© Zoological Institute, St.Petersburg, 2004 Review of North American species of the genus Lymeon Foerster, 1869 (Hymenoptera: Ichneumonidae: Cryptini) D.R. Kasparyan & E. Ruíz Cancino Kasparyan, D.R. & Ruíz-Cancino, E. 2004. Review of North American species of the genus Lymeon Foerster, 1869 (Hymenoptera: Ichneumonidae: Cryptini). Zoosystematica Rossica, 13(1): 53-79. A key to 33 North American species of Lymeon is given. Twenty eight species are record- ed for Mexico; new data on their distribution are given. Nine new species from Mexico are described. New synonymies are established: Lymeon imbecillis (Cresson, 1868), M = L. leucosoma (Cameron, 1886), F, syn. n.; L. patruelis (Cresson, 1873), F = L. rarus Cresson, 1873, M, syn. n. Two species are transferred from Lymeon: Rhinium absolutum (Cresson, 1873), comb. n. and Acerastes accolens (Cresson, 1873), comb. n. D.R. Kasparyan, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St.Petersburg 199034, Russia & División de Estudios de Postgrado e Investi- gación, UAM Agronomía y Ciencias, Universidad Autónoma de Tamaulipas (UAT), Cd.Victoria, Tam. 87149, Mexico. E. Ruíz Cancino, División de Estudios de Postgrado e Investigación, UAM Agronomía y Ciencias, Universidad Autónoma de Tamaulipas (UAT), Cd.Victoria, Tam. 87149, Mexico. Lymeon Foerster is a very large genus with pre- – Propodeum at least basally black (in L. transilis only dominantly Neotropical distribution, from Mex- transverse basal groove and front margin of propo- ico to Argentina; three species (L. cinctiventris deum black) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 Cushman, L. nasutus Porter and L. orbus Say), 3. Abdomen black with white bands . . . . . . . . . . . . . . 4 – Abdomen entirely rufous . . . . . . . . . . . . . . . . . . . . 12 are described from the USA. 4. Mesoscutum rufous; thorax rufous, white on front The types of North American species have been margin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 examined by the senior author. The types of new – Mesoscutum black with two white median stripes. – species and all materials (except if stated other- Hind and middle coxae with a white spot dorsally. wise) are deposited in the Insect Museum, UAT, Hind tibia uniformly rufous. – Fig. 2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26. L. rufotibialis sp. n. Cd. Victoria, Tamaulipas, Mexico. Two species 5. Front wing hyaline with two fuscous bands: one at are transferred from Lymeon: Rhinium absolu- apex and another distad of nervulus and of basal tum (Cresson, 1873), comb. n. and Acerastes ac- vein. – Hind tibia with white basal spot (Figs 1, 3, 5) colens (Cresson, 1873), comb. n. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 The following abbreviations have been used – Front wing without distinct fuscous bands, or only with apical one, or entirely fuscous . . . . . . . . . . . . . 9 for collections containing types of Mexican and 6. Head rufous. – Mesopleurum with subvertical white Neotropical species: AEI – American Entomo- spot anteriorly. Legs rufous; hind tibia white subba- logical Institute, Gainesville, Florida, USA; sally. Tergites 1-3 with apical white band, tergites 4- BMNH – Natural History Museum, London, 5 black, two apical tergites white . . . . . . . . . . . . . . . U.K.; ANSP – Philadelphia Academy of Natural . . . . . . . . . . . . . . . . . . . . . . . . 2. L. adjicialis Cresson Sciences, Philadelphia, USA; USNM – U.S. – Head black and white . . . . . . . . . . . . . . . . . . . . . . . 7 7. Head black (Fig. 5), only frons with white orbits. National Museum of Natural History, Washing- Tergites 3-5 black (Fig. 5). – Hind coxa reddish; hind ton D.C., USA. Other abbreviations: SMSC – tarsus light brownish (Fig. 5) . . . . . . . . . . . . . . . . . . “selva mediana subcaducifolia”, SMSP – “selva . . . . . . . . . . . . . . . . . . . . . . . . 25. L. rufoniger sp. n. mediana subperennifolia”. – Head at least on orbits entirely white (Fig. 3). Tergites In the “Material”, Spanish words are cited like 4-5 black (Figs 1, 3) . . . . . . . . . . . . . . . . . . . . . . . . 8 8. Petiolus reddish with dorsal black spot between spir- in labels. acles (Fig. 3). Hind coxa entirely red; hind tarsus ru- fous (Fig. 3). Pronotum on dorsolateral margin and Key to North American species of Lymeon scutellum entirely reddish . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23. L. rufithorax Cameron 1. Females . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 – Petiolus black with extreme base reddish (Fig. 1). – Males . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 Hind coxa reddish with dorsal white spot and brown- 2. Propodeum red (sometimes on apophyses white). – ish pattern surrounding the spot (Fig. 1); hind tarsus Hind coxa red . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 predominantly white (Fig. 1). Pronotum on dorsola- 54 D.R. Kasparyan & E. Ruíz-Cancino: North American Lymeon (cid:127) ZOOSYST. ROSSICA Vol. 13 teral margin and scutellum dorsally yellowish . . . . . covering tubercles. Hind coxa white, black antero- . . . . . . . . . . . . . . . . . . . . . . . 22. L. rufinotum sp. n. basally and dorsoapically. Hind tibia uniformly yel- 9. Head red; orbits white, interrupted on vertex. Anten- lowish. Petiole black . . . . . . . . . . . . . . . . . . . . . . . .18 na blackish, without white band; scape reddish. Meso- 18. Mesoscutum at lateral lobes mostly smooth, without pleurum with large white central spot. Front wing punctures. Speculum black; mesosternum entirely brownish. – Cuba and Florida . . . . . . . . . . . . . . . . 10 black (Fig. 16). Front wings weakly dusky at apex. – Head black and white. Antenna in female and often Hind femur varied in coloration: usually black and red in male with white band . . . . . . . . . . . . . . . . . . . . . 11 or black and yellow, but sometimes red with yellow 10. Metapleurum entirely red. Abdomen beyond tergite lateral spot; hind trochanter I black with yellow apical 2 predominantly reddish with white dorsal spot at margin (Fig. 17). Hind tarsus white, except for seg- apex of tergite 7 . . . . . . . . . . 5. L. bicinctus Cresson ment 5. Upper valve of ovipositor behind of nodus – Upper division of metapleurum with white spot in its about 5 times as long as high at nodus (Fig. 35). Pro- lower half. Abdominal tergites black, banded with pleurum black or white . . . . . . 1. L. acceptus Cresson white . . . . . . . . . . . . . . 28. L. subflavescens Cresson – Mesoscutum evenly coarsely and closely punctured. 11. Temples black with orbits white only close to vertex Speculum yellow; mesosternum yellow or black with and in their lower 0.3, where white coloration ex- yellow spot just beneath sternaulus. Front wing hya- pands on cheeks. Face white with vertical median line. Hind femur uniformly reddish; hind trochanters black spot; clypeus white, margined with black. White entirely reddish (or partly darkened). Hind tarsus usu- band on tergites 4-6 slightly interrupted dorsally (in ally only with segments 2 and 3 white. Upper valve the type) (Fig. 6) . . . . . . . . . 10. L. ingenuus Cresson of ovipositor behind nodus thin, about 6-7 times as – Temples white (Fig. 4); on head, only median band, long as high at nodus. – Fig. 68 . . . . . . . . . . . . . . . . from upper margin of antennal sockets to occiput, . . . . . . . . . . . . . . . . . . . . . . . 7. L. cinctiventris Cush. and upper part of occiput are black (and partly red- 19. Abdomen predominantly reddish; tergites without ba- dish). White band on tergites 4-5 interrupted dorsal- sal black bands (at most only tergite 3 partly brown- ly (Fig. 4) . . . . . . . . . . . . . . . . . 21. L. rufatus sp. n. ish basally). Tergites 3-6 without white apical band. 12. Head black (except for mandibles). Thorax entirely – Hind coxa predominantly red. Mesoscutum with red (except for white front margin of pronotum, small two yellow median stripes . . . . . . . . . . . . . . . . . . . 20 marks on subtergular ridge and on apophyses). Scape – Abdominal tergites black with apical white or light reddish brown; flagellum with about 4 segments white yellow band, or (in L. sexlineatus) sometimes almost (in the type, antennae missing). Front wing broadly entirely whitish with narrow basal blackish band only fuscous just distad of nervulus and basal vein. Legs on tergites 2 and 3. In L. patruelis, abdomen some- uniformly reddish, only last tarsal segments brown- times almost entirely reddish brown, but tergites 3-6 ish . . . . . . . . . . . . . . . . . . . . . 9. L. imbecillis Cresson with distinct apical white band . . . . . . . . . . . . . . . . 23 – Head red, or red and white . . . . . . . . . . . . . . . . . . . 13 20. Front wing with two brownish bands. Scape, hind 13. Propodeum with two transverse carinae without tu- corner of pronotum, speculum, mesosternum, post- bercles. Clypeus not projecting downward. Front wing scutellum, metapleurum (except posterior white spot) with cloud on basal vein, on areolet and at apex. Pet- black. Tergite 7 white dorsally. Hind coxa entirely iole yellow at base . . . . 15. L. mexicanus Cameron reddish, without white dorsal spot. – Figs 8, 37 . . . . – Propodeum with strong crests at apical transverse ca- . . . . . . . . . . . . . . . . . . . . . 31. L. tinctipennis sp. n. rina. Clypeus strongly projecting downward. Front – Front wing hyaline, without fuscous bands. Scape, wing with a median and subapical lightly infuscate hind corner of pronotum, speculum, mesosternum band. – USA: New Mexico . . . 17. L. nasutus Pratt white; postscutellum white or brownish; metapleu- 14. Mesoscutum entirely black . . . . . . . . . . . . . . . . . . . 15 rum (except on margins) whitish yellow or reddish. – Mesoscutum with one or two white (or yellow) me- Tergite 7 reddish (at most hind margin white). Hind dian marks . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16 coxa red, usually with dorsal white spot . . . . . . . . 21 15. Dorsolateral margin of pronotum medially with broad 21. Mesopleura medially with oblique black band (from conical tooth (see dorsally) projecting laterad. Flag- mesopleural pit to subtegular ridge). White flagellar ellum with 5 black basal segments before white band. segments brownish ventrally; scape reddish beneath. Mesoscutum coarsely punctato-rugose . . . . . . . . . . . Postscutellum white. Propodeum black between apo- . . . . . . . . . . . . . . . . . . Rhinium admonitum Cresson physes; metapleurum whitish yellow. Hind tarsus with – Dorsolateral margin of pronotum without conical tu- basitarsus completely reddish. Flagellar segments 1 bercles. Flagellum with 4 black basal segments be- and 2 combined about 1.3 times as long as maxi- fore white band. Mesoscutum without punctures on mum diameter of eye. – Ovipositor beyond nodus lateral lobes; its posterior half medially with strong strongly compressed (Fig. 38), 1.05 times as long as longitudinal rugae. – Figs 13-15 . . . . . . . . . . . . . . . . segment 2 of hind tarsus . . . 11. L. junctus Cresson . . . . . . . . . . . . . . . . . . . . . . . . . . . 4. L. atrator sp. n. – Mesopleura medially without oblique black band. 16. Mesoscutum with one central pale spot . . . . . . . . . 17 White flagellar segments not darkened ventrally; – Mesoscutum with two (or more) median marks . . 19 scape white beneath. Postscutellum brownish. Pro- 17. Propodeum without tubercles or spines. Scape white podeum reddish between apophyses; metapleurum beneath. Head (except black spot on vertex and oc- reddish. Hind tarsus with basitarsus in apical part ciput), thorax (except for mesoscutum, lateral regions white. Flagellar segments 1 and 2 combined about of scutellum and basal band at propodeum) and legs 1.6-1.7 times as long as maximum diameter of eye. (except tarsal tips) yellowish white . . . . . . . . . . . . . . Groove between metanotum and propodeum shallow . . . . . . . . 6. ?Lymeon [?Ischnus] centralis Cresson and broad, below postscutellum with vertical wrin- – Propodeum with a pair of distinct blunt tubercles kles . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22 (apophyses) sublaterally. Scape black. Median band 22. Pronotum on hind (vertical) margin black, except be- on frons, hind half of vertex and occiput entirely fore tegula. Mesoscutum black, with two median yel- black. Mesopleurum on prepectus, on its upper and low stripes. Propodeum with two sublateral white hind margins black; mesosternum sometimes entire- spots covering apophyses and extending to apex . . . ly black; propodeum black with two large white spots . . . . . . . . . . . . . . . . . . . . . . . . . . 29. L. sulsus Cresson ZOOSYST. ROSSICA Vol. 13 (cid:127) D.R. Kasparyan & E. Ruíz-Cancino: North American Lymeon 55 – Pronotum on hind (vertical) margin widely white. White sublateral spots of propodeum extend from Mesoscutum reddish, with black margins and two basal transverse carina to apex. Ovipositor stout . . . median yellow stripes. Propodeum only on apophys- . . . . . . . . . . . . . . . . . . . . . . . . 30. L. tantillus Cresson es white . . . . . . . . . . . . . . . . 32. L. transilis Cresson – Black band on hind margin of mesopleurum not ex- 23. Hind coxa tricoloured: ventrally red, dorsally white pands below mesopleural pit, except for thin line on with apical black spot (Fig. 33). Mesopleurum whit- mesopleural suture (Fig. 19). Hind coxa rufous with ish yellow (front corner entirely whitish); prepectus white dorsal spot (Fig. 19); petiolus white. White partly black; speculum entirely or partly black (but sublateral spots of propodeum anteriorly not extend black coloration from speculum not extends below to basal transverse carina (Fig. 26) . . . . . . . . . . . . . . mesopleural pit, except for mesopleural suture); . . . . . . . . . . . . . . . . . . . . . 14. L. mandibularis sp. n. mesepimeron whitish. Petiolus black, except for ex- 32. Propodeum red (usually on apophyses white and so- treme base. Ovipositor short, ovipositor sheath 0.4 metimes with light brownish spots at base) . . . . . . 33 times as long as hind tibia. [In the type of L. flavova- – Propodeum predominantly black or at least with two riegatus, ovipositor sheath about half as long as hind basal blackish or dark brown spots . . . . . . . . . . . . 39 tibia. Areolet large (Figs 7, 9)] . . . . . . . . . . . . . . . . 24 33. Abdomen entirely reddish. – Mesoscutum entirely – Hind coxa not as above: black and white, or red, or black. Antenna black with scapus yellowish red. Head red with white, etc. . . . . . . . . . . . . . . . . . . . . . . . . 26 black with clypeus and mandibles partly white. Pro- 24. Apophyses absent (weak crests). Lower part of meta- notum black with broad white band on front margin. pleurum black . . . . . 8. L. flavovariegatus Cameron Prepectus and anterior half of mesopleurum black; – Apophyses present (but not strong). Lower part of posterior part of mesopleurum and metapleurum com- metapleurum not darkened . . . . . . . . . . . . . . . . . . . 25 pletely reddish. Scutellum white dorsally to apex. 25. Mesoscutum yellow with black longitudinal bands Legs uniformly reddish; fore coxae and trochanters (Fig. 21) . . . . . . . . . . . . . 27. L. sexlineatus Cameron white . . . . . . . . . . . . . . . . . . . 9. L. imbecillis Cresson – Mesoscutum black with two median yellow stripes . – Abdomen with black and white marks, sometimes . . . . . . . . . . . . . . . . . . . . . . 33. L. tricoloripes sp. n. partly reddish . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 26. Hind coxa white with black spots (Fig. 34) . . . . . . 27 34. Mesoscutum entirely black. – Antenna black; scape – Hind coxa light reddish (sometimes with white dor- and pedicel white ventrally. Head black with face, sal spot) (Figs 31, 32) . . . . . . . . . . . . . . . . . . . . . . 28 clypeus, mandibles and orbits white. Pronotum black 27. Lateral lobes of mesoscutum almost smooth, without with broad white band on anterior margin. Meso- punctures. Coloration of thorax similar to that in L. pleurum reddish with upper front corner black (Fig. orbus (Fig. 70) . . . . . . . . . . . 16. L. moratus Cresson 45); metanotum reddish brown. Petiole light reddish – Lateral lobes of mesoscutum of male (female un- with a blackish dorsal spot between spiracles; abdom- known) distinctly and rather coarsely punctate . . . . . inal tergites 2-6 black with broad apical white band . . . . . . . . . . . . . . . . . . . . . . . . 12. L. lassatus Cresson on tergites 2 and 3, with whitish or reddish lateral 28. Mesoscutum with yellow stripes widened anteriorly sides on tergite 4, and with tergite 7 entirely white; and hooked back along lateral margin (Fig. 18). Meso- parameres brownish. Hind coxa and femur red, tro- pleurum entirely white, only mesopleural suture chanters brownish, tibia fuscous with subbasal white blackish (Fig. 18). Propodeum usually partly yellow- ring; tarsal segments 2-3(4) white . . . . . . . . . . . . . . ish at base (Figs 24, 25). Second abdominal tergite . . . . . . . . . . . . . . . . . . . . . 23. L. rufithorax Cameron pale in basal 0.15. Hind coxa entirely red . . . . . . . . . – Mesoscutum entirely red (sometimes with weak . . . . . . . . . . . . . . . . . . . . . . . . . 3. L. adultus Cresson brownish marks) . . . . . . . . . . . . . . . . . . . . . . . . . . 35 – Mesoscutum with yellow median stripes not hooked 35. Second tergite entirely black, tergite 4 white basally anteriorly and without sublateral yellow stripes . . 29 (Fig. 46). Hind coxa as in Fig. 63. Thorax entirely 29. Dark basal band on tergites 2-6 usually brownish and red with white propleura and front margin of prono- in its hind part lighter (reddish brown); tergites 5-6 tum . . . . . . . . . . . . . . . . . . . 24. L. rufoalbus sp. n. sometimes entirely reddish brown dorsally. Prepec- – Second tergite black with white apical band. Hind tus black. Mesopleurum with a broad black spot un- coxa differently coloured . . . . . . . . . . . . . . . . . . . . 36 der subtegular ridge, which expands on front half of 36. Mesopleurum with large anteromedian white spot. speculum; mesepimeron yellowish (Fig. 20). Front Scutellum and postscutellum white. – Front margin part of metapleurum black. Hind coxa with weak of pronotum, small spot on its dorsolateral margin, white dorsal spot on posterior margin; hind trochant- subtegular ridge, spot on apophyses whitish. Head er I fuscous; hind tarsus with apical half of 1st seg- white and red . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 37 ment and segments 2 and 3 entirely white. Hind – Mesopleurum without large anteromedian white spot. tibia with subbasal white spot. Scape brownish, some- Scutellum or postscutellum (or both) reddish. – times (in type) narrowly white beneath; pedicel with Tergites 1-3 black with white apical band; tergites white dorsal spot at base . . .20. L. patruelis Cresson 4-6 entirely black, apex of abdomen white. Flagel- – Dark basal band on tergites 2-6 black. Other charac- lum blackish. Hind tibia blackish with basal 0.2 ters not entirely as above . . . . . . . . . . . . . . . . . . . . 30 white; hind basitarsus in apical half and tarsal seg- 30. Propodeum predominantly pale basally and white at ments 2-4 white . . . . . . . . . . . . . . . . . . . . . . . . . . . 38 lateral black stripes; its median black mark narrows 37. Discoidella completely absent. White spots on upper apically and expands to basal transverse carina (Fig. division of metapleurum and in hind corner of its 28). Mesopleurum entirely pale yellowish (except lower division. – Tergite 3 in apical half and tergite 4 small black spots above and beyond subtegular ridge). entirely reddish. Hind tarsus fuscoferruginous. – Hind coxa pale rufous . . . . . 18. L. novatus Cresson Southern Florida and Cuba . . . . . . . . . . . . . . . . . . . . – Propodeum before basal transverse carina, a lateral . . . . . . . . . . . . . . . . . . . . . . . . 5. L. bicinctus Cresson stripe on pleural carina, prepectus and speculum on – Discoidella present. Metapleurum entirely reddish. – mesopleurum black . . . . . . . . . . . . . . . . . . . . . . . . 31 Other characters not as above. – Mexico . . . . . . . . . 31. Mesopleurum broadly and completely black on hind . . . . . . . . . . . . . . . . . . . . . . . . 2. L. adjicialis Cresson margin (to middle coxa) (Fig. 22). Hind coxa and 38. Hind coxa red with dorsoapical dark brown spot (Fig. petiole of 1st abdominal segment entirely rufous. 61). Propleurum white; pronotum reddish beyond the 56 D.R. Kasparyan & E. Ruíz-Cancino: North American Lymeon (cid:127) ZOOSYST. ROSSICA Vol. 13 epomia. Propodeum with the carinae reduced (except distinct punctures. Brachiella long . . . . . . . . . . . . . . for basal transverse carina) . . . . . 21. L. rufatus sp. n. . . . . . . . . . . . . . . . . . . . . . . . . .18. L. novatus Cresson – Hind coxa tricoloured (Fig. 62). Propleura and pro- 47. Tergites 3-7 reddish with black basal third (black ba- notum predominantly black. Propodeum with more sal half in tergite 3). Antenna with flagellar segments or less distinct areola and with both transverse cari- 8-12(13) white (fuscous ventrally). Hind coxa dor- nae distinct . . . . . . . . . . . . . . 22. L. rufinotum sp. n. sally tricoloured: red and white dorsolaterally and 39(32). Mesoscutum and mesosternum black. Orbits with dorsobasal black spot (Fig. 56). Hind tibia uni- white, widely interrupted with black on temple. An- formly reddish brown. Brachiella absent . . . . . . . . . tenna black (including scape). Propleura, scutellum . . . . . . . . . . . . . . . . . . . . . . . . . 11. L. junctus Cresson (except apically), fore and middle coxae and trochant- – Tergites differently coloured; other characters not as ers whitish yellow. Metapleurum yellow with its front above . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 48 and lower part black . . . . . . . . . . . . . . . . . . . . . . . . 40 48. Hind coxa almost entirely black or brown dorsally – Mesoscutum with one or more yellow marks central- (except for extreme apex), whitish laterally and ven- ly; mesosternum rarely black . . . . . . . . . . . . . . . . . .41 trally (Fig. 57). Hind tibia with white subbasal band. 40. Hind coxa dorsally yellow and black, sometimes pre- Propodeum black, except for two apical lateral yel- dominantly black (Fig. 52); hind femur blackish; hind low spots (these sometimes confluent). Mesopleurum tibia yellowish with blackish base and apex (Fig. 48). usually black in upper third (except for subtegular Tergites 4-7 reddish in basal half and black in apical ridge and usually for posterior part of speculum), but half (Fig. 47). Pronotum on dorsolateral margin white. sometimes coloration of mesopleurum lighter. Meta- Lateral lobe of mesoscutum in hind half polished, pleurum usually black on front upper corner. Petiole shiny . . . . . . . . . . . . . . . . . . . . . . 4. L. atrator sp. n. black except ventrally; tergite 4 usually reddish; ter- – Hind coxa and femur predominantly reddish with dor- gite 7 white dorsally . . . . . . 20. L. patruelis Cresson soapical brown spot (Fig. 50); hind tibia blackish with – Hind coxa not so strongly darkened dorsally, usually white subbasal ring. Tergite 3 in apical 0.2 and tergite anteriorly or (and) posteriorly with basal dark spot. 4 entirely reddish; tergites 5-6 entirely black. Prono- Hind tibia without white subbasal band. – Other tum on dorsolateral margin black. Mesoscutum mat, characters not entirely as above . . . . . . . . . . . . . . . 49 with fine indistinct punctures . . . . . . . . . . . . . . . . . . 49. Antenna with about flagellar segments 7-15 white. . . . . . . . . . . . . . . . . . . . . . 13. L. maculipes Cameron Mandibles swollen basally (Fig. 23). Mesoscutum 41. Mesoscutum black with a central whitish yellow spot uniformly densely and distinctly punctate. Apophys- . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 42 es distinct. Hind coxa tricoloured (Figs 58-60) . . . . . – Mesoscutum black with a pair of median yellow . . . . . . . . . . . . . . . . . . . . 14. L. mandibularis sp. n. stripes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 44 – Antenna with flagellum blackish, without white band. 42. Hind coxa entirely reddish or sometimes with dorso- Mandibles basally rather flat. Mesoscutum usually basal brownish spot (Fig. 66). Mesoscutum with cen- without distinct and dense punctures (except for L. tral spot subquadrate and bifurcate anteriorly. Meso- lassatus). Apophyses absent . . . . . . . . . . . . . . . . . . 50 pleurum usually whitish, bordered with black except 50. Mesoscutum rather coarsely and densely uniformly ventrally (Fig. 22) . . . . . . . . 30. L. tantillus Cresson punctate. Propodeum with distinctly bordered areola – Hind coxa dorsally black and white (Figs 51, 53). (as in L. rufinotum – Fig. 67). Scape white ventrally. Mesoscutum with round central spot . . . . . . . . . . . 43 Mesepimeron predominantly or entirely black. Peti- 43. Mesoscutum densely uniformly punctate. Hind coxa ole white dorsally (black laterally). Hind coxa with white and black (Fig. 53, 69) . . . . . . . . . . . . . . . . . . dorsoapical dark spot, white at base (Fig. 54) . . . . . . . . . . . . . . . . . . . . . . . . . . 7. L. cinctiventris Cushman . . . . . . . . . . . . . . . . . . . . . . . . 12. L. lassatus Cresson – Mesoscutum on lateral lobes almost impunctate. Hind – Mesoscutum in posterior 0.6 almost impunctate coxa tricoloured (Fig. 51): black and white dorsally (smoother on lateral lobes). Propodeum without are- and reddish yellow ventrally . . . . . . . . . . . . . . . . . . . ola. Scape reddish or brown ventrally. Mesepimeron . . . . . . . . . . . . . . . . . . . . . . . . . 1. L. acceptus Cresson whitish yellow. Petiole black. Hind coxa differently 44. Hind coxa entirely red or yellowish (Fig. 64) . . . . 45 coloured . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 51 – Hind coxa with black marks or reddish yellow with 51. Areolet large, about as long as portion of second re- white dorsal spot (Figs 54-60, 65) . . . . . . . . . . . . . 47 current vein above the bulla. Orbits interrupted with 45. Orbits white, broadly interrupted with black on tem- black in upper part of temple. Mesoscutum on lateral ple; mesosternum black; mesopleurum black with lobes mat, distinctly granulate. Speculum black; large submedian white spot (Fig. 43). Tergite 4 en- mesopleurum whitish with small black spot under tirely reddish; hind tibia fuscous with subbasal white ridge. Submetapleural carina usually black. Hind coxa spot (Fig. 43) . . . . . . . . . . . . . 25. L. rufoniger sp. n. anteriorly reddish, dorsally whitish at base, blackish – Orbits white, without an interruption on upper part at apex (Fig. 65) . . . . . . . . . 33. L. tricoloripes sp. n. of temple; mesosternum and mesopleurum almost – Areolet smaller, about 0.5-0.7 times as long as por- entirely light yellow. Tergite 4 black with apical white tion of second recurrent vein above the bulla. Orbits band. Hind tibia yellowish, weakly darkened at base not interrupted on temple. Mesoscutum on lateral and distinctly so at apex . . . . . . . . . . . . . . . . . . . . . 46 lobes and in hind half polished, without granulate 46. Antenna without white median band. Hind corner of sculpture. Speculum white; mesopleurum whitish pronotum before tegula yellow; no black spot under with oblique black band from mesopleural pit to up- subtegular ridge. Black spot between apical trans- per front corner (Figs 70, 71). Submetapleural carina verse carina and apex of propodeum broadened to white. Hind coxa with black dorsal band from base apex. Mesoscutum with fine superficial punctures. to apex (Fig. 55). Tergite 4 usually reddish, darkened Brachiella reduced . . . . . . . . . 3. L. adultus Cresson at apex . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 52 – Antenna with white median band interrupted ventral- 52. Hind coxa black and white, usually with an anterior ly. Hind corner of pronotum before tegula black; black and a posterior black basal spots (Fig. 55). – Mexico spot under subtegular ridge. Black spot between api- . . . . . . . . . . . . . . . . . . . . . . . . 16. L. moratus Cresson cal transverse carina and apex of propodeum wider – Hind coxa usually tricoloured, anteriorly reddish, with to transverse carina (Fig. 28). Mesoscutum with dense weak black mark (Fig. 71). – USA . 19. L. orbus Say ZOOSYST. ROSSICA Vol. 13 (cid:127) D.R. Kasparyan & E. Ruíz-Cancino: North American Lymeon 57 1.Lymeon acceptus (Cresson, 1873) 2.Lymeon adjicialis (Cresson, 1873) (Figs 16, 17, 35, 49, 51) Cresson, 1873: 172, F (Hemiteles; type: F, Mexico, Oriza- Cresson, 1873: 154, F [Mesostenus (Mesostenus); lecto- ba); Cameron, 1886: 253; Townes, 1946: 31 (Lyme- typus: F, Mexico, Orizaba]; 1916: 13 (lectotype); on); Townes & Townes, 1966: 88. Townes, 1946: 31 (Christolia); Townes & Townes, 1966: 88 (Lymeon). Material. Mexico. Veracruz: M and F (type) (ANSP). Male (nov.). Coloration rather similar to that Material. Mexico. Nuevo Leon: 1 F, La Estancuela al S of female, but anteromedian white spot on mes- Monterrey, 22.VIII.1986 (E. Ruíz); Tamaulipas: 1 F, Cd Victoria, canyon Novillo, 840-920 m, bosque pino-enci- opleurum larger and scutellum and postscutel- no, 13.III.1988 (J. de León); 1 F, Gomez Farias, 10.III.1999 lum with white marks (in female, only postscutel- (K. Flores); 2 M, Gomez Farias, Los Cedros, 300 m, tr. lum with white mark). Malaise, 13.III-3.IV.1999 (S. Hernández A.); 1 F, same data, Diagnosis. L. adjicialis can easily be distin- but 31.VII-7.VIII.1999; 4 M, Gomez Farias, Alta Cima, 940 guished from other species of Lymeon with red m, 27.III-5.VI.1999 (S. Hernández A.); Veracruz: 1 F (lec- totype) (ANSP); 1 F, Tuxpan, Juana Moza, citricos, thorax, dark marks on fore wing, and white spot 13.X.1995 (R. Herrera); Yucatán: 1 F, 9 km N Teya Pueb- at base of hind tibia, by the red head (with entirely lo, Selva Baja Caducifolia, 28.X.1999 (D. Burgos). white orbits) and large subvertical white spot on Male (nov.). Fore wing 4-7 mm long; flagellum mesopleurum; tergites 4-6 black, 7-8 white. 24-27-segmented, segments (11)12-17(18) with a linear tyloid on each. Antennae entirely black, ex- 3.Lymeon adultus (Cresson, 1873) cept for small yellow mark on scape beneath. Col- (Figs 18, 24, 25, 31) oration of body similar to that in female, but clypeus not bordered with black and mandibles (except for teeth) white, whitish spots on apophy- Cresson, 1873: 173, F (Hemiteles; type: F, Mexico: Oriza- ses confluent and extend to hind margin of propo- ba); Cameron, 1886: 254; Townes, 1946: 32 (Lyme- on); Townes & Townes, 1966: 88. deum. Fore and middle coxae and their trochant- ers I white; legs beyond the trochanters pale ru- Material. Mexico. Veracruz: type (ANSP); Tamauli- fous; tarsi with segment 5 blackish; middle tarsus pas: 1 F, Hera, 44 km S Cd Victoria, matorral, Trampa with segments 1-4 light brownish, pale at extreme Malaise, 18-25.XI.2000 (Kasparyan); 3 F, Gomez Fari- base and apex; segments 3 and 4 sometimes whit- as, Altas Cimas, 900 m, 1.XI.1990 (H. Ruíz C.); 1 F, ish. Hind coxa tricoloured (Figs 49, 51): reddish Gomez Farias, Los Cedros, 300 m, Tr. amarilla, 26.II.1995 (S. Niño, J. Coronado, S. Hernández). ventrally and posteriorly, black and white dorsal- Diagnosis. L. adultus is easily recognizable by ly; hind femur red with blackish base and apex; the yellow spot on mesoscutum hooked anterior- hind tibia black with pale subbasal spot (some- ly, strong reduction of black pattern on meso- times very weak); hind tarsus white with base of pleurum (Fig. 18), and yellow spots at base of segment 1 and apex of segment 5 black. propodeum (Fig. 24, 25). Hind coxa reddish; hind The male can easily be distinguished from oth- tibia and tarsal segments 1-2 (and base of seg- er males of Lymeon by the central white spot of ment 3) uniformly yellowish; segments 3-5 dark mesoscutum and by coloration of mesopleurum brown. First abdominal segment white with fus- (Fig. 16) and hind leg (Fig. 49). The male of L. cous spot between spiracles; second tergite whit- acceptus differs from that of L. cinctiventris in ish basally (before wide subbasal black band). the almost impunctate lateral lobes of mesoscu- Scape blackish or reddish ventrally. Pterostigma tum, black speculum and mesosternum, and tri- pale. In the type, flagellar segments 1 and 2 com- coloured hind coxae. It differs from that of L. at- bined 1.45 times as long as maximum diameter rator in the completely white orbits (in L. atrator, of eye; proportion of segments 2, 3, 5 of hind orbits at vertex interrupted with black), predomi- tarsus 3.3 : 2 : 2.6; ovipositor tip beyond the nantly red hind femur, black tibia, and abdominal nodus 0.8 times as long as segment 2 of hind tergites 1-7 black with whitish yellow apical part tarsus. Lateral lobes of mesoscutum mat, with (in males of L. atrator, segments 4-7 reddish in very fine dense superficial punctures. basal half and black apically – Fig. 47). Note. In the type of L. adultus and some other Diagnosis. L. acceptus can easily be separated specimens in the material examined, there are two from other Mexican species of Lymeon with a anterolateral yellow spots on central lobes of central white spot on mesoscutum by the black mesoscutum. This character and basal pale band speculum and mesosternum (Fig. 16) and smooth of tergite 2, pale coloration of mesopleurum, yel- and almost impunctate lateral lobes of mesoscu- low spots on propodeum before spiracles resem- tum; tip of upper valve of ovipositor distad of ble characters of L. sexlineatus, but in L. adultus nodus shorter than in related species (Fig. 35). the hind coxa is reddish (tricoloured in L. sexlin- Hind femur varies in coloration but usually red eatus) and yellow spots at base of propodeum and black or black and yellow (Fig. 17); hind tar- are wider. sus almost completely white. 58 D.R. Kasparyan & E. Ruíz-Cancino: North American Lymeon (cid:127) ZOOSYST. ROSSICA Vol. 13 1 2 3 Figs 1-3. Lymeon, female, colour pattern of body and hind leg. 1, L. rufinotum sp. n., 2, L. rufotibialis sp. n., 3, L. rufithorax Cameron. 4.Lymeon atrator sp. n. Description. Female (holotype). Fore wing (Figs 13-15, 36, 47, 48, 52) about 8.7 mm long. Antenna with 26 flagellar segments; flagellar segments 1 and 2 subequal Holotype. F, Mexico, Tamaulipas, Gomez Farias, Alta and combined 1.5 times as long as maximum di- Cimas, 900 m, tr. Malaise, 12-19.VIII.2000 (Kasparyan). ameter of eye; segments 5-9 weakly compressed; Paratypes. Mexico. Tamaulipas: 1 M, Gomez Farias, segment 7 about 1.1 times as long as wide and Alta Cimas, 900 m, tr. Malaise, 29.V-5.VI.1999 (S. 1.5 times as wide as segment 1in its middle. Malar Hernández); 3 M, same locality, 1-8.V and 4-26.VIII.2000 (Kasparyan); 3 M, Gomez Farias, Los Cedros, 300 m, tr. space 0.7 times as long as basal width of mandi- Malaise, 13-20.II and 17.IV-1.V.1999 (S. Hernández). ble. Central part of face transversely punctato- ZOOSYST. ROSSICA Vol. 13 (cid:127) D.R. Kasparyan & E. Ruíz-Cancino: North American Lymeon 59 rugose; clypeus with large punctures separated coxa black and white (Fig. 14); hind trochanters by 1-2 their diameters. Temples strongly nar- and femur predominantly black (Fig. 15); poste- rowed beyond eyes and strongly broadened to rior yellow spot of femur smaller than anterior mandibles (Fig. 13); in profile, their maximum one. Hind tibia yellowish, slightly black basally length about 2.5 times the minimum length (at (Fig. 15); hind spurs yellowish; tarsal segments level of upper 0.25 of eye). Occipital carina 1-4 almost completely yellowish white, segment straight and sharp before connection with hypo- 5 reddish brown in basal half and brown in api- stomal carina; oral carina just beyond mandible cal half. Tergite 2 black with apical third yellow; weakly raised as low lobe. Epomia vertical, weak, tergites 3-7 dorsally black about in basal half and obscured by parallel vertical wrinkles. Notauli yellow in apical half; epipleura of tergites 3-8 not deep, with coarse transverse rugae. Front part entirely yellow; tergite 8 yellow, black dorsally of mesoscutum mat, with distinct punctures be- to hind margin and reddish yellow basolaterally. tween notauli; lateral lobes of mesoscutum mat, Sternites yellow; sternites 2-5 with black basola- smooth, impunctate (except anteriorly); hind half teral spots. of mesoscutum shiny, strongly longitudinally stri- Male. Fore wing about 8 mm long. Antenna ate. Mesopleurum coarsely and densely longitu- with about 28 flagellar segments; flagellar seg- dinally striate; speculum polished, large. Scutel- ments (13)14 to 17 with a linear tyloid. Habitus lum and postscutellum smooth and almost pol- much slenderer than in female; hind femur 6.8- ished. Metapleurum coarsely punctato-rugose. 7.5 times as long as wide; proportion of segments Propodeum before basal transverse carina with 1-5 of hind tarsus 8 : 3.5 : 2.1 : 1.1 : 1.5, 2nd superficial rugosity, partly smooth, polished on segment 2.9 times as long as wide. The main di- anterior margin, coarsely rugose beyond trans- agnostic characters of coloration (orbits interrupt- verse carina, smooth around apophyses. Apophy- ed with black on vertex, mesoscutum entirely ses rather strong, subconical (Fig. 14). Areolet black) same as in female. Other characters of rather small, half as long (high) as second recur- coloration of head and thorax also similar to those rent vein above the bulla; nervulus strongly an- in female (Figs 13, 14), but flagellum black, with- tefurcal; postnervulus intercepted at upper 0.38; out white band; face, clypeus (except for brown- nervellus in hind wing intercepted at lower 0.27; ish lower margin), cheeks mandibles, and pro- brachiella distinct and long. Hind femur 5.1 times pleurae light yellow. Propodeal apophyses short- as long as wide; proportion of segments 1-5 of er, and yellow spots surrounding apophyses hind tarsus 8 : 3 : 1.7 : 0.5 : 2.4. Second tergite smaller. Tergites 1-3 black with apical 0.2 yel- 1.2 times as long as wide, mat, evenly granulate, lowish; tergites 4-7 reddish in basal half and black with very short, inconspicuous, scarce hairs. Tip in apical half (Fig. 47). Fore and middle coxae of upper valve of ovipositor beyond the nodus and trochanters entirely white; anterior basal about 0.8 times as long as segment 2 of hind tar- black spot on hind coxa confluent with dorsoapi- sus and 4.5 times as long as height on nodus (Fig. cal one; hind trochanters black with ventrobasal 36). Ovipositor sheath 0.78 times as long as hind yellow spot on trochanter II. Hind femur black, tibia. at extreme base reddish yellow; hind tibia yel- Body black with whitish yellow spots and lowish with blackish basal 0.15 and apical 0.35 bands. Antenae black; flagellar segments 5-10 (Fig. 48); hind tarsus white with apex of segment completely and segment 11 dorsally white. Head 5 black. predominantly whitish yellow with wide median Diagnosis. L. atrator is the only species known stripe on frons, margins of clypeus and mouth, from Mexico with mesoscutum entirely black malar space, mandibles (except yellow basal (sometimes small sublateral marks or in male a spot), vertex almost entirely (except partly yel- pair of small central obliterated yellow stripes low on orbits anteriorly), hind margin of temple, can be distinguished). L. atratus most closely and occiput entirely black. Head, thorax, hind leg resembles in coloration (except for mesoscutum) and base of abdomen marked with black and L. acceptus, from which it differs in the white white (or whitish yellow) as in Figs 13-15; meso- orbits on vertex interrupted with black (Fig. 13); scutum entirely black centrally, with minute yel- it also tends to have a slightly longer tip of ovi- low sublateral spot just before the tegula; hind positor (Fig. 36). tegulae black with yellow margin. Fore wing hy- aline, weakly infuscate on distal margin. Pter- 5.Lymeon bicinctus (Cresson, 1865) ostigma blackish brown. Fore and middle coxae white with minute dorsoapical brownish spot, Cresson, 1865: 24, F [Hemiteles; type: F, Cuba (Muse- their trochanters whitish with blackish spots; fore um Havana)]; Townes & Townes, 1966: 89 (Lymeon; and middle legs beyond the trochanters reddish = leiponeuron; bibl.). – leiponeuron Townes, 1962: yellow, femora black posteriorly, tarsi brownish 397, F, M (descr.; figs 333 f, g) [type: M, USA, Florida dorsally, their segment 5 entirely brown. Hind (?AEI, ?USNM)]. 60 D.R. Kasparyan & E. Ruíz-Cancino: North American Lymeon (cid:127) ZOOSYST. ROSSICA Vol. 13 4 5 6 7 8 9 10 11 12 Figs 4-12. Lymeon, female. 4, L. rufatus sp. n., colour pattern of body; 5, L. rufoniger sp. n., colour pattern of body and hind leg; 6, L. ingenuus Cresson, colour pattern of abdomen, dorsal view; 7-10, areolet and second recurrent vein: 7, L. sexlineatus Cameron; 8, L. tinctipennis sp. n.; 9, L. tricoloripes sp. n.; 10, L. mandibularis sp. n.; 11, 12, posterior part of hind wing with nervellus: 11, L. rufoniger sp. n. (brachiella reduced); 12, L. rufotibialis sp. n. (brachiella lacking). ZOOSYST. ROSSICA Vol. 13 (cid:127) D.R. Kasparyan & E. Ruíz-Cancino: North American Lymeon 61 Diagnosis. This species is closely related to L. Description. Female (type). Fore wing 6.6 mm subflavescens; both species differ from other long. Antenna with 26 flagellar segments; seg- North American species in having antennae of ments 1 and 2 combined 1.52 times as long as female without white ring, fore wing uniformly maximum diameter of eye. Malar space about brownish, discoidella in hind wing absent, head 0.75 times as long as basal width of mandible. and thorax predominantly red. Lower margin of clypeus impressed and sharp. Remarks. This species is distributed in Cuba Mesoscutum granulate, without punctures on lat- and Florida (USA). Head and thorax predomi- eral lobes. Mesopleurum mat, granulate, rather nantly reddish with white marks, tergites 1 and 2 densely punctured, without rugae. Speculum predominantly blackish with apical white band; large and shiny. Scutellum mat, very finely gran- (3)4-8th tergites of female reddish (weak white ulate. Propodeum without apophyses. Areolet dorsal marks on tergite 7); coloration of male large, 1.3 times as high as 2nd recurrent vein tergites similar to that in L. rufoniger (Fig. 43); above the bulla. Ovipositor rather thin. Oviposi- hind tarsus of male brownish red (in male of L. tor sheath 0.51 times as long as hind tibia. rufoniger, predominantly white). Antennae of Antenna fuscous with flagellar segments 5- male and female without white band; lateral lobe 11(12) white. Head whitish yellow; clypeal fovea, of mesoscutum polished, with very sparse small spot between antennal sockets, broad longitudi- punctures; discoidella in hind wing absent. nal band on frons, hind 0.4 of vertex and of up- per half of temple, upper half of occiput black. 6.? Lymeon [or ?Ischnus] centralis (Cresson, Prothorax yellow with black on prosternum, at 1873) base of propleura, and above transverse groove of pronotum. Mesoscutum black (except for yel- Cresson, 1873: 174, F [Hemiteles, type: F, Mexico, Oriza- low 1! submedian stripe laterad of notaulus and ba]; Townes, 1946: 34 (Lymeon); Townes & Townes, very weak marks on prescutellar carina); surface 1966: 89 (bibl.). around yellow scutellum black. Mesopleurum yellow (including entirely yellow front corner); Material. Mexico. Veracruz: type (ANSP); Tamauli- prepectus completely, speculum, sutures above pas: several localities (Mus. UAT). Remarks. This species has unclear taxonomic it and below (along pale epimeron), sternaulus position. It has a large areolet, propodeum with- and small spot in hind lower corner of meso- out apophyses and some other characters of the pleurum black. Mesosternum yellow with black subtribe Cryptina and maybe belongs to Ischnus. median posterior spot; hind margin of postpec- tus black. Metapleurum entirely yellow on upper 7.Lymeon cinctiventris (Cushman, 1929) division; its lower division yellow with black on (Figs 53, 68, 69) front suture and on lower part (widely posterior- ly). Propodeum black with two broad yellow spots Cushman, 1929: 32 (Diapetimorpha; nom. nov. pro Mes- extending from basal transverse carina to hind ostenus laticinctus Cresson); Townes & Townes, 1962: margin of propodeum. First sternite black, but 39 (Lymeon; key, description; south-east of USA and yellow at extreme base and beyond spiracles. Texas; bibl.); Porter, 1977: 83. – laticinctus Cresson, 1878: 208, F (Mesostenus; type: F, USA, Louisiana), Tergites 2-5 black in basal half and pale yellow nom. praeocc., non Walker, 1874. in apical half; second tergite with basal narrow yellow band, which is connected by a yellow line Material. USA: F (type of laticinctus) (ANSP). Mex- with thyridia; tergites 6-7 yellow in visible part; ico, Tamaulipas: 2 F, Cd Victoria, jardin, 3.VI.1984 and tergite 8 dorsally brownish with yellow hind 10.VI.1985 (E. and H. Ruíz); 2 F, 14 M, same data, but margin and entirely yellow laterally. Sternites 31.XI-2.XII.1986 and 20-25.IX.1987; 7 F, 1 M, Gomez yellow. Fore and middle coxae whitish; hind coxa Farias, Los Cedros, 300 m, tr. Malaise, 7.I-7.VIII.1999 (S. Hernández); 7 F, 9 M, Gomez Farias, Alta Cima, tricoloured (reddish with dorsobasal white spot 940 m, tr. Malaise, 24.VI-21.III.1999 (S. Hernández). and dorsoapical black spot – Fig. 33). Trochant- Diagnosis. L. cinctiventris can easily be dis- ers partly brownish, hind trochanters red. Legs tinguished by the presence of a central white spot beyond trochanters pale rufous; hind tarsus pale on mesoscutum and by distinct and dense punc- yellow with basal half of segment 1 pale rufous; tation of mesoscutum; the body and hind coxae apical segments absent; last tarsal segment of are black with white marks. other tarsi brown. Male: unknown. 8.Lymeon flavovariegatus (Cameron, 1886) Diagnosis. L. flavovariegatus most closely re- (Fig. 33) sembles L. sexlineatus and L. tricoloripes, from which it differs in the propodeum without apo- Cameron, 1886: 254, F (Hemiteles; type: F, Guatemala, physes and darker coloration (clypeal foveae and San Jeronimo); Townes & Townes, 1966: 90 (Lymeon). lower part of metapleurum blackish in L. flavo- Material. Guatemala: type (BMNH). variegatus). 62 D.R. Kasparyan & E. Ruíz-Cancino: North American Lymeon (cid:127) ZOOSYST. ROSSICA Vol. 13 13 14 15 16 17 18 19 20 21 22 Figs 13-22. Lymeon, female, colour pattern of different body parts, lateral view (except for Fig. 21). 13-15, L. atrator sp. n. (13, head; 14, thorax and base of abdomen; 15, part of hind leg); 16, 17, L. acceptus Cresson (16, hind part of pronotum and mesopleurum; 17, hind leg); 18, L. adultus Cresson, thorax; 19, L. mandibularis sp. n., thorax; 20, L. patruelis Cresson, thorax; 21, L. sexlineatus Cameron (holotype), mesonotum, dorsal view; 22, L. tantillus Cresson, hind part of pronotum and mesopleurum.