J. New York Entomol. Soc. 105(3—4);199—205, 1997 THE ANT (HYMENOPTERA: FORMICIDAE) TYPES IN ARGENTINIAN COLLECTIONS Donat Agosti Department of Entomology, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024-5192 — Abstract. Type specimens of 114 species and subspecific ant taxa in Argentinian insect collections are listed, including 14 holotypes and 5 lectotypes. Most ofthe types are Neotrop- ical, with the exception ofa series of Karavajev types from South East Asia. The two collection at the Division Entomologia, Museo Argentinao de Ciencias Naturales in Buenos Aires includes (MACN) and the Fundacion e Institute Miguel Lillo, Universidad Nacional de Tucuman in Tucuman (IMLA), house all ant types in Argentina. The collection of 50 Kusnezov types at IMLA, the Bruch collection MACN and thirteen Karavajev types from SE-Asia in is the most important holding because of the 14 holotypes and 2 lectotypes. The latter includes one holotype of each W. L. Brown, Jr., Th. Borgmeier, and W. Kempf. The Karavajev types are in a small separate box in MACN, which is labeled as follows: “Donacion Dr. Alejandro A. Oglobin” and “Hormigas tejedoras, Formici- dae, gen. Oecophylla, Polyrhachis, tipos del mirmecology Karawajew”. The box includes also 19 further species identified by Karawjew. Bruch’s collection is at MACN, but his extensive photographic documentation is at the Museo de la Plata in La Plata (MLPA). Type designation in myrmecology has in the past been rather loose. Ants, being social, led to a long upheld practice ofnot designing a holotype, but rather designate syntypes, types, including a series of specimens. The list is organized alphabetically following subfamilies and genera. Only the often fragmentary information provided on the labels are listed. The taxa are listed following the valid combination. If this does not agree with the label data, the orig- inal genus used on the label is provided in brackets. Types for which no correspond- ing description can be found are nevertheless listed. Further information on the species can be found in Bolton (1995). The abbreviations of the collections follows Arnett et al. (1993), which includes also the addresses of the collections. CERAPACHYINAE Acanthostichusfemoralis Kusnezov Syntypes 17 workers, #10449; Argentina, For- mosa, Ing. Juarez, N. Kusnezov, viii. 1958, IMLA [manuscript name?]. Cerapachys silvestrii Mann Cotype 3 workers PR. Rio Pideras, Insular Expedition Station, F. Sein Jr., Feb. 5, 30. IMLA. In soil in sugar cane [manuscript name?]. DOLICHODERINAE Dolichoderus {Monads) andinus Kempf. Paratypes 2 workers, 1145, Peru Pichita, Caluga, 2,150 m, Weyrauch, 18.9.1960. IMLA. 200 JOURNAL OF THE NEW YORK ENTOMOLOGICAL SOCIETY Vol. 105(3-4) Dolichoderus (Monads) lobicornis Kempf. Paratypes 3 workers. Blumenau, M. Wit- te, 10.12.1921. IMLA. Dorymyrmex (Dorymyrmex) emmaericaellus Kusnezov. Holotype and paratypes 6 workers. Bolivia, Cochabamba, La Paz. IMLA. Forelius (Neoforelius) tucumanus Kusnezov. Syntypes 2 females. #5617, Tucuman, N.Kusnezov. IMLA. ECITONINAE Edton mexicanum argentinum Borgmeier. Holotype 1 female. #32890, Roque Saenz Peria (Chaco), F. Ohnehider. MACN. Edton setigaster ^orgmdcY. Cotype 1 female. #11804, Bolivia. MACN. Neivamyrmex Edton hetschkoi oglobini Santschi. Typus 1 female. Argentina, Mi- siones Dr. A. A. Oglobin. MACN. Neivamyrmex hetschkoi Mayr. Cotype 1 worker. #2001, Argentina, Misiones, Est. Exp., Loreto, Dr. A. A. Oglobin. MACN. Neivamyrmex impudens W. M. Mann. Cotypes 3 workers. Honduras, Progresso, W. M. Mann. IMLA. Neivamyrmex laevigatum Borgmeier. Cotypes 2 workers, 1 male. #2580, Argentina, Rogne (?), S. Pena, N. Kusnezov, 13.6.48. IMLA. Neivamyrmex (Edton) pseudops grandipseudops Bruch. Typus 1 female. Argentina, Alta Gracia, La Granja, Sierras de Cordoba, C. Bruch. MACN. EORMICINAE Acropyga (Rhizomyrma) wheeled Mann. Cotypes 3 workers. Honduras, Lombardia. IMLA. Camponotus aguilerai Kusnezov. Syntypes 10 workers. Argentina, Jujuy, 6.1.1949. IMLA. Camponotus goeldii Forel. Syntype 1 worker. Brazil, Rio de Janeiro. IMLA. Camponotus guayapa Kusnezov. Holotype and paratypes 6 workers. LaRioja, Guay- apa, 12.8.1948. IMLA. Camponotus phytophilus Wheeler. Cotypes 4 workers. #2834/sk., Mexico, Guema- vaca, E. Skwarra, 6.26.1929. IMLA. Camponotus querdcola M. R. Smith. Paratypes 6 workers. USNM paratype#62025, Los Angeles, Tambark flat, T. C. Laurence, 7.15.1952. IMLA. Myrmelachista ina Kusnezov. Holotype and paratypes 36 workers. Argentina, Mi- siones, Manuel Belgrano, N. Kusnezov, 7.26.1949. IMLA. Myrmelachista schachovski Kusnezov. Syntypes 2 workers. Argentina, San Martin de Los Andes, S. Shajovskoi. IMLA. Myrmelachista ute Kusnezov Holotype and paratypes 20 workers. Argentina, Mi- siones, M. Belgrano, N. Kusnezov, 7.27.1947. IMLA. Myrmelachista vicina Kusnezov. Holotype (male), paratypes 1 male, 2 females, 1 worker. Neuqueu, Lago Cacar, Schachovskoi. IMLA. Camponotus mucronatus hirsutifrons [manuscript name?]. Types 6 workers. Guate- mala, Zacapa W. M. Wheeler, 12.13.1911. IMLA. Paratrechina dichroa Karavajev. Typus 1 worker. #2399, Prinseneiland, Karavajev. MACN (Karawajew). 1997 ANT TYPES IN ARGENTINA 201 Paratrechina koningsbergeri Karavajev. Typus 1 worker. #3869, Java, Jibodas, Ka- MACN ravajev. (Karawajew). Polyrhachisflavoflagellata Karavajev. Typus 1 worker. #2875, Java, Buitenzorg, Ka- MACN ravajev. (Karawajew). Polyrhachis keratifera Karavajev. Typus 1 worker. #2505, Amboina, Karavajev. MACN (Karawajew). Polyrhachis mystica Karavajev. Typus 1 worker. #2353, Java, Buitenzorg, Karavajev. MACN (Karawajew). Polyrhachis ochracea Karavajev. Typus 1 worker. #35090, Java, Buitenzorg, Kara- MACN vajev. (Karawajew). Polyrhachis rastellatajavana Karavajev. Typus 1 worker. #2875, Java, Buitenzorg, MACN Karavajev. (Karawajew). Polyrhachis rotundiceps Karavajev. Typus 1 worker. #3004, Wammar, Ins. Am, Ka- MACN ravajev. (Karawajew). Polyrhachis schang gracilior Karavajev. Typus 1 worker. #2503, Amboina, Kara- MACN vajev. (Karawajew). Polyrhachis tibialis orientalis Karavajev. Typus 1 worker. #2439, Tifoc (Bocroc (?)), MACN Karavajev. (Karawajew). Polyrhachis tibialis robustiorKaravajev. Typus 1 worker. #2583, Wammar, Ins. Am, MACN Karavajev. (Karawajew). Polyrhachis tubifex Karavajev. Typus 1 worker. #2359, Java, Buitenzorg, Karavajev. MACN (Karawajew). MACN Pseudolasius carinatus Karavajev. Typus 1 worker. #2537, Aroe, Karavajev. (Karawajew). MYRMICINAE Atta opaciceps Borgmeier. Paratype 1 worker. Brazil, Tapera, Pemambuca, F. Car- valho, 1.941. IMLA. Atta robusta Borgmeier. Paratypes 2 workers. Brazil, Rio de Janeiro, Sao Bento, F. Torres, 15.X.935. IMLA. Basiceros singularis F. Smith. Lectotype (designated by W. L. Brown) 1 worker. B. G., Kumpung, H. C. Lang. IMLA. Basiceros squamifer Borgmeier. Syntypes 3 workers. #5510, Rio de Janeiro, Angra dos Reis, Jussaral, Lopes and Lent. IMLA (ex coll. Borgmeier). Chelistruma lilloanus (Brown). Holotype and paratypes 3 workers. Argentina, Tuc- uman, N. Kusnezov, iii.20.48. IMLA. Crematogaster hyperphyes Kusnezov. Paratype 1 female. #7080, Bolivia, Chapare, Marcus, 1.7.49. IMLA. Crematogaster hyperphyes Kusnezov. Paratypes 1 worker and 1 male. #7079, Bo- livia, Chapare, Marcus, 27.viii.49. IMLA [no type labels, but same blank red standard labels]. Lepthtoraxpergandei Emery. Cotypes 2 workers. USNM Cotype #54078, USA, DC, Washington, 9.10. IMLA. Monomorium minimum cyaneum Wheeler and Mann. Paratypes 3 workers. #128, Mexico Hidalgo, Guerrero Mill, Mann. IMLA. 202 JOURNAL OF THE NEW YORK ENTOMOLOGICAL SOCIETY Vol. 105(3^) Monomorium (Monomorium), minimum emersoni Gregg. Paratypes 3 workerUSNM type #57292, USA, Texas, Austin, A. Emerson, iv.6.1937. IMLA. Neostruma zeteki Brown. Paratypes 2 workers. #5059, Panama, Barro Colorado Is- land, Zetek, Feb-March 43. IMLA. Pheidole eglobini Kusnezov. Paratypes 2 workers. #4699, Argentina, Misiones, Ya- cuy. IMLA [manuscript name?]. Cyphomyrmex hillbramus Kusnezov. Paratypes 2 workers. Argentina, Salta, R. So- ladillo, R.55-km 1442, 25.1.948. IMLA [manuscript name?]. Cyphomyrmex ulemli Kusnezov. Syntype 1 worker. #8849, Argentina, El Rey, Suita, 6.2.53. IMLA [manuscript name?]. Ochetomyrmex {Brownidris) argentinus (Kusnezov). Syntypes 1 female, 5 workers. Argentina, Misiones, Loreto, Dr. A. A. Oglobin. IMLA. Ochetomyrmex {Brownidris) argentinus (Kusnezov). Syntype 1 female. Tucuman, Trancas, ii.l947. IMLA. Ochetomyrmex {Brownidris) bolivianus (Kusnezov). Holotype 1 female. Bolivia, en- tre Wames y Montero, Sta. Cruz, Wygodzinsky, 27/28-1-1958. IMLA. Octostruma stenognatha Borgmeier and Kempf. Paratype 1 worker. Brazil, Nova Teutonia, 27°11'S, 52°23'W, 3-500 m, Fritz Plaumann, vi.l957. IMLA. Oligomyrmex {Erebomyrma) eidmanni (Menozzi). Syntypes 6 workers. Resistencia Chaco, 6.17.948. IMLA. Oligomyrmex {Erebomyrma) eidmanni (Menozzi) Syntypes 3 workers. Brazil, Men- des, H. Eidmann, 3.10.33. IMLA. Oligomyrmex stenopterus Kusnezov. Types 2 females. #6087, Argentina, Misiones, Est. Exp., Loreto, Dr. A. A. Oglobin. IMLA. Orectognathus clarki Brown. Paratype 1 worker. Australia, Victoria, Femtree Gulley, W. L. Brown, 1.1.58. IMLA. Oxyepoecus {Martia) inquilina (Kusnezov). Holotype 1 worker. #1832, Argentina, Tucuman, Jardin Ins. M. Lillo, N. Kusnezov, 2.v.1948. IMLA. Oxyepoecus {Martia) minuta (Kusnezov). Holotype 1 worker. Tucuman, Quebrada Cainzo, N. Kusnezov, 8-iv.l948. IMLA. Pheidole bambusarum Forel. Cotypes 9 workers. Brazil, Sao Paulo, Grand bambou foret. Lutz. IMLA. Pheidole carapunco Kusnezov. Syntypes 1 soldier, 6 workers. #1295, Argentina, Tucuman, Carapunco, 2,600 m, N. Kusnezov. IMLA [1 worker lectotype desig- nated by E. O. Wilson]. Pheidole descolei Kusnezov. Lectotype and paralectotypes 9 workers. Argentina, Salta, Apolinario, Saravia, E E. Luna, 1948. IMLA [1 workerLectotypedesignated by E. O. Wilson]. Pheidole gavrilovi Kusnezov Lectotype 1 worker, paralectotypes 1 female, 6 work- ers. #5137, Argentina, Misiones, Loreto, 19.vi.949. IMLA [1 worker Lectotype designated by E. O. Wilson]. Pheidole {Elasmopheidole) lilloi (Kusnezov). Holotype 1 female. Tucuman, 7.i.48, IMLA. Pheidole minutissima Kusnezov. Lectotype worker, paralectotypes 5 workers. #5348, Argentina, Misiones, Eldorado, N. Kusnezov. IMLA [1 worker Lectotype desig- nated by E. O. Wilson]. Pheidole {Elasmopheidole) subaberrans (Kusnezov). Paratypes 2 workers. #1679, 1997 ANT TYPES IN ARGENTINA 203 Salinas, 16.4.48; 1 female. #4586, Salta, Oran, 23.11.48; 2 females. Rio Calera, 25.4.48; 1 worker. #1792, ?. IMLA. Pheidole (Eriopheidole) symbiotica (Kusnezov) Paratypes 1 female, 1 male, 7177, La Picaga, N. Kusnezov, v.1951. IMLA. Pogonomyrmex anergismus Cole. Paratypes 2 females. #2911, USA, New Mexico, A. C. Cole, Sept. 12, 1961. IMLA. Pogonomyrmex guatemaltecus Wheeler. Cotypes 3 workers. MCZ-Cotype 20—24 20577, Guatemala, Zacapa, W. M. Wheeler, Dec. 12, 1911. IMLA. Pogonomyrmex longibarbis andinus Kusnezov. Paratypes 3 females, 2 males, 3 workers. #5712, San Antonio de los Cobres, 22.12.49. IMLA. Pogonomyrmex longibarbis andinus Kusnezov. Paratypes 3 workers. La Rioja, San Refugio del Penon, 4,000m, Monros, 1.47. IMLA. Pogonomyrmex marcusi lobatus Kusnezov. Paratypes (?) 3 workers. Bolivia, Co- chabamba, La Paz, N. Kusnezov. IMLA [unlabeled standard red type label]. Pogonomyrmex meridionalis leonis Kusnezov. Paratypes 4 workers. #5834, Santa Cruz, Canadon, N. Kusnezov. IMLA. Pogonomyrmex odoratus Kusnezov. Paratypes 3 males, 6 workers. #3553, Cerro Olto, 6.2.49. IMLA. Pogonomyrmex odoratus Kusnezov. Paratypes 1 female, 1 worker. Bariloche, 6.2.49. IMLA [unlabled red type label]. Pogonomyrmex rugosus Emery. Cotype 1 worker. USNM type #54071, USA, Cali- fornia, S. Jacinto, T. Pergand. IMLA. Procryptocerus marginatus Borgmeier. Paratype 1 worker. Brazil, Bahia, Uru^uca, P. Silva, 1947. IMLA. Procryptocerus montanus Kempf. Paratypes 12 workers. #1645, Brazil, SP, Campos do Jordao, Kempf, 14.xi.l956. IMLA. Solenopsis (Labauchena) acuminata (Borgmeier). Cotype 1 female. Luna. IMLA. Solenopsis basalis major Forel. Cotypes 6 workers. #12085, Argentina, La Plata, Bruch, 9.10.1912. IMLA. Solenopsis clyptemnestra bruchi Forel. Cotypes ? #12093/92, Argentina, La Plata, Bruch, 1892. IMLA. Solenopsis clyptemnestra orestes Forel. Cotypes 22 workers. Brazil, Ceara, Roche. IMLA. Solenopsis (Synsolenopsis) egregia (Kusnezov). Paratypes 4 females, 11 workers. #1298 Argentina, Tucuman, Tafi del Valle, N.K., 3.10.48. IMLA. Solenopsis {Synsolenopsis) eximia (Kusnezov). Syntype 1 worker. #2411, Uriburu, Formosa, 6.21.48. IMLA. Solenopsis globularia Forel. Syntypes 2 workers. IMLA. Solenopsis hammari carhuensis Forel. Cotype 1 worker. Buenos Aires, Bruch. IMLA. Solenopsis latastei hojfmanni Forel. Syntypes 5 workers, 7 females. IMLA. Solenopsis (Bisolenopsis) sea (Kusnezov). Syntypes 4 males, 4 females. #752, Ar- gentina, Santa Fe, Villa Ana, H. Willink, 2.1946. IMLA. Solenopsis {Paranamyrma) solenopsidis (Kusnezov). Paratypes 2 females. #7122, La Picaga, E. Rios, N. Kusnezov. IMLA. Solenopsis {Paranamyrma) solenopsidis (Kusnezov). Paratype 1 female. #4996, Mi- siones, Iguazu, N. Kusnezov, 25.7.49. IMLA. 204 JOURNAL OF THE NEW YORK ENTOMOLOGICAL SOCIETY Vol. 105(3^) Solenopsis spei Forel. Cotypes 6 workers. Esperanza, A. Forel, in nest of Atta ce- phalotes. IMLA. Strumigenys planeti Brown. Paratypes 2 workers. Bolivia, Beni, Huachi, W. M. Mann, 1921-22, MuIIford Exp. IMLA. Strumigenyspraecava Brown. Paratypes 3 workers. MCZ-Paratype#29293, Panama, Barro Colorado Island, W. M. Wheeler, 6.27.24. IMLA. Tetramorium (Triglyphothrixl) desertorum (Forel). Syntypes 8 workers. IMLA. Wasmannia williamsoni Kusnezov. I worker. #6060, Gral, Pico. IMLA [unlabeled red type label] Zacryptocerus (Paracryptocerus) borgmeieri (Kempf). Holotype and paratypes 6 workers. #5330, Misiones, Iguazu, N. Kusnezov, 29.7.49. IMLA. Zacryptocerus {Paracryptocerus) borgmeieri (Kempf). Paratype I worker. Misiones, Iguazu, N. Kusnezov. IMLA. Zacryptocerus {Paracryptocerus) cojfeae (Kempf). Paratype 1 female, I worker. Co- lombia, Cundinamana, Tibacuy, 18.iii.952. IMLA. Zacryptocerus {Paracryptocerus) cojfeae (Kempf). Paratype 1 worker. Colombia, Cundinamana, 1500m, Bemel + Mendoza, 13.3.1952. IMLA. PONERINAE Ectatomma regis Kusnezov. Holotype 1 worker. Argentina, Salta, El Rey, Nogues N. Kusnezov, 14.ii.1953; paratype 1 worker. #6185, Argentina, Tucuman, Ruta 9 km 1335; paratype 1 worker, Argentina Rio Sali, 40 km of Tucuman 10.1.48 IMLA. Metapone hewitti Wheeler. Paratype 1 worker. F946, Malaysia, Borneo, Kuching, J. Hewitt. IMLA. Metapone tillyardi Wheeler. Cotype 1 female. MCZ-Cotype 1-2 20787, Mt. Tam- bourine, A. M. Lea. IMLA. Pachycondyla {Neoponera) goldbachi goyana (Kusnezov) Holotype and paratype 2 workers. #4792, Argentina, Misiones, Iguazu, 7.24.1949. IMLA. Probolomyrmex brujitae Agosti. Holotype, female. Argentina, Jujuy Aguas Blancas- Yaculica (Argentinian-Bolivian frontier), 22°43'44"S 64°22'25"W, 460 m, 25 Oc- tober 1994, D. Agosti & J. M. Carpenter. PSEUDOMYRMECINAE Pseudomyrmex acanthobia cocae Santschi. Typus 6 females, 3 males, 7 workers. Argentina, Alta Grande, La Granja, Sierras de Cordoba, C. Bruch, 1.922. MACN. Pseudomyrmex arborissanctae symbioticus Forel. Cotypes 6 workers, #12010, Co- lombia, Dibulla. IMLA. Pseudomyrmex {Pseudomyrma) championii haytiana paulina (Forel). Cotypus 1 fe- male, 1 worker Argentina, Bruch, 989. MACN. Pseudomyrmexflavidula laevivertexlizeri Santschi. Cotype 1 worker. #1144, Bolivia. MACN. Pseudomyrmex {Pseudomyrma) gracilis argentina (Santschi). Typus 2 workers. #2026, Argentina, Misiones, Loreto (Est. Exp.), Dr. A. A. Oglobin. MACN. Pseudomyrmex {Pseudomyrma) gracilis glabriventris (Santschi). Typus 1 worker, cotypus 1 female. #1170, Bolivia. MACN. 1997 ANT TYPES IN ARGENTINA 205 Pseudomyrmex (Pseudomyrma) gracilis sericata (Santschi). Typus 4 workers. Par- MACN. aguay. Pseudomyrmex oglobini Santschi. Cotypes (?) 9 workers. #2088, Argentina, Mi- siones, Loreto, Dr. A. A. Oglobin; Nido ne las tamas de lapacho (Tecoma Ipe) de 35 mts de altura - Patoreo (Grande) IMLA. ACKNOWLEDGMENTS This work was supported by a grant from NSF to Platnick. Willems was very helpful in organizing the visit in Tucuman as well as discussing the remain ofother ant collections in Argentina. This compilation is part of a survey of Argentina, supported by the Argentinian National Parks through the necessary collecting per- mits. LITERATURE CITED Arnett, R. H., Jr., G. A. Samuelsen, andG. M. Nishida. 1993.TheInsectand SpiderCollections ofthe World. Elora and Eauna Handbook 11:310pp. Bolton, B. 1995. A NewGeneralCatalogue ofthe Ants oftheWorld. HarvardUniversityPress, 504 pp. Received and accepted 12 December 1997.