THREENEWSPECIES OF TIPORUS WATTS (COLEOPTERA: DYTISCIDAE) WITH REDESCRIPTIONS OFTHE OTHERSPECIES IN THEGENUS WATTS C. H. S. WATTS, C. H. S. 2000. Three new species of Tiporus (Coleoptera; Dytiscidae) with redescriptions of the other species in the genus. Records of the South Australian Museum 33(2): 89-99. Three new species of Tiporus Watts, 1985 (Coleoptera: Dytiscidae) are described and figured: Tiporus georginae, T. lachlani and T. moriartyensis. The original descriptions ofthe existing species were based on a small number ofspecimens. Recent collections have allowed these descriptions to be reviewed and, where necessary, corrected and/or enlarged. A key is giventothe 11 knownspecies. C. H. S. Watts, South Australian Museum, North Terrace, Adelaide, South Australia 5000. Manuscriptreceived 11 October 1999. QM The genus Hypodes Watts, 1978, later changed Queensland Museum, Brisbane due to preoccupation to Tiporus Watts, 1985, was SAMA SouthAustralian Museum, Adelaide erected for a group of small to moderate sized WAM WesternAustralian Museum, Perth Australian Hydroporini related to Antiporus QDPIM Queensland Department of Primary Sharp. Members of Tiporus are distributed across Industries, Mareeba coastal Northern Australia where they appear to be restricted to rivers and streams, or to the small pools which form in their beds in the dry season. Systematics In these habitats some species, such as T. undecimmaculatus and T. josepheni, are often Thegenus TiporusWattsbelongs tothatsection abundant. ofAustralianHydroporini with pseudotetramerous The genus has not been reviewed since my protarsi and densely rugose/punctate bodies 1978 paper. Many more specimens are now including the legs. It can be distinguished from available, among them the three new species the other genera in this section, Antiporus Sharp described below. The taxonomy of the genus is andSekaliporusWatts, bythe formofthe humeral based on the males, which have distinctive, angle of the elytron and the three-segmented species-specific genitalia and secondary sexual protarsi in the male (see Watts 1997 for more characters on the legs. Females are more difficult details). taxonomically but the additional material has Within the genus there are two morphological allowed these to be tentatively keyed for the first groupings, one group characterised by: lack of time. Unfortunately, in the case of T.josepheni, I pronotal ridges/grooves, moderate to strong mis-associated the sexes in my original dorsal colour pattern, a tendency for the edge of description, which has led to confusion over the the elytron to bulge or extend outwards near the identification ofthis species. The type material of tip, protibia of the male with a small tubercle all species except T. collaris and T. on the outside near the base, and the other undecimmaculatus hasbeen restudied. group characterisedby: pronotal ridges/grooves, The collections from which specimens were dark colouring with only vague reddish patches, examined are listed under the following edges of the elytron serrated but not extended abbreviations: and the male protibia having a broad spine in ANIC Australian National Insect Collection, the middle or towards the apex on the outside. Canberra To what degree these groupings reflect NTM Northern Territory Museum and Art phylogenetic relationships has yet to be Gallery, Darwin determined. 90 C. H. S. WATTS Keyto TiporusWatts Extreme front of elytron only slightly — narrower than adjacent pronotum, 1 Disc of pronotum with a ridge/groove difference <= width of mid segments of running backwards parallel to and at a antenna; edge of elytron with moderate littledistancefromsides; edgeofelytron bulge near tip; males with either short serrated in apical half, neverwith lateral stout proclaws or with moderately long extensionneartip;sidesofpronotumdark, butthin proclaws (Figs 13, 14) 6 atmostdiffusely lighterincolour 7 — 6 Pronotumweaklyridgedatsides;edgeof Disc of pronotum smooth, at most with elytra straight immediately prior to it weak ridges; edges of elytron usually meeting pronotum; male proclaw short, smoothbutoftenwithroundedortriangular thickenedbasally (Fig. 13 ) extension near tip; sides of pronotum T. denticularis(Watts) usually with well-defined light coloured Pronotum smooth; edgeofelytraslightly area 2 curved immediately prior to it meeting — 2 Female with edge of elytron near apex pronotum; male proclaw elongate, thin with large, triangularextension reaching (Fig. 14) T. centralis (Watts) beyondtipofelytron(Fig.8);malewithout 7 Large (> 5.0 mm long); male proclaw suchanextension,protibiawithtriangular thick (Fig. 2), protibia stout, with basal spineonoutside nearmiddle spine (Fig. 3); apical segmentofprotarsi — T.josepheni (Watts) large,deeplybifid(Fig. 3);Pilbararegion Both sexes with elytron with at most a ofW.A T. lachlani sp. nov. modest bulge near tip or with the tip Smaller (< 5.1 mm long); males not as extended; males with small tubercle on above 8 outsideofprotibia atbase 3 — 8 Tipofelytronwithsmallbutdistinctpoint mm 3 Larger (4.5 long); postcoxal lines (except in some T. alastairi); elytral narrow, reaching metathorax; outer serrations usually strong; male protibia metatibialspineinmalegreatlyelongated, withprominent,triangularenlargementin about twice the length of the inner (Fig. apical half 9 12); (femaleunknown) Tip ofelytron rounded; elytral serrations — T. moriartyensis sp. nov. weak; male protibia with small spine/ mm Smaller(<4.1 long); postcoxal lines tubercle oninsidecloseto base wide,oftenpoorlydefinedtowardsfront; T. collaris(Hope) outermetatibialspineinmalenormal,not greatly differentin lengthto inner 4 9 Viewedlaterally,theedgeofelytracurves — forwardforshortdistancepriortomeeting 4 Femalewithtipofelytronclearlyextended pronotum; male protibia with large (Fig. 15);edgeofpronotumweaklysinuate triangular expansion on front edge near infront;malewithmetatibiaexpandedon apex; male proclaw squat insidenearapex T. alastairi(Watts) — T. tambreyi(Watts) Viewedlaterally,theedgeofelytranot,or Tipofelytronnot,oronlyweakly,extended onlyslightly,curvedimmediatelypriorto (edge of elytron may be moderately meeting lateral edge of pronotum; male expanded near, but not at, tip); sides of protibia with small triangular expansion pronotum with only a hint of sinuation; on frontedge in middleortowards apex; male withnormal metatibia 5 maleproclaw thin 10 — 5 Extreme front of elytron considerably 10 Malewithtipofcentral lobeofaedeagus narrower than adjacent pronotum, pointed(Fig. 20);protibiawithtriangular difference > width of mid segments of expansionjustbeyondcentre antenna(Fig.4):lateralextensiontoelytron T. giuliani (Watts) neartipabsentorweak(Fig.4);malewith Malewith tipofcentral lobeofaedeagus verylongdagger-shapedproclaw(Fig.7) broad,weaklytridentate(Fig. 16);protibia T. georginae sp. nov. with triangular expansion on front edge, SPECIES OFTIPORUS 91 eitherinmiddle(N.Qld.)ortowardsapex ventrally, apical segment cylindrical. Mesotarsi (N.T..WA) much narrower. T. undecimmaculatus (Clark) Remarks Resembling T. centralis and T. denticularis in Tiporusgeorginae sp. nov. size and colour but with the mis-match of the humeral angleofthe elytron andthe posterolateral Types angle of the pronotum more noticeable (Fig. 4). Holotype: male: 'W. Aust. Mitchell Plateau. Some specimens of T. centralis have this feature 14°40'S 125°44'E 23 Sept 1982. B. V. Timms', to a degree but the extremely long, dagger-like dissected and mountedon card, SAMA. male proclaw in T. georginae (Figs 6, 7) readily WA CruPsahreart'ypCesA:L5M, "Si1t4e°592/1'S4 1k2m5°S50b'yE W Min'iTnhge sTeippaorruast.esThmealaeedsepaegcuismiesndsisftirnocmtivtehi(sFiga.nd5).otThheer Camp Mitchell Plateau 2-6 June 1988 I. D. species is known only from the Kimberley region Naumann", ANIC; 4, "14°25'S 126°40'E CALM whereit isbroadly sympatric with T. centralis. Site 4/3 14 km S by E Kalumburu MissionW. A. 3-6 June 1988 T. A. Weir", ANIC (3), SAMA Distribution (1). All labelled as collected at light in open The Kimberley regionofWesternAustralia. CALM forest except for one in closed forest at site4/3. Tiporus moriartyensis sp. nov. DescriptionFigs4-7 Length 3.5 - 4.0 mm. Elongate-oval, wider in Types middle, front edge of elytron considerably Holotype: male, 'Moriarty ck N.T. Reardon 8/ narrower than rear edge of pronotum (Fig. 4). 82', SAMA. Pronotum smooth, without lateral ridges or grooves; edgeofelytronturns forwardsharplyjust DescriptionFigs 9-12 before meeting edge of pronotum, very weakly Length 4.5 mm. Elongate-oval, widest in serrate towards rear, usually slightly expanded middle. Pronotum narrower than elytra. Dorsal near but not at tip, quite strongly acuminate. surface dark brown-black, rear of head, lateral Whole body strongly rugose/punctate. Dorsal margins and rear margin of pronotum, three surface dark testaceous-black, front margin of small lateral spots and two larger spots near head, sides and parts of front and rear margins of suture on elytron testaceous. Ventral surface pronotum, three lateral spots andoften some spots mottled dark and light testaceous, appendages inwards from these on elytron, testaceous. Ventral including pro- and mesocoxae much lighter. surface dark testaceous-black, appendages Whole body covered with rugose punctures, including pro- and mesocoxae, much lighter. punctures smaller and less rugose than in other Pronotal process narrow, keeled, bluntly pointed. Tiporus species. Disc of pronotum smooth, Metacoxal lines relatively close in hind half, without lateral ridges or grooves. Side ofelytron widening to a bit more than two times their curves forward immediately before meeting edge narrowest width in frontquarter. of pronotum, very weakly serrate towards tip, Male. Protarsi three-segmented, with adhesive apex obliquely truncated. Pronotal process setae ventrally, anterior parts of segments relatively broad, keeled, tip blunt. Metacoxal expanded, small spine on hind apical corner of lines close, diverging to about two times their second segment, third segment about 2.5 times minimum width towards front, reaching length of second, single claw extremely long, mesosternum. expanded in middle (Fig. 6). Protibia with small Male. Protarsi three segmented, segments tubercle on anterior edge at base (Fig. 7). expanded in front, with dense adhesive setae Mesotarsi as for female. Metatibia with a slight ventrally, second segment with small tooth on thickening on inside near apex. Apical ventrite hind margin, apical segment 2.5 times length of with small, sharp, apical keel/spine. Median lobe second; claw short, rapidly narrowing to about ofaedeagus broad incentre, very narrow in apical half its width in middle (Fig. 10). Protibia with quarter(Fig. 5). weak tubercle/spine near base on anterior edge. Female. Protarsi five-segmented, first three Metatibia with inner apical spine, twisted, moderately expanded with adhesive setae flattened and broadened slightly at tip, about two WATTS 92 C. H. S. 8 (] 10 11 15 FIGURES 1-15. 1-3,dorsal viewofmedianlobeofaedeagus, apical view ofproclaw andanteriorviewofprotibia and tarsi ofT. lachlani; 5-7, ditto T. georginae; 9-11, ditto T. moriartyensis; 4, dorsal outline ofT. georginae; 8, dorsal outline ofelytron of T.josepheni; 12, metafemur of T. moriartyensis; 13, apical view ofproclaw and third tarsal segmentofT. denticulatus; 14,ditto T. centralis; 15,dorsaloutlineofelytronofT. tambreyi. SPECIES OFTIPORUS 93 times length of outer, which is simple (Fig. 12). squat, broad but narrowing to sharp point (Figs 2, Apex oflast abdominal ventrite broadly depressed 3). Protibia relatively short, thick, with strong in middle, rear edge with two small spines in tubercle on outside atbase (Fig. 3). Apical ventrite middle. Median lobe ofaedeagus broad inmiddle, with short well-marked keel near apex. Median abruptly narrowingnearapex (Fig. 9). lobe of aedeagus relatively narrow, parallel-sided Female. Not known. (Fig. 1), ending in pick-like beak (not visible dorsally). Remarks Female. Protarsi five-segmented, first three Although only one specimen is available this is segments weakly expanded anteriorly, densely enough to show that T. moriartyensis is a very covered with adhesive setae ventrally, fourth distinctive species. It belongs to the group of segment very small, apical segment thin, species lacking lateral pronotal ridges but is cylindrical, shorterthan third. Innerapicaledgeof noticeably larger and less strongly punctured than metatarsal segments extended, extension other members. No other Tiporus has the inner accentuatedby anumberofstrong spines. apical spine on the metatibia modified as in this species. The female is unknown but it is likely Remarks thatthischaracteris restrictedto the male. Tiporus lachlani differs from other Tiporus with ridged/grooved pronotum by its large size Distribution — and strongly narrowedpronotal-elytraljunction, as Known only from the type locality Moriarty wellas in malecharacters. Creek, 160°4'S 129°12'E—in the Northern Territory. Distribution The Pilbara/Gascoyne region of Western Australia. Tiporus lachlani sp. nov. Types Redescriptions OfTheOther SpeciesInThe WA Holotype: male, 'Wooramel R. 25°47.52'S Genus, AfterWatts (1978) 115°17.44'E 26.8.94 S.A. Halse', SAMA. (ListedInAlphabeticalOrder) Paratypes: 1, female, as for holotype, SAMA; 1, male, 'Gregory Gorge Fortescue R W.A. 2 xii.74coll. K. F. Walker', SAMA. Tiporus alastairi (Watts, 1978) Description Figs 1-3 DescriptionFig. 23 Length 5.6 mm. Elongate-oval, widest behind Length 4.0 - 5.2 mm. Oval, convex. Dark middle, outline strongly indented at junction of red-brown, underside lighter. Strongly and pronotum and elytra. Lateral edge of elytra not densely rugose-punctate all over. Pronotum curved forward immediately prior to meeting with a distinct raised ridge parallel to and a pronotum. Pronotum with well-developed lateral little distance inwards from each side, area ridge/groove on each side, edge ofelytron weakly just inside ridge depressed, ridge and serrated towards tip, tip weakly pointed. Dark depression strongest anteriorly, weak testaceous, rear edge of head, lateral edges of posteriorly. Elytron weakly margined, weakly pronotum, small patches at side of elytron near serrated towards apex, usually sharply pointed apex and appendages slightly lighter. Densely (rounded in male holotype). Edge of elytron rugose/punctate throughout. Pronotal process curves forward for short distance immediately narrow, keeled, bluntly pointed. Metacoxal lines prior to meeting edge of pronotum. raised, relatively wide apart, strongly diverging in Prothoracic process narrow, strongly convex, anterior half to about 2.5 times their narrowest roundly pointed at apex, only slightly width, raised lines not reaching metasternum. constricted between procoxae. Metacoxal Epipleuron evenly narrows until close to apex lines strongly raised, well separated, quite whereitabruptly ends. strongly diverging in anterior two-thirds, not Male. Protarsi three-segmented, anterior quite reaching mesosternum. portions greatly expanded and densely covered Male. Protarsus three-segmented. Anteriorsides with adhesive setae ventrally, third segment as of segments of protarsus moderately expanded, a long as first two; claw shorter than third segment, little expanded on mesotarsus. Single claw on WATTS 94 C. H. S. protarsus short, broad, weakly curved, near base. Tip of apical abdominal sternite broadly dorsoventrally flattened, anterior edge deeply groovedinmidline. SeeFig. 24foraedeagus notchedjust beyond middle. Protibia with a large Female. Protarsi five-segmented; basal three sharp tooth on outside just anterior to middle. segments of roughly similar size, weakly Apical sternite strongly carinate for a short expanded anteriorly, ventral surface covered in distance in middle near apex. See Fig. 23 for adhesive setae; fourth segment very small; aedeagus. apical segment narrow, cylindrical, about same Female. Protarsus five-segmented. Anterior length as third segment; claws weakly sides of pro-and mesotarsi less expanded than in developed. male. Apical abdominal sternite with ashort sharp ridge inmiddlenearapex. Remarks Tiporus centralis is very—similar to T. Remarks denticularis and T. georginae see discussion Females are difficult to separate from those of underthose species. T. undecimmaculatus or T. giuliani but are larger Originally known only from the male holotype, thanmostspecimensofthese species andtheedge the species is now known to be reasonably of the elytron curves forward for a short distance common in the Northern Territory and into immediately before it meets the edge of the Western Australia. It seems to have a more inland pronotum whereas it is straight or almost so in T. distribution than either T. collaris or 7". giuliani and T. undecimmaculatus. The same undecimmaculatus, the other common Northern applies to the males but in addition these have a Territory species. much shorter and broader proclaw and a much narrowercentral lobetotheaedeagusthan in these Distribution species. The species is so farknown only from the Northern Territory northofWesternAustralia. 2, 46 km SSWBorroloola, ANIC;W1,48 km SW by S Borroloola, ANIC; 3, 45 km Borroloola, Distribution SAMA; 1, Davenport Ranges, 40 km NE Murray WesternAustralia W Downs Station, SAMA; 1, Elkedra Homestead, 2, Derby, SAMA; 3, 4 km King Cascade, NTM; 6, Gosse River, Murchison Ranges, NTM; 15°38S 125°15'E, ANIC; 1, Kings Sound, ANIC. 2, Kakadu Highway, 31 km from Pine Creek, SAMA; 1, 35 mi NLarrimah, SAMA; 2, Moriarty Creek, SAMA; 1, Skull Creek (Victoria River), Tiporus centralis (Watts, 1978) SAMA; 10, VictoriaRiver, SAMA. DescriptionFig. 24 WesternAustralia Length 3.2 - 3.9 mm. Oval, convex. Black; 1, Stumpy Creek 'Glenroy', 16°55'S 125°34'E, SAMA. sides of pronotum and appendages reddish, sides and portions of middle of pronotum and base of elytron with vague red patches. Strongly and densely rugose-punctate throughout. Reticulate. Tiporus collaris (Hope, 1842) Pronotum not flanged. Elytron with a weak shallow stria near the suture, edge curves forward Description Figs 18, 21 slightly immediately before meeting pronotum, Length 3.3 - 4.1 mm. Oval, convex. Black; edge weakly to moderately expanded for a short undersides and appendages a little lighter. distance near, but not at, tip which is pointed. Strongly and densely rugose-punctate all over. Prothoracic process narrow, ridged in midline, Reticulate. Pronotum widely flanged at sides, apex rounded, weakly constricted between lacking raised ridges between disc and flange procoxae. Metacoxal lines strongly raised to except very slightly anteriorly. Elytron weakly metasternum, quite strongly divergent in anterior margined, moderately serrate towards apex which two-thirds. is rounded, base with vague longitudinal Male. Protarsus three-segmented. Anterior sides impressions. Prothoracic process narrow, strongly of segments of protarsus moderately expanded, convex, roundly pointed at apex, not constricted mesotarsus slightly expanded. Single claw on between procoxae. Metacoxal lines raised, slightly protarsus long, sharply curvednearbase, apicaltwo- to moderately diverginganteriorly. thirds straight. Protibiawith asmalltoothon outside Male. Protarsus three-segmented. Anterior sides SPECIESOFTIPORUS 95 FIGURES 16-24. 16, dorsal and lateral views of median lobe of aedeagus of T. undecimmaculatus; 17, ditto T. tambreyi; 18, ditto(lateral view) T. collaris(holotype); 19, ditto T.josepheni; 20,ditto T. giuliani; 21,ditto(dorsal view) T. collaris (holotype); 22, ditto T. denticularis; 23, ditto T. alastairi; 24, ditto T. centralis. From Watts (1978). WATTS 96 C. H. S. of segments of protarsus moderately expanded, pronotum, some small patches on elytron and those on mesotarsus alittle expanded. Single claw appendages yellowish. Strongly and densely on protarsus thickandcurved, strongly toothed on rugose-punctate throughout. Pronotum not underside near base. Protibia with a small tooth flanged, with a weak ridge with deep groove on on outside near base. Tip of apical abdominal inside, parallel to and some distance inwards from sternite moderately tuberculateinmiddle. SeeFigs sides. Apex ofelytron pointed. Epipleuron with a 18, 21 foraedeagus. small triangular expansion just before apex. Female. Protarsus five-segmented. Anterior Prothoracic process relatively broad, ridged in sides of segments of protarsus a little less midline, apex rounded, moderately constricted expanded than in male. Protarsus with two simple between procoxae. Metacoxal lines straight in claws. Apical abdominal sternite simple. posterior half, rapidly diverging in anterior half, notreaching metasternum. Remarks Male. Protarsus three-segmented. Anteriorsides Tiporus collaris is very similar to T. giuliani of segments of protarsus moderately expanded. and to T. undecimmaculatus. Males are readily Mesotarsus a little expanded. Single claw on distinguished from both these species by the basal protarsus short, flat, spindle-shaped, rapidly position ofthe tubercle on the protibia rather than narrowingnearapex on inside. Protibia with small the central or more apical position in T. giuliani tooth on outside near base. Tip of apical sternite and T. undecimmaculatus. In addition, T. collaris with small weak ridge in midline. See Fig. 22 for is slightly more rounded, has slightly weaker aedeagus. pronotal ridges/grooves and has rounded rather Female. Protarsus five-segmented, basal thansharply pointedtips totheelytra. segments somewhatexpandedoninside. The species occurs in small temporary streams in stony country across the north of the Northern Remarks Territory and Western Australia. Like T. Although the species has a weak ridge/groove undecimmaculatus, with which it is often onthe sideofthepronotum, the testaceous margin collected, it appears to be more coastal than T. of the pronotum, smooth elytron edge, lateral centralis. expansion of the epipleuron near the tip and the basal positionofthe tubercleon the male protibia, Distribution ally it to the T. josepheni group of species. NorthernTerritory Tiporus denticularis is very similarto T. centralis 8, 5 km SE Mt Borradaile, SAMA; 4, 6 km SE and females are difficult to separate, but they Mt Borradaile, SAMA; 1, 19 km E by S Mt differ from this species by the greater Borradaile, ANIC; 1, Canon Hill, Kakadu development of the pronotal ridge/groove and by National Park, SAMA; 1, CooWper Creek near Mt the extreme front edge ofthe elytron immediately Borradaile, SAMA; 17, 1 km Gubara, Kakadu before it meets the edge of the pronotum being National Park, SAMA; 1, 20 km SSW Jabiru, slightly curved in T. centralis but straight in T. SAMA; 1, Kambolgie Creek, SAMA; 4, 10 km E denticularis. by N Mt Cahill, ANIC; 1, Nawurlandja, Kakadu Originally only known from the Cairns- NationalPark, SAMA; 1, 6 km SWby S Oenpelli, Cooktown region of North Queensland the SAMA. additional specimens extend its known range into theNorthernTerritory. WesternAustralia 1, Carson Escarpment, 14°49'S 126°49'E, Distribution ANIC; 1, Drysdale River, 15°02'S 126°55'E, Queensland ANIC; 2, Kimberley, SAMA; 4, Mitchell Plateau, 1, 34 km NW Chillagoe, SAMA; 8, 70 km SW 14°40'S 125°44E, SAMA; 1, UpperCamp Creek, Greenvale, SAMA; 7, Lakeland Downs, SAMA; 14°49'S 125°51'E, SAMA. 3, Laura, SAMA; 4, Mary Creek, 16°33'S 12°5E, ANIC; 2, 11 km WSWPetford, QDPIM; 2, Walsh River near Chillagoe, QDPIM; 1, Windsor Tiporus denticulatus (Watts, 1978) Tableland, QDPIM. DescriptionFig. 22 Northern Territory Length 3.4 - 3.6 mm. Oval, convex. Dark red- 1, Bullita Outstation, 16°07'S 130°25'E, NTM; brown; sides, base and portions of anterior of 1,VictoriaRiver, SAMA. SPECIESOFTIPORUS 97 Tiporus giuliani (Watts, 1978) Distribution Northern Territory DescriptionFig. 20 4, Kakadu Highway, 31 km from Pine Creek, Length 3.6 - 4.2 mm. Oval, convex. Black; SAMA. appendages, some vague reddish areas at base of pronotum and base and sides of elytron, lighter. WesternAustralia Moderately strongly rugose-punctate throughout. 1, Beverley Springs, WAM; 2, Duncan Pronotum with a raised ridge parallel to and a Highway, WAM; 2, nearDampierDowns, WAM; little distant inwards from each side, area just 4, Logues Springs, 102 km SE by E Broome, inside ridge depressed, ridge and depression ANIC; 1, 163 km SEby S Broome, ANIC. strongest anteriorly, weak posteriorly. Elytron weakly serrated towards apex which is pointed. Prothoracic process narrow, strongly convex, Tiporusjosepheni (Watts, 1978). bluntly pointed at apex, only slightly constricted between procoxae. Metacoxal lines relatively DescriptionFigs 8, 19 close, strongly diverging in central third, weakly Length 3.5 - 4.1 mm. Oval, convex. Black, in anterior and posterior third, reaching sides of pronotum narrowly vaguely reddish, metasternum. appendages dark red-brown. Ventral surface Male. Protarsus three-segmented. Anteriorsides moderately and densely rugose-punctate, of segments of protarsus moderately expanded, dorsal surface closely and moderately weakly soon mesotarsus. Singleclaw on protarsus punctate. Pronotum not laterally flanged or short, narrow, with slight notch on underside in ridged. Tip of elytron weakly pointed. middle. Protibia moderately strongly toothed on Prothoracic process narrow, strongly convex, outside inmiddle. Apical sternite stronglycarinate only slightly constricted between procoxae, for a short distance in middle near apex. See Fig. apex rounded, midline virtually impunctate. 20 foraedeagus. Metacoxal lines well separated, moderately Female. Protarsus five-segmented. Pro- and diverging posteriorly and in middle, mesotarsi less expanded than in male. Apical subparallel in anterior quarter, reaching abdominal sternite very weakly carinate in middle metasternum. nearapex. Male. Protarsus three-segmented. Anterior sides of segments of protarsus strongly Remarks expanded. Mesotarsus a little expanded. Single Tiporus giuliani is close to T. claw on protarsus short, flat, narrowing abruptly undecimmaculatus and T. collaris. It can be on inside near apex, base with a large rounded separated from T. collaris by its stronger pronotal expansion beneath. Protibia with small tooth on ridges/grooves, sharply pointed elytral tips, outside in middle. Tip of apical sternite stronger colour pattern and the central position of depressed slightly in midline. See Fig. 19 for the spineon the male protibia. aedeagus. The only clear separation from T. Female. Protarsus five-segmented, robust, basal undecimmaculatus is the shape of the tip of segments somewhat expanded on inside. Elytron the median lobe of the aedeagus which is with very well developed lateral subapical spine broad with three small spines at the tip in T. (Fig. 8). undecimmaculatus (Fig. 16) and narrow and bluntly pointed in T. giuliani (Fig. 20). In the Remarks Northern Territory and Western Australia the Originally described from Beverley Springs and more apical position of the spine on the male Wittenoom Gorge in the Kimberley and PiIbarra protibia and the more pronounced dorsal regions of Western Australia respectively, the colour pattern will separate it from T. species is now known to be much more undecimmaculatus. However specimens of T. widespread, occurring right across coastal undecimmaculatus from Queensland resemble Northern Australia. It is particularly common in T. giuliani both in the central position of the North Queensland. It seems to favour larger and spine on the male protibia and in the well morepermanentriversthan other Tiporus. developed dorsal colour pattern. Unfortunately I mis-associated the sexes in my A less common species than some and possibly original description. (The female paratype in more inland thancoastal in distribution. ANIC belongs to T. tambreyi.) WATTS 98 C. H. S. Distribution sternite at apex raised for some distance either NorthernTerritory side ofgroove. See Fig.17 for aedeagus. 2, CooperCreeknearMtBorradaile, SAMA; 7, Female. Smaller, more densely punctate. MagelaCreek, SAMA; 2, MagelaCreek, 12km E Protarsus asymmetrically expanded as in male but Jabiru, SAMA; 1, Nourlangie Creek, 20 km SSW not to same extent. With small triangular Jabiru, SAMA. extension at tip of elytron, diverging from each otherwhenelytraclosed (Fig. 15). Queensland 1, Barron River, QDPIM; 52, 8 km N Remarks Bluewater, SAMA; 1, 25 km N Coen, SAMA; 2, A relatively large species known only from the 2 km NW Daintree, QDPIM; 2, Emu Creek 5 km Pilbara region of Western Australia, readily W Petford, QDPIM; 8, Eubenangee Swamp near distinguished from all other Tiporus by the Babinda, SAMA; 2, Helenvale, SAMA; 18, expanded male metatibia and the triangularly Lakeland Downs, SAMA; 1, Laura, SAMA; 2, extended elytral tips in the female. The sides of Mazlin Creek Atherton area, QDPIM; 1, 2 km S the pronotum straighten slightly in front giving Mt Molloy, SAMA; 2, 20 km S Townsville, thepronotum aslightly sinuate outline. RSiAvMerA;341k,m37NkWmCShiTlolwangsovei,llSeA,MSAA.MA; 7, Walsh Distribution WesternAustralia WesternAustralia 1, 17 kmNby ECaneRiverHomestead, ANIC; 1, Beverley Springs, WAM; 1, Stumpy Creek 15, Gregory Gorge, Fortescue River, SAMA; 16, 'Glenroy' 16°55'S 125°34'E, SAMA. MWbiyAllMWst;rMe9ia,lml,1stkArNmeIaNmC,;MAi1lN,lIMsCitl;rlesa1tm,r,eTaAamNm,IbPCrae;yl4,m,PW3oAkolMma;NreWa1,, Tiporus tambreyi (Watts, 1978) WittenoomGorge, ANIC. DescriptionFig. 17 Length 3.5 - 4.4 mm. Oblong oval, strongly Tiporus undecimmaculatus (Clark, 1862) convex, pronotum a little constricted in anterior quarter. Black, extreme anterior of head, sides DescriptionFig. 16 ofpronotum narrowly, a basal spot and extreme Length 3.4 - 4.6 mm. Oval, convex. Black or lateral markings on elytron and appendages dark red-brown; underside and appendages a little reddish. Moderately strongly and very densely lighter, baseofpronotum and sidesofelytron with punctate all over, punctures on pronotum and vague red patches in many. Strongly and densely elytron with short setae. Reticulation strongest rugose-punctate all over. Reticulate. Pronotum anteriorly. Pronotum and elytron weakly with a distinct raised ridge parallel to and a little margined, margins of elytron weakly serrated distance from each side, area just inside ridge towards apex, extreme apex not margined. depressed, ridge and depression strongest Prothoracic process lanceolate, quite strongly anteriorly, weak posteriorly. Elytron weakly expanded behind procoxae, narrow between margined, strongly convex, rounded towards apex procoxae, weakly flanged, strongly carinate. which is pointed. Prothoracic process narrow, Metacoxal lines strongly raised, rapidly strongly convex, roundly pointed at apex, little if diverging in anterior half, subparallel at all constricted between procoxae. Metacoxal posteriorly. lines raised, slightly to moderately diverging Male. Anterior sides of segments of protarsus anteriorly. moderately expanded, posterior sides Male. Anterior sides of segments of protarsus unexpanded. Basal segment of mesotarsus moderately expanded, those on mesotarsus a little weakly expanded. Single claw of protarsus expanded. Single claw on protarsus weakly greatly expanded in basal half, flattened curved, slightly thickened and with a small tooth dorsoventrally. Metafemur robust, curved. on underside nearbase and anotherjust before the Metatibia angularly thickened on inside near middle. Protibia with a small sharp tooth on apex. Protibia with a small tooth on outside close outside in apical half. Tip of apical abdominal to base. Last abdominal sternite widely and sternite weakly to strongly tuberculate in middle. weakly grooved in midline with a small knob in SeeFig. 16foraedeagus. middle of groove at extreme apex, edges of Female. Anterior sides ofsegments ofprotarsus