" ' 45 ,s.iZi'si 46 pm 1/ Figs 35-37 .Male genita]i aof Capila sttmatrana Maruyama & Uehara, sp. nov,, holutyp e.35. Ddorsal view. 36. Lateral view, with lcf tvalva removed. 37. Innc rvicw of lef tvalva. Figs 3840. Male genitali aof Capil apieridoide ssojLt Evans, 1934, Ma]ay Pen. 38. Dorsal view. 39, Lateral view, with lef tvalva removed, 40. Iinner view of lef tvalva, Figs 41-43 .Male genital oifa Pintara prasobsuki lao Maruyarpa & Uehara, subsp, nov., holotype, 41. Dorsal view 42. Latera vlicw, with lef tvalva removed. 43. Inne rview of lef tvalva. Figs 4446. Male gcnitali aof Ct{pi lpaattripuneatat atamdaoensis Devyatin, 1996, Tam Dao, N, Vietnttm .44. DorsaL view. 45. Lateral view, with lef vtalva removed, 46. Inner view ofleft va]va. Figs 47-49. Malc gcnitali aof Coladeni akoiwava tMaruyama & Uchara, sp. nov., holotype. 47. Dorsa lvi'e", .48. Latera lview, with lcf vtalva removed. 49. Inne",iew of lef tvalva. NII-Electronic Library Service TThhee　 LLeepipdiopdteorpoltoegiroaollogical　 SSoooiceityety 　ooff　 JJaapapnan 290 Kiyoshi　MARuYAMA 　and 　Jir　oUEHARA Acknowledgments Maruyama　expresses　his　deepes　tgratitud　eto　Mr 　Arbaimun，　Mr　Rick　Effendy，　Mr　Satoshi Koiwaya 　and 　Mr　Hiroyuki　Wakahara．　Mr 　Arbaimun　and 　Mr　Rick　Effendy　assisted　with collecting 　butterfli　eisn　Sumatra　and　supplied　many 　materials 　from　lndonesi，　aMr　Koiwaya brought　the 且rst　materials 　of　Coladenia　koiwayai　to　Maruyama 　and 　gave　him　a　chance 　to study　them．　Mr　Wakahara 　assisted 　Maruyama　to　visit　Phu　Pan　and 　supplied 　the　materials 　of Coladenia　koiwayai．　Maruyama 　is　greatly　indebted　to　Mr　R　R，　Ackery（BMNH ）and 　Mr Campbell　Smith（BMNH ）for　the　opportunity 　to　examine 　the　hesperii　mdaterials 　in　BMNH inl990． Also，　Uehara　expresses 　his　deepes　tgratitud　eto　Mr　Shilo　Osada　and 　Mr　Khamboune ．　 Mr Osada　gave　the　specimen 　of　Capila　sttmatrana 　to　Uehara　and 　Mr　Khamboune　assisted 　with collecting 　butterfl　iien　sLaos　and　offered　many 　materials 　from　Laos　to　Uehara． References E　va　ns（〈，　JWat．　uHra．　，tH1i9s4tθ9r，　yA）：　C1−厩2，α8’o4ge−t8e8　，qプ11th2e−　1H1e8s，pei’iid‘le　fro丹1　Ett广oρe，　Asia　and 　A“stralia 　in　the　Britis　hMuseum 　　　　　inl9，13903−91．93N6e：w16　s3p，ecies　and 　subspecies 　of　Hesperiidae（Lepidopte）roabtained 　by　Herr　H ．　H6ne　in　China ELkee−cAhkJlnn．　uHa，y，，1　8P9．，32−0108694、．8　B厩u昭tt酬e（εf65li（）fe　Tshfar　iClohaminnda：7，　4J6ap，　paln．　a3nd42　．Co厂ea ：568−569，　pl．41． Moore，　FJ878 ．　Dcscription　of　new 　Asiatic　Hesperiidac：687，　p1．45． 摘 要 スマ トラ及びラオスか ら発見されたチャマ ダラセセ リ亜科の 2 新種及び 1新亜種 （丸山　清 ・ 上原二郎） スマ トラからオバ ケセセ リ属 〔Capila）の新種を記載した．大陸からマ レー半島 ボルネオ島に広く分布 ， する♂の斑紋が白い ピエ リ ドイデス オバ ケセセリ （Capil　apieridoides）の 各亜種とは，♂につ い ては 前 翅に黒紋があること，後翅 （裏面）の中室に黒紋がない こ と，バ ルバ の形状が異なるこ とにより区別，が で き，♀につ い て1よ ピエ リドイデス オバ ケセセ リ各亜種の既知の斑紋が褐色であるに対し，新種は白 色で あるこ とに より区別で きる、 ラオスか らプラソブス ク トラガセセ リ（Pintar　parasobst｛ki）の新亜種P．　t）rasobsuki 　laoを記載 した．北 タ イから記載された原名亜種（J’intara　prasobsuki（Ek−Anmuay，2006））とは，前翅中室の透明班が新亜種は 2個であるのに対 し，原名亜種は1個であるこ とにより区別で きる．なお．プラソブス ク トラガセセリは， 当初，パ ウリプネタタオバケセセ リ（Capil　apauri．p“netata ）の亜種として記載された もので ある．また プ ， ラソブス ク トラガセセ リの外観はパ ウリプネタタオバ ケセ セ リ（Capila　pauripunetata）と酷似する，し か し，プラソブス ク 1・ラガセセ リは下脣鬚第2節が前面に突出し第3節が長い こ とに より，下脣鬚第2 節が直立 し第3節が短いパ ウリプネタタオバ ケセセ リと区別で きる．また，下脣鬚第2節下部の色彩 は，プラソブ スク トラガセセ リが白っ ぽい の に対 し，パ ウリプネタタオバ ケセセ リは薄黄色となる．さ らに，♂交尾器は著しく異なる． ラオス からクロ テ ンセセ リ属 （Coladenia）の新種を記載した．　P1．1国西南部か ら記載 されたビ トレアクロ テンセセ リ及びシエ イラクロ テ ンセセ リに似 るが，ビ トレアクロ テ ンセセ リのが灰色，シエ イラクロ テ ンセセ リの下脣鬚第2節下部がオレ ンジ色であるの に対 し，新種は白色を帯びる点が異なる．ま鵡 新 種の後翅外縁部は白色 となる点によ り区別で きる． （Accepte　dMarch 　17，2008） Published　by　Ihe　Lepidopterologicu　lSociet　yof　Japan， 5−20，　Motoyokeyama　2，　Hachioji，　Tokyo，192−0063　Japan 一 NNI工I工-EElleoetcrotniroonic　 LLiibrbarryary 　Service