© ZooloIgnisctaiSltt u.tPee,t2 e0r 0s2burg, StudoineN se arcstpiecoc Lfie eusc (oDpiisp Cthearmaa:e m yiidae). IT.hree descorifpNteiarLocentu iccpo upbisl biesfh1oe9rd6e 5 VN..T anasijtshuk TanasijVt.sN2h.0u 0kS2,t. u doinNe esa rscpteicocif le esu co(pDiisp Cthearmaa:e ­ myiiIdT.ah ere)e .d escorfiNpetaLireocuntc ipocup isb lbiesfoh1re9ed6Zo 5o.s ystematica Ross1i1c(a11,9) 3:- 207. ReviosfiNoenca t rycpoteLfise uc(opDiispC thearmaa,e mdyeisicdbraeiefob)1re 9ed 6 5 warse alTiyspeaednsc.d o rresmpaotnedwrieinrsagetl usd foirte hfodel loswpiencg ies: L.ameriMcaalnla1o 9c2hL1,.a; t r!faAclider1si9 c2Lh5.,;fl aviAclodrrn1ii9cs1Lh 4.,; maculTahtoam p1s9o01Ln; .m, aj oMra ll1o9c2hL1,.m; i nMoarl l1o9c2hL1,p.;a ra/le/a Mall1o9c2hL1,.p; e mphMiagalel1 o9c2hL1,.p; i nipMearldla1o 9c2hL1,.s; i mplex Loe1w8,6 9. V.N.T anasijZtsohoulko,Ing sitciaRtluu stseAi,ca ande moyfS cienUcneisv,e rs itetskaya nabI.S, t .Pet1e9rs9b0uR3rgu4 s,se i-am;ab irla:c h@z in.ru Introd uction ofm ansyp ecuiseifesnm ga laerdseo omteobd e uncerutnatsiionlmc el eadrelfiync erdi taer reia Thsep ecoifLe esu cMoepiisg (eDni pCthear­ad:e velaonpided de ntwiifitpehad r tiscpuel caire s maemyiwihdoalseae)r pvraeoeynA phiadna de whosmea lwea sc orredcetslcyrF irubtehde.r ­ Cocciadswa eeal,slP syl(lSitdeave1e 9n6s 7o;nm,o raem ,a lheolt oy wpiet fehm aplaetr yap (oe rs McLea1n9,9 8p)l,aa yv eriym porrtoalin net v icvee rqsuai)ot fetb eenl otnodg i ffesrpeen­t manbyi oceHnoowseesvste.ur do ifte hgsir so upci es. havbee eunn eviendn i fferreegnitAo lnmso .s tIi ntteopn udb lasi esrhoi fae rst iocnNl eeasr c­ 70s pecoifte hsgi esn aurske n owfrno mt h etiLce uctohpoiinssie d;s e vottsoep de cwihei sc,h Palaeabrulctet tsihsca 2,n0 s pecwieerdse­e werpeu blibsehfoertdeh w eor ko fM cAlp ine scrifrboemtd h Nee ar(cMtciAcl p1i9n6 e5,,( 19p6u5b)l iisn"h Ace adt aolfto hDgei ptoe fra 197179,7T 8a;n asi1j9t9sn6ho)aut,lko l,fw hi chAmerniocraot fMh e xicToh"id.sa tweac sh o­ currebnetllotynot g h giesn Tuhsi.cs ab ne e x­sebne caduessec ripputbiloilnsasht weeedr r e plaibnyte hdfae c tth daett ermionfma ot sitotny piccaarlrloiyuie tndm ordee twaiitdlhr aw­ specoifLe esu cboype ixst emrnoarplh ol ogiincogafstl hmea lgee ni talia. featuirsae lsm oismtp osbseicbalouefts he e ir uniforTmhietrye.afo d rees,c roifop ntseip oen­ Matearnimdae lt h ods ciceasbn es imitlota eornt h(eo rfcs o utrhse er e arseo msep ecwiietdshi stifenacttuitrvhe eas t Durimnysg t ianyt hUeS Ai n1 99I2v ,i s ited areea styod etermsiomn oess)tp, e cciaebns e thceo lleocftt hiMeou ns eoufmC ompar ative deteromnilbnyyep dr epaorfta htmeia olgnee niZ­ooloHgayr,vU anridv e(rMsCiZtw)yh, e Ir e talSiiam.id leatre rmiunsaifetnmigao lntee r­studtihteeyd op fLe e ucsoipmipLsol eeWwxh. i le minailnsio wate dlelv elaottph meeod m ebnut t,i nC anaidn1a 9 9I4s ,t udtihveead cs otl le ction hapso tenatsdi eamlo,n sbtyrR aats(ep1di9 85o,fL euciontp hiCesa nadNiaatniC oonlalle ction 198169,9 109,9 6R)a,s& p Bie rto(l1i9n 9i3o )f,lnsOetctta(swC,aN CD)rJ;.. M Fc.A lpgian vee Bescho(v1s9k9ai5n )Bd,e scho&v Msekr iz meh idsr amfta nuscarbioLpuettus c ionpcilsu d­ (199H8o)w.e avfe emra leev,ei cnfo llaenc dteidnt ghs ep echiehe assst udTiheedms.ae n us cripts reaornet dhs ea maep haisds sp ecX,i ceasbn e ­inclauk deeuyds ienxgt emrnoarlp hoblu otgy, lontgos pecYi;te hsee x cepatrimeoo nnso ­withgoeunti sttarluiccT thueIrnw e o.r kfoer d phagsopuesc sipeesc,ci leesad relfiyn aobn let hrmeoen twhistt hhc eo lleocfLt eiuocniosn p is extearnpaple araansndpc eec,ri eeasir nce udl ­thNea tiMounsaelou fNm a tuHriasltW oarsyh,­ tufrroem k nowpna rebnymt est hdoedssc riibnegd(tNoMnN Ha)nt dh Iel liNnaotiuHsri aslt ory byG aim&aT ruirn (e1r9 9S7od),e. t ermin aStuirovnUesryb,a( nIaN HISr )e.c eailvases ode r ies 194 VN.T anasijNtesahrusckpt:ei ccoi fle esu coIp• iZ sO.O SYSRTO.S SIVoClA1. 1 oft yspep ecifrmoemDn rsW. .P ulawCsakliiw,­i thhoeuatwd i)t,th hs ea mlea b"e4l65 6A8c"c.o rding fornAicaa deomfSy c ie(nCcAeSsB) e.s idteors e coart!d NsH Sl,a b"e4l6 5m6e8a""n Dsi pdtee­ra thessoemm,ea tewreirsaetl usdi itneh cdeo llesct­roaypihnigd s; IEvlMalan.ys 1 t,51o 9n1,D2 .. K . McMil lan". tioofnC SIR(OC anbAeursrtara,an ltdih ae) DescrFiepmtaiBlooendl..ye n2g.t1hm- m2 ..2 ZooloIgnisctoaifltt huRetu es sAicaand oe mfy Bodlyi gghrteH ye.a1 d. 5t-i1m.ae6shs i ag sh ScieSntcP.ee st,e r(sZbIuINrnt)g o. It s atlu dileodnFgr.o2 n.s7 -t2i.mn8ea sr rtohwhaeenr a d. morteh 2a0n0 s0p eci mens. Ocelpllaawrte ea kmlayr kfreodn;st tarldi a prek Aftreert utrnoSi tn.gP etIer rescbeuair vge,d grewyi,tt whdo a brrko swtnr oinip tesssi O dT e. signipfiocrat(nimtoo tnrh ea7 n0s 0p ecimens) ind1e.xL6 u.n umleed ium-Asnitzeaendndn .a oft hceo lleocLfte iuocnbosep liosnt gCoiN nC ga ribsltaa3c rkad,r isttohmrteeiremel e osn ger anNdM NHp,r ovitdhfiern agm eowfto�i r skt han 2nhde.i 4gGt heitnm laee sts hsea yne stutdoby eg reaextplayn AdsDe rdM..c Alpme heigMhitn.ub trei satrlreasin,nog nesedp a rse (perscoonmamlu niacsastuiimoatenp d)p, e ars rowp,r esoengn etn aan;t ebrriiolsrat rlgee st. ratehxetre nsive. Lateraolfm esstornigopotelusdbm ir soehwx n­, McAlp(i1n9ee6 n5u) me1r5as tpeedco i fes tendtiodn igs 1t/ao4lfm esonoMteudmi.a n Leuc(ot phaies snu bgoeLfne uusc otphwiaest r) e strgirpeAeyrs.e b aest wseterindp eevsoo bfiri ds ­ descrfriobmte hdNe e aracntodin cfre o mt h e tloevs2e /ro3 fm esonTowtpoua mi.or fds o rso­ nortNheeornt r(oCpuibFcais)ov .fteh e(mb elclean btrriaspltr leesLsee ngbtsl. aa cpki;oc fefes­ Loe1w8,6 b6e,l lWuillal i1s8t8bo9in,v, i tmtoartaaanb da soetfs i byiealelo onfow rl;ee tgasr si Mall1o9c4ho0,c, e lMlaalrli1os9c 4ah0n,v d e r­neaernltyib rleaolcnymk i,d hainnldde fi grss t ticMaallilso1 c9h4w,0e )r sei ngoluet d btywtoa rsoymeelrlOeonsww i.n vgesi,Mn asn d McAlp(i1n9e6i 0ntt)ho se e paorcaetlel aRr icso­nviednri gse1t /at3l1p, . t3i mleetssh sa n groTuhpe.gi ern eric caormbebe imngad 4ti+i5s sotenacslto ifCo uLn.a rbgleas cpkco otv enng chanbgyTe anda sij&tG sahium(kaip rnri e ts os) synte1r+gu2ip tt ope o s?tree drgitew;ro ?a etr thgee nusLeuacloowpniig1tn 5nha e ,sw p e cisemsa.slplo ts opntr eers3ge;1dn taterm ke dian Ouotft hree mai1n1oi,nn (ego rbMiatlall oicsshp ,optrse soenan ntt eerdeigo oftr e rg4i-t5e.s 192i1as)s ynoonfyL m.a rgentZiectotlelrTi­asx onnoomtTiehcsde .e scroifMp atliloonc h ste1d8t4(,8M cAl&p Tiannea si1j9t7sI2h )u.k , (1 92i1is)n disFtoeirxn acmthp.ewl reo,"t Teh o­ havset udtihteeyd pa enosdt hmeart eorfti h ael rawxi otwrhi thtowbuort o vwint tbuaated " d,e d 10r emaisnpienctgih reees s;bu elittnshsg eu b- thtahtte y psiecra"ilhe tashst e h ovriatx'.·tI ante' jeocftt h sitsu dy. reatlhipetr ye,s oeran bcseeo nflc aets etrrailp es Unfortunantoaetllo lyft , ht ey psep eci mens ivse sreyr tiaoxuosn cohmaircaa ncLdta. em re,r i­ wermea lweh;ai mtso r-seo metsipmeecsi mens cansae nMsaul lmoacbyhec onceaiascv oemd­ that wearsme a licenit theeddie rs cr iptions ploefsx p ecSiteusd.ty hiteny gsp eer diined os t turnoeudtt bo efe malTehe.r erfoerdee,s c ription soltvhpeer ob-laelpmla ratwyeprfeeem sa le, ofs evteyrpailens se ictohmeprnl oesrta et isfac­ soa naloyfms ailgsee niitisam lpioas sible. tory. Neatrht ey psee riinIe NsH iSsa t uwbei th Malgee niwtearsleti uadi ginle ydc derraiwn­; lab"e4l6 5(6t8sh"ae ma est ypaent)dh see c ond inwgesrm ea diegn l ycjeerlTilemnyri.n �oyl o-lab"eLle ucaompeirsiM caalnla<o lcJeh. t,R. . accorMdciAnlgp(1 i 9n)8eO.1 T (o cetlnlaanr­ MalloIcctoh n"t.5a m ianleaedns1 d 4fe malIe s. glei)n dreexfet rots h rea toifto hd ei stbaen­ce prepaaltrlhe emdsa ela ensddi scotvhteahrtee yd twepeons toecreialonltrdih d ei stbaentcw ee en beltoLon. gg lyp hiniTvaonraas i(jTtasnhauski j­ anpyo stoecreilaolnrtud h sae n teorn ieo.r tsh1u9k51 89,8 I6it)ts .h erperfoorbeta h balte thtey sppee ciamlebsneosl toont ghs ip�sie e s, Descriptions anLd.g lyphinisvhoobrueaal cdc eapastJ eudn ­ iosry noonfyL m.a meriBcuattnh aei.rns eo Leucoapmiesr iMcaanlal1 o9c 2h1, distbianstciots porrro evjteeh ctiths e sis. DistriIblultiIintfooh nsii.pss e. c iiisde esn ti­ Mall1o9c2h31,5: ;41 94207:2 . cawli Lt.hg lyphi-nneiavcroolrsyam opolitan. Modeo fl ifAet.l eaisnpt a rlta,rfe veaode n Typperse seirn/v NeHdHS o.l otsyppeec i(mwehni cah phiItdfh ssi.ps e ciiisde esn wtiiLtc.gha l ly phi­ wagsl uoenpd a pterri alnogseltme,p)p t iyln a be"l#l ed nivoirtlasa, r feveadoe n n eaornlheyu ndred 46568""L,e ucopis Maaml-eltr.yi pc(eaw"nr aii tnt en penc"iTly)Lp,ee u coapmiesr iMcaalnlado "cr'he,ld a ­specoifae psh ids. _ be"lI NHTySp e#0 122O"n.t hsea mpem a rteh r ee puJaarniadsa m atlulcb oen tatihnaeib ndgo imngeln y c­Leucoaptirsi faAclider1si9 c 2h5, eriTnh.ia sb do(mweint dhoouufrbtot m,h olostpyepcei (­Fi1g-s5 ) meni)fes m alneom,ta laesM alltohcohu ght. Paraty1p esc;,l:,a b"e4l6s5 6"8la"l,o tlyepuec opis ameriMcaalnla<;>o "c",h! NHTSy p0e1#2 32" ;(<;> o neA ldr1i9c2h15,5: 2 . ZOOSYSRTO.SC SAIV o1l1.• V N. TanasijNtsehaurksc:pt eiccoi fLe esu coIp is. 19 5 2 4 3 Fgis1 -L5e.u csoa ptirifaAcliders1i ,2c ,h-.h 3e,ma adl;ge e niitnap lrioafi4 l,ee p;a ndfrroimub me hi5n,ad p;eo xf aedeafrgoumbs e loSwc.a0 l.em1:m . TypeHso.l ot<;>y,pp ree,s eirNnv MeNdHL .a be"lTsy:p eS ta1t2ed' :,7 <;> ,G oldenAduag2l.0e1 95e9m A.p r. #2 82U1S2N M"",S aFnr ancCiaslci"of.E "W.,a lthe1r9 6p0r,e Add.e lpgiecse( aRe.M Gi.t chIed' l,2l <;> ),; col"lI.cn"o ,n newcittThiheo cno dippilnois-irsa ditahsteaa mepe"l ,aacnecd o lle"ccto8ol rl2-,.0- 59"C.a li­ "LeucaotpriifsaA ciidPe"as.r atNyMpNeHs6:: (c;>" al­forniaI:cl ,S aFnr ancJias1nc.9o 2,o8 n,A delsgp e.s lotyrpeeap"la,lr ya atny5dp" ep aratwyipttehhss ea" m)e onP in(uEsW. a lth1e r(9)? B;)a ,l dLw.i6 n5 0S0.' B,. labtehlCesa ;l ifoArnciaad eomfSy c ie(nCcAeSs4) c;> ,, Co.1,3V 195(5W .RM.aMs.o Inc;> ),A;l baAnlya,m eda thsea mlea bels. Co1 5V I19 5a8d,u clotl leocnMt oendt eprien(yeF r ed Othmeart eerxiaamli CnAeSd3:d'. , 2 c;> ,S aFnr an­D.BenneIt<;> t,t) hs;e a mlea bbeulVt,I 1I9 538d' ;,3 cisCcaolI 2. 7 2.5 p,r ebsyE. .W altchoel(rlo o.nn d'e 9,t hsea mpel aacnecd o llePciutnsro ard,i VaI1t 9a5,8 ; penlcaibl"e Lle ucboep/iOls.a S .a?n"ad n otlhaebre l2 <j> ,S aRna faeMla rCion1, 9 V I1 95p8r,e odnP. i neus "LeucaotpriifsaA ciidDe.es tF .. BDe.n n1e9t5t8a "n)d, o nM onteprien(yea slple cidmeetnesr mbiyFn .eDd. I'i' ,t hsea mlea bbeulXt,1 81.9 2C6N.C W:a shi ngtBoennnaesLt .at t rifaMceixeisIc)<jl o.,3: m iE E lS aito, 19 6 V.NTa.n asijNtesahrusckpt:ei ccoi feLse ucIo• pZ iOsO.S YSRTO.S SIVoClA.1 1 DgoM ex8.2 0J0u'l,4y ,1 96(4J .MFc.A lpi5n9 e,1) 0; Leucfloapviisc Aolrdnri1is9c 1h4, miW ElS aitDog,o .9,0 00J'u n5e- 161 96 4(Fi6g-s 8 ) (J.F.McAZlIpN2i:c! n ,Ae l)b;a Cnayfl,A.i l ameCdo a VI1 95o8nP inruasd i(aFtraDe .Bd e nne(trte)c eived fromD rJ.. MFc.A lpiinen xec hanCgSeI)Rl.Oc! :, 1 Aldr1i9c1h44,0: 4 . 9,G oldGeantP ekS .FC.a,l ifoFreneidaoi,nnP ign eus spb.r,e Jdu n1e9 3(8F .GJa.y ). Typesp reseirnNMv NeHd. H olotd'y.pL ea,b e ls: DescriMapltefei,mo anl.Be o.dl ye n1g.t7h- "BrownTsevxiM.la ly1e 0 1,9 10""R;AV .i ckCeorly­ lect"o#5r "4;r;" e lda b"eTly Npoe1 84U2.8S .N .M."; 2.m2m .B odgyr eHye.a( dF i1g-s2o )fc! 1. 5 relda b"eHlo loLteyupceo jlpaivsi coArlndirsi ch''. timaeshs i gahsl onogf9, 1 .(3a verAangtee)­. Paratlyd' p,5e 9 s,t: h sea mpel atciema,en c do llector. ripoaror ffrt o nasn guilpnar ro fiFlreob.na sr elCyNCl:9 ,t hsea mpel atciema,en c do llector. wideannetde irnic! a,2d .;1 -t2i.mn2ea srr o werDescriMapltefei,mo anl.Beo .dl ye n2g.t0h- thhaena idn9, ,2 . 2-t2i.mO4er sb.is ttralilip gehs2t .m6m .B odlyi gghrteH ye.a1 d. 7-t2i.ma0e s s grewyi,d eannetde Orciealpdll.aa ertl eo ntgrait­hei agshl onFgr.os nosm ewwhiadte annitneg r iad; angusllairge,hl telvyFa rtoensdtt.ar alil plseio g h2t.6-t2i.mn7ea sr rtohwahener a Fdr.ou nssu ally greFyr.ow nismt ahn tyh wihni htaeie rssp,e c imaolnloytgorneheyo ;w evmearpy,o sstewsdosa rk nealru nuFlaecl.eu ,n aunlpdea orftg endaa r kg,resyt riflpoewsia nrgo uoncde lpllaaartn ed neabrllayLc ukn.us lheod rotr,bs oarlcd uerr v erdeaclhuinnuWgl heenfr. o nmso notoocnee l,l ar sligOhTti lnyd1.e. xA6 n.t ebnlnaa3c rkad,r istpol­abtaer oeultyl iinnse pde;c iwmietdnhas r k mer4e-t 5i mleosn tghea2rnn dG.e nhai g2h.,4 -fronsttarli appepseo,ab rlso ng-tOrcie alnlgiular. 2.t6i mheisg thheearny h ee igMhitn.ub trei stulneesvsepnalfrcyoe mde acoht hOeTri ,nd veaxr y­ ong enaarr anignre odwa ;n tebrriiolsrat rl gee isntbg,e tw1e.ea5nn 2 d. S0c.a apnepd e dilci eglh t oftseunrr oubnyds emdab lrli sPtallbeplsia. c kb.rowbna,s oflagealnlador miyesertleal roaw­; Mesonowtiutmhs oturtic poevse,br uentdo , t tiboe twleeennog ft3 hra dn 2dn adr issetga­l densbeylr ye,g ulloanrg itruodwoisfsn maal l lmenvtasr frioemd1 .2t:o21 . 5L:u1n.uh lie g h, briessTt.wl op aiorfds o rsocbernits rtadlloe rsesdaglse e mi-roGuennhdaee id4g..h 5t - 5.0 presFeenmtob.rl aaw cikty he llaopwi cteisb ;i ateimleests hsea ynh ee igShetv.eb rrailsfo trl..me s blawcikty he llboaws aelst,h osuogmhe t imienisgrr egruolpwar re soenen atcg he naan;t e rior femoraan tdi beinatei brlealccyok l;o roa ftiborni sltalregPsea sldtpa.irb kr owGnr.em ye d ial tarvsair iea.bgel.ne,t, i brlealocyrfik r,ts atr sos­trsilpiegsdh atrltkyhel arin gg hrteb ya ckg round meroenms i ddalnhedi nldeb glsa acnksd l ig hptrleysoenmn ets onottougmew tihtwehir d lea t­ yellboaws aolrfil ryts,at r soymeelrloeonsaw l lersatlr oifps eism illiagrchlotyl oreaxttieonnd ­ legOsnw. i ngvse,iM n asn Rd4, 5 sligchotnl­yi nbge yomnidd odlfme e sonoAtruebmae.st w een verigdnei s1t/oa3rla rpea ratlvple erslyh. oi r ntm edisatlr idpeevsoo ifbd r isatrlebeaes t;w een compawriidtsihos nst eaclto ifCo urn;a ti2is.o 0 -medainaldla tsetrrawilip tsehms a blrli sotn lleys 2.A2b.d omsiynnatle l+r 2ga iltmeoe sntt i reexltyentdoai nntge 1r/iTw3oo.r p aiorfds o rso­ browtnhc;io sl orsaptrietoatonde sr 3gb iytt we o centbrrailsp trleessFe enmto.br laa acpki, c es waveRsa.t hweirdm ee dilaonn gitsutderi inayaell ltoiwb;yi ealel moiwd,do lfae n tetrii boira presoentn etr g3i-to5er 4s - 5D.a rfike lod fs darketnaeyrdes;li l eoxwc,ae ppittc aarls o meres modimfiiecdr otarbiscfrehonimlta a et eedrga elsd arkeOnnwe idn.g vse,iM n asn Rd par allel 4+5 offi rasbtd omtienragilincl t . es ors omewchoantv enregaairpn igc tep1s. .1 -1.3 Malgee ni(tFail3gi-sa5E )p.a ndmraisusm i vtei,msehso rtthedarin s ptoarlto ifCo uAn.b domi­ extenidnivtneogrs yh ocrutr,ve eddiH tyepsa.n n­asly nte1+r 2dg airtkeae nnteedr tiwosorm l ayl;l drinuema relryeA cetd.e algounswg i,d e nebdlaocrgk o ldsipsohst osm eteivmiedsoe nnt apictaulrnlueypd,G. o nopaobdssep natr,a m etreers3gs,ii tmeit sloam ram leld sipaolot nts e r gites larsgmeo,o tthalpye red. 35-.T hecoafme i nubtrei sotnal bedso minal TaxonnoomtIiencgs e.n isttarlut chtseup ree­t ergeintceisrb cysl meabdlu lnt o ticbelaabclke ciiessc loosnelt yoL .p inipMealrdlaoI ct hss.p oDtasr.fik e lodfms o dimfiiecdr otornichiae determbiyne axttieaornpnap le airssai nmc pel lea;teedrgaoelffis r asbtd omtienragolifm t aelse s coloroaftt hiheoe nai dnc ombinwaittaihbo ­nn oetv ident. senocfse t roinpt ehmsee sonaortveue mrc yh ar­Malgee ni(tFai6lg-isa8E )p.a ndmreiduimu m­ acterBiuistitt ns ie cc.e stsocao rnyst ihdpeeo rs ­sizweidt8,h- 1p2a iorfls a rbgrei satftleneros ­; sibiolfai m tiys tianck oem parwiistdohan r kt­abcloen streixctteiinondnmts,eo d ium -sized, facesdp eciomfLe p.ni sn ipfreordmwa h,i Lc.h ratthheeirdn i Htyepsa.n dwriiduemin nae ndt e­ atrifdaicffioeenrslis ytn h set ruocftt humera e l eripoarrs tl,i gchutrlvAyee dd.e awgiudsei nne d geni talia. baspaalsr tm,o ottahpleytr oiwntaghr sidln i, g htly DistriUbSuA(t Wiaosnh.i Cnagltiofon rn,i rao)u,nadpee dx . Mexi co. TaxonnoomtiLejlc.sa .v iccoarbnned i est er­ Modeo lfif Lea.r vparee daotnAo dresls gp eps.m ineeads biylt yhy ee llcoowl oofut rhb ea so­ onP in us. flagellTohmiecsrh ea.r aicsut neirqa umeo ng ZOOSYRSOTS.CS AIVo l1.1 • V. N. TanasijNtesahrcustkpi:ec c oiLfe esu coI pis. 197 Fig6s-L 8e.u copisjlAalvdirc6iom,cr ahnlgi.ees n iitpnar loifi7aal, ee d;e fraogmbu esl 8o,w ;g onaonppdao rdas m eres froma boSvcea.0l .em1:m . stuNdeiaerdsc pteioccLfi e eusc oOpnti hcse.o n­othPearlbspl.ia L cakt.se trraoilfp meess onotum trasrtyr,u ocfgt eunrieit nsao lctih aa rac tienrciosntsipocirhc aurocduolsny s pimceudo iuasl, andde termoinnolayntt hiciohsna rcaacbnt e ers trfaiipnentsol tyi coenanlbeyala nert eerdi goer. unreliable. Mesonsoutrafaelcv ee nly bcymo ivne urteed DistriTbhusitpsie ocinknis.oe wson n frloy m brisatrlebeaesst, ws eterni poeust lTwnioon te d. Tex as. paiorfds o rsobcrenitsFrtealmleob srl.aaw cikt h Modeof lifuen known. yelalpoiwtc iebsai;nat dea yr eslilO onww i.n gs, Leucompaicsu lTahtoam p1s9o1n0, vtehdaiinnas snpRtMdo4+:a rp ltoa 1frC0 autOnl.1pn l a . eb1.ldt-4;oi1 ( mm sFeehi9nsog) r.,t er (Fi9g-s1 3) synteI+r 2ga iltmeeon stti brreolwryno ;u n ded paisrpeodpt rse soentn etr g3i-ts5eo;sm et imes Thomp1s9o1n20,3: 8 . simillaarorgnlte ye rg3i-mt4ie,ns ou ntt ee r gite 5,b uuts uaelvledyni lmyi npiosshtieOnrng i ad. 'J:yppreess eirNMNHnv .e d Holot9y,"p Cea,lN veebrt. t erg3i-t5e,s t brliaacnskgp uoaltlaspsr ro e sent 10/15"/9,0 Ty9p/e"1#,3 14P1a"r.a tI9y ,pt ehssea: m em ediLaalclkydi.an firgek l odmfso dimfiiecdr o­ labIecl l,t; h sea mlea bpellau,ds d itliaob"neBalrl e d tricohnei dageoe ffis r asbtd omtienragil in tes fromEricoopleolrtaid(seIt nnhds eeis s"c. rTihpotmipo­n, sownr ottheta ytsp eer wiae3ss a cln 1d )9. mal es. Othmeart eerxiaamli CnNeCdC:.a naAdlab:e2 rtcl a,: Male g(eFnii1gt0sa- l1Ei3pa)a .n drium bear­ Scan9dV iI1aI9, 5 (60P .e cMka)n.i tI9o ,Wb ian:n ipieng1 g0p aiorfbs r isotnpl oessts eirdpieao,sr s ing Sept1.9 2e7m,. 131.9I2Xe7 x,g rassacsl (eA . V.smooitnhdtlioys traolulneydd eidHt yepsa.n d­ MitchCe.nHCe.ur rr,a 4ncl ),5;9 ,W pg1.8 .2X7g ,r assr iusmh omrats,s piavsesfr,io nmbg e neianttho coc(cpirdo Ab.aMVbi.lt yc h4ecl n,2e9 r,W) p;gF .e b .29, paramcelroesfuses lewydi itthg; o nocpoomd­s 192(8A .MVi.t chNeMnNeHrI:)n .d i5acl n,1a 9 :,L a FayeVtIt-e9(,-J 1.A5Ml .d rich). plerteedluyc ed.s lAiegbdheetnsaltmgy,ou ost h­ DescriMpatlifeoe,nm .aB loed.ly e n2g.t2h- lyt apetroiwnaagpr edx . 2.mm8. B odlyi gghrteH ye.a1 d.t 6i maeshs i gh TaxononmoitcTe hsse.p ecieisae ssdi ilsyt in­ asl onFgr.os nosm ewdhialtaa ntteedr iadg;u iasviheenarx b­tleearn paple abryta hnpecr ee s ence agi2n.gt6 i mneasr rotwhaenr Ohrebaidt.oa fpl a isrpeoodtn ts e rg3i-wt5ie,ats lh m coosmt­ strliipggehrset Oy c.e lpllaaortu et loinnble yyd pleatbes eonflc aet setrraoilfpm eess on otum. ligghretcyo lorpartoicoetneo,dw ealdru dnsu lGee nisttarulciotcfLu .mr aec uliuasnt ac ommon asn arrtorwi aFnrgolnsett.ar glir peey , dfoarLr ekuecrog poinso;pc oodmsp lreetdeu lcye d. coloFurroedndes.n sceolvyew riectdoh l o urlLe.msasculhaaatsnaa lmcoosmtp laentaelloy­ haiOrTsi .n dex1 .nA5en.at re annnadar i sgtoaucso untientrp haPera tl aeraerpcrteisce nted bla3crakdr, i st5to immeaerlsseo a n2sgn Gde. n abyL .sz epliget1i9i3sA7ofc, arz ree lp,ofr rtoemd hei3g.h1t- 4.l0e tsthsiea mynhee es i gBhrti.st ­hree gfrioomHn u ngtaoMr oyn go(lTiaan asij­ tloengs e nvaa riiannbu lmeba errr,a in1ng- e3dt sh1u9k8,I6i t)ds .i stinfrgouLmi.m sa hceu dl ata irrergouwalsna;tr e rior britsh talnient hnpeor teasboelflny ac telse atrrraogilfemp ree sso notum 19 8 VN.T anasijNtesahrusckpt:ei cocif Le esu coIp• iZ sO.O SYSRTO.SS I CVoAl 1. 1 / 12 Fig9s- 1L3e.u cmoapciusTl ahtopams o9na,. b dom1e0mn,a; lg ee niitnpa rloifi1al1 ae,e; d eianpg ruosfi 1l2ae,e; d eagus fromb elo1w3p,;a ramfreormbe e low.0 .Sm1cm a.l e: (t hohuigghvh alryyii ningn teoncfso oilrt ayt ioDni)s,t ribCuatnia(odAnal. b eMratnai,t oba), narrgoewneaarn, id nd ifferiengn ecneist aUlSiA(c N ebrIansdki aa,n a). stru-cptaurrtei ncaurlrasoruwlreysr,lt oyb leasrM odoelf if Lea.r avrapeer edoanEt roirosp eltis anade deagus. Hoowpfea viesrrpe,od t spscr ae lseesn.ce ona bdomtienrag3li-a t5ne sdsi mibleatrwieteyn basgiecn icthaalriac(c mtaesrsrsio vuen,d Leedu copisM amlalj1oo9c r2h 1, sursltoyblgeaesrn, e raolfa esdheaaapgbeu­ s(,F i1g41s-6 ) senocfge o nopaoridens d)i coafact liovrseee ­ latiobnesthwtiehpee snsp ee cTiherese .l ation­ Mall1o9ch23,15 :2 . shbiept wteheesnsp ee cmiaeaysl b seoj udged byt hfaec tth tahtel iarrp vraeoeyns cafrloems Typesp reserivnINe HdS H.o loty'i'p.Lae b,e "lSst:. Joseph, cerepaalrtsi,co unsl paeroclEfiyr e iso pEevlit­IilsMla..3y ', 1 4""S;aF lotr" kLe"u;c ompaisjM oarlT lype." (in dentthleyss,pe e chiaevoser igirnealtaetdip veenc"liTylyL )pe;eu cmoapjisMo arl l<;,o"(c rehld a be"lNH)S; recefrnotmalc yo mmaonnc easntsdoe rp,a raTtype e#1 d2 4P"ara type3s'i' :,th esa mel abOenlpe.ar aitypse greasoenwdeit, h ohuetadA .l lomtyepnet iboyMan leldao b­ch duteog eograipshoil caatli on. sePnrto.b iastbe lrvfoyer pd r epaoragfte inointa lia. ZOOSYSRTO.S SIVCoA1l 1.• V. N. TanasijNetasrhcustkpi:ec c oi/eLucseo pIi s. 199 16 Fig1s4 -L1e6u.c opmiajsoM ra lhl.1o 4ch,e a1d5;a, b dom1e6mn,a; l gee ni(taaflMtiaealr l och). DescriFpetmiaBoloned.l.y e n2g.t4hm- m2.. 6 TaxononmoitcMe sa.l l(ooccphi. tw .r)o te Bodlyi gghrteH ye.a( dF i1g41.). 4t-i1m.ae 6ss " tyfepmea lael,l oatnyfodpu erfe, m alpea ra­ hiagshl onFgr.od niss twiindcetanlneytd e ritaydp,easnp"dr, e setnhsteke edt dcrhaywo ifn g 2.2t-i2m.ne6as rr otwheharen a Hde.as dm oothmlaylg ee ni(tFai1lg6iA.)ac . c ortdtoih dnirgsa w­ rounidnpe rdo fiFlreow.ni stm ha nlyi grhetl,ai ­nLg.m, aj ohra asr atmhaesrs eipvaen drium, tivleolhnyag i presr,p entdofri ocnuOslr.ab ri tmaolderwaitdeeedl iyat c eusr,hv yepda nda rium, strilpieggshr tes yl,i gwhitdleyan netde rilaagdrea. edeaapgoadle amneadr, a tshmearl l Frontaild esnttlriiicwgpahheletv ln iy e frwoe md aedeaBgututh sdi.rs a wiiisnn ge xtahpcert e;p a­ fronbtu,fat rd arwkheervn i ewfreodm aborvaetu.is ofoenrdt hdirsa wwiannsgo fo tu nidtn h e Ocelpllaanrto reta idsiesdt,i nognyulb i yshIeNdHc So llectiotnhm.eo seBtxe psriedsessi,v e ligchotl orAanttieoornci.eo lsrla umssei az se charafocrtt hseepr esca irteehl sei gfrhotnw si th posteornieOosTr.i nd1e.x4 -L1un.u6la.er c­thhiani rresl,a twiivdgeeel nayan t dh rpeaei rs shapdeodr;ps aawrlit t sha mheas ia rsfr on so,fde (rautnhceorm cmhoanr aMcatlelerov;c e hn proclAinntaebtnlena.aac r ki,s tbar od3warnrd;kw roatbeo "utth arnesdeo metfoiumrde osr so­ arist4o.m0e-tr4ie.m a5esl s o nags2 ndG.e nac entrUanlfosr"t)u.an maotNneegla yr,Lc etuicc­ hei3g.h3tt- i3m.le4ets sh sea ynhe e iMgihntu.t oep isse ebnym eI,h avneofo tu nmdo rsep eci­ bristlesi no narerer gaurnlogaowern dg enam;e nwsi stuhca ch o mbinoafct hiaornaa cntde rs somaen tebrriiosfortrm lientsgi cglhuts wti etrhc,o unlomdta kper eparoafmt ailgoeen nsi talia. onlea rbgreirPs atllbpeli.aL cakt.es rtaroli f pesD istriIblultiinooni. s. mesongootludmi sh bronwanr, r orwMe,oldaoetfl i ifuvene klnyo wn. extentdopoi sntge praioiofrdr eG .r emye d ial strriaptephsae lerex ,t entdmoii dnodgfl mee so­ LeucompiinsMo ar ll1o9c 2h1, notuamr;eb aest wseternui spueadsle lvyoo fid (Fi1g7s- 24) briosnta lnetse1 r/oi4fo mre soTnhroetpeua mi.r s ofd orsocbernitsprtralelesF see nmtob.rl aa ck; Mall1o9c2h31,5: 41 ,9 4207:4 . tibbilaaweci kty he lalpoiwcfr eesq,u eynetll­y loown i nnseurr fatcaery;se il ltowwao,p ical Holotyd'p,pe r,e seirvn!e NdH LSa.b e"lDsBu:o iIsl,l . tarsodmaerrkeeOsnnw eidn.vg esiM,na snR d4+5 Aug9.,' 17""L;e ucmoipnMisoa rlT yple."( ipne nc"iTlyp)e; conviednri gs1et/ at2p1l; . 3-t1i.ml6ee5tss h s a nLeucmoipnisMo arl ld'o"c( hr e"dI)N;THyp Se #0 12 5". dissteaclto ifCo unO. n a bdom(eFni1 g5 .) ,DescriMaplteiB.oo ndl.ye n1g.tm5hm .E n­ synt1e rga+il t2mee o nsttid rae(rlgkyo lwdhiesnht ibroed gyr eaistecsdo ,l, oa ucrcdoirntg o viewfreodma boavned bfrloambc ekh i nMda)l.lloecahd,-H gera(edFy i1.g7 1.). t 8i mae ss Sligohbtllipyqa uiserp eoodtnt s e r3gc iotleo urheidagl sho nFgr.on nowsti deannetde 3rt iiamde,s identPiacmlaeeld lilyao.nn gisttuprdriieans aelnn atr rtohwahener a Odr.b iatnfradol n sttarli pes ont er3g;ibt reim gehdti satnrp iraeeso e nnetq uadlalrSykh .ot rhtbi rni sptrleesosnefr not n s terg4ai nt5de. s anddo rpsaaorlft l uOncuellpell.aac rta er d iform, 200 VN.Ta nasijtNesahruckspt:ei cci ofe Lse ucoIp• iZ sO.O SYSRTO.S SIVoClA1. 1 2 0 ., :-··.·. :- : · t 21 ·...... ......... I 18 23 2 4 Fig1s7 -L2e4u.c ompiinMsoa rl lo1c7hh,e. a 1d8;1, 9 m,a lgee niitnpa rloifiaaln fredo mb el(oawft MecrA lp2i0n,e ); aedeaagnaudps o deimnpe r ofi2l1ea,;e deafrgoumbs e lo2w2s;,u rsltoyblieanpsr r ofi2l3es,;u rsltoyblfreao srm behi2n4gd,;o nopaondpdsa ramfreormbe esl Socwa.0l .em1:m . ZOOSYSRTO.S SIVCoA/l! . IV. N•.Ta nasijtNsehaurksc:pt eiccif leesu cooI pis. 201 sliglhitfaltryeo dua nhdt eorcieolOrlT ui sn.d exTaxononmoitcDe est.e rmionfta htsiipsoe n­ 1.8O.c ellii nse iqzuLeau.ln ualrec -s hacpiebedysd. amatgyepdse p eciwmietsnhh, od ret­ Antenna3 rabdrl iasctk6o;t m iemaresles o nags scriopftM iaolnla oncpdha rdtelsyt rgoeyneid­ 2n.dG enhae i5g.ht2ti mleests hsae nyh ee ightta.li isva e ruyn relAimaobnlmgea .t erfrioaml s Minubtrei sptrleesosen gn etn iar,r egaurl­aCralnyaa dnaUd S Aa rsee vesrpaelcw iietsshi mi­ rangoendle,a rgPeaslbtpl.ia Nceki.t lhaetre rlaaglre nitbautlthi eahy,a vlea taenrmdae ld ial nomre dsitarlvi ipseoisnbg lree amseesdo notsutmr;oi npt ehmsee sono tum. Malldoicndho d ti sctuhcsihssa raicnht ideser ­ DistriIblulti inooni.s . scripMtciAolnpw;ih nsoea ,tw h tey sppee cimeMno dofe l ifuen kno wn. befoprree paorgfa etniiowtnra ol(tiileani , tt ht a.t) "mesonwoittuhdmoo urts ocsetnrtiTwrpoae ls "L.e ucpoapriasMl allell1oa9c 2h1, paiorfds o rsocbernitsprtraleles sFe enmto .r (aFi2g52s,6 ) blawcikty he llaopwi cteisbb;il aaewc ikty he l­ lobwa sfoerse;t aresnit ibrlealficyrk s,1t - 2 Mallo1c9h23,15: 3 . tarsoymeelrleoosnmw i ddalnehd i ntda rsi. Holotypc;>e,p, r eseirn!v NeHdLS a.b e"lMsu:n cIil el,. Wincgrsu mpvleeindno,dst i sceArncicbolred.­ Ju5i,' 14""S;t oCnry." "L;e ucpoapriaslM laelllao ch, intgoM allotchship,sie echsa "st hsep ootns typ(eip"ne nc"iTly)Lp;ee u cpoapriasl Mlaelllaoc;> "c h, secotnedr gsimtaaeln lwd i desleyp artahtee (dr,e" d!)N;HT Sy p#e 012 8". postmearrigooirffin r tsetr ggriaptyue b es cenDte"scriFpetmiaBoleon.d.I y7 . 5mm lonlge ,a d (actually ptohiean rtsresye nfetre1er+ng2ci egt ree Hye.a( dF i2g51.). t4i maeshs i gahsl ong. antde r3g) i.te Frosnlsi gwhitdleaynn etde 3rt iiamndea,sr r ower Malgee nitIra elcieafri.ov mDe rdJ. . 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