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Diffusion – Publications scientifiques Muséum national d’Histoire naturelle CP 41 – 57 rue Cuvier F-75231 Paris cedex 05 (France) Tél. : 33 (0)1 40 79 48 05 / Fax : 33 (0)1 40 79 38 40 [email protected] / http://sciencepress.mnhn.fr © Publications scientifiques du Muséum national d’Histoire naturelle, Paris, 2018 ISSN (imprimé / print) : 1280-9659/ ISSN (électronique / electronic) : 1638-9395 Presence of the foraminifer Chapmanina gassinensis Silvestri, 1931, in the Eocene (Lutetian) of the Grignon “falunière” (Yvelines, Paris Basin). The genus Chapmanina, its species and world distribution Armelle POIGNANT 5 rue Léon Bloy, F-92260 Fontenay-aux-Roses (France) [email protected] Submitted on 6 July 2017 | accepted on 3 April 2018 | published on 30 August 2018 urn:lsid:zoobank.org:pub:1336C880-D09D-4C20-8BB1-52B58B378192 Poignant A. 2018. — Presence of the foraminifer Chapmanina gassinensis Silvestri, 1931, in the Eocene (Lutetian) of the Grignon “falunière” (Yvelines, Paris Basin). The genus Chapmanina, its species and world distribution. Geodiversitas 40 (17): 461-470. https://doi.org/10.5252/geodiversitas2018v40a17. http://geodiversitas.com/40/17 ABSTRACT KEY WORDS Chapmanina gassinensis (foraminifer) has been observed as it seems for the first time in the Lutetian Foraminifer, Eocene, of the “falunière” of Grignon (Yvelines, southwestern Paris Basin). It is widespread in the Eocene of Oligocene, western Europe, also present in the Tethysian domain l.s. and even across the Atlantic. In the Oli- Paris Basin, gocene, it becomes very rare and has been only found in some places of Mediterranean Europe and palaeobiogeographical distribution. disappears in the Miocene. The Paris Basin seems to be its northernmost occurrence. RÉSUMÉ Présence du foraminifère Chapmanina gassinensis Silvestri, 1931, dans l’Éocène (Lutétien) de la falu- nière de Grignon (Yvelines, Bassin Parisien). Le genre Chapmanina, ses espèces et sa répartition mondiale. MOTS CLÉS Chapmanina gassinensis (foraminifère) a été observée pour la première fois, semble-t-il, dans le Luté- Foraminifère, Éocène, tien de la falunière de Grignon (Yvelines, sud-ouest du Bassin de Paris). Elle est largement répandue Oligocène, dans l’Éocène de l’Europe occidentale, présente aussi dans le domaine téthysien l.s. et jusque dans le Bassin de Paris, continent américain. À l’Oligocène, elle ne s’observe que dans quelques points de l’Europe méditer- distribution paléobiogéographique. ranéenne et disparaît ensuite. Le Bassin de Paris apparaît comme sa localisation la plus septentrionale. 461 GEODIVERSITAS • 2018 • 40 (17) © Publications scientifiques du Muséum national d’Histoire naturelle, Paris. www.geodiversitas.com Poignant A. fig. 1. — Chapmanina gassinensis Silvestri, 1931 (Lutetian, Grignon falunière, Yvelines), in lateral (A) and apical (B) views (MNHN.F.F62410). Scale bars: 200 µm. INTRODUCTION associated can be up to 4 mm in diameter, Chapmanina gas- sinensis can’t exceed 1 mm. According Smout (1954) opin- Several samples of the Lutetian marl-pit of Grignon (western ion: “Chapmanina has a very small test compared to most Paris Basin) have been examined for a research of miliolids. complex species”. In one of these samples (probably from unit 4, see Fig. 2), I found two specimens of Chapmanina gassinensis Silvestri, 1931 (MNHN.F.F62410; Figs 1, 2). This foraminifer is easy THE GENUS CHAPMANINA AND ITS SPECIES to identify and can’t be confused with any other one. It is well known in the Eocene of France (Aquitaine Basin) and also According Loeblich & Tappan systematics (1987), the genus observed in the Eocene of various Tethysian European countries Chapmanina belongs to the family Chapmaninidae Thal- and of the Tethysian domain l.s. It is only pointed out in the mann, 1938, superfamily Rotaliacea Ehrenberg, 1839 and Oligocene in France and a few European countries. Until now, its type species is Chapmanina gassinensis, because pointed Ch. gassinensis had never been mentionned in the Paris Basin. out at Gassino, in Torino area (Silvestri 1905a). The genus Sometimes called Chapmanina sp. or cf. gassinensis, it is very Chapmanina first appears with the name Chapmania described probably Chapmanina gassinensis because of the typical associ- by Silvestri & Prever (in Silvestri 1904). In 1931, Silvestri ated microfauna. A new species of Chapmanina Silvestri, 1931 changed this name in Chapmanina, two species having been has been described in the Paleocene of Australia (Quilty & described with the name Chapmania: Packham 2006), it will be treated further. Chapmania galea Silvestri, 1923, type species of the genus Before studying the genus and its species, it is useful to give Preverina Frizzell, 1949, considered by Loeblich & Tappan some precisions: one of the aims of this work is to show the (1987) as a synonymous of Chapmanina, species of the italian wide Tethyan distribution of Chapmanina gassinensis, but it Tortonian (surroundings of Reggio d’Emilia), represented only is probably not an extensive recense of all the localities where by an axial section, but the holotype in unknown (Cita & this species has been found; in the same way, all the references Scipolo 1961), as well as the topotypes. concerning the localities of its presence in a particular country Chapmania sertata Silvestri, 1929, from the Lutetian of have not been reported, when they are too many, such as is Ancona, Italia, illustrated by a bad section (Frizzell [1949] the case of the Aquitaine in France or of Turkey. did not think it is a Chapmanina). Moreover, it is important to remark that Chapmanina gas- Quilty & Packham (2006) described Chapmanina conjuncta, sinensis has not always been considered as larger foraminifera, in the Australian Paleocene (see further). and so, it has probably been omitted several times in the In 1918, Halkyard described Patellina conica which is in literature dealing with large benthic assemblages containing fact Chapmanina gassinensis. for example: Nummulites, Alveolina, Discocyclina, Pellatispira, Loeblich & Tappan (1987: pl. 775, figs 1-9), reported Fabiania cassis (Oppenheim, 1896) with which it is frequently the genus Chapmanina from the Lutetian until the mid- 462 GEODIVERSITAS • 2018 • 40 (17) The foraminifer Chapmanina gassinensis in the Grignon “falunière” (Yvelines, Paris Basin). The species of the genus Chapmanina nits ntial u e nits ub- equnits U S Su a Batillaria and milliolids limestone (0.1 m) 1 A10 b Lucinids and miliolids limestone, erosive base (0.2 m) a Seraphs limestone (0.5 m) 2 b Coral and mollusc fine limestone (0.1 m) c nt Seraphs limestone (0.5 m) o a A9 estern fr Lwaitmh iOnarbteitdo laitneds ,p dairffteiarleyn ltiitahli fcieodm lpimaecstitoonn e(0.6 m) 3 w Laminated and partialy lithified limestone with rare b glauconite grains, differential compaction (0.6 m) Milliolids and Orbitolites calcareous sand with a rare glauconite grains (1 m) Milliolids and molluscs (Chama) calcareous sand with 4 b A8 rare glauconite grains (0.3 m) nt o n fr Milliolids and Orbitolites calcareous sand with c easter rcahraen gnlealuincgo nfiiltlein gg raatin tsh.e H bigashely (1b iotoc l1a.s5t imc,) N A TI E Glauconitic calcareous sand, T a U highly bioturbated (0.6 to 0.9 m) L E 5 L D D A7 Glauconitic with bioclasts and grains of quartz MI b filling micro-channels, Campanile giganteum, lithified agregates (0.8 m) Glauconitic calcareous sand, a large quartz grains, oblique bedding (0.4m) b Glauconitic calcareous sand, 6 large quartz grains, bioturbated formations (0.4m) c Glauconitic calcareous sand, large quartz grains, flint flakes, oblique bedding, partially lithified (0.7 m) a Calcareous sand with traces of quartz n pit and glauconite, barely no macrofossil (0.4 m) b her ort Calcareous sand with traces of quartz, micropebbles, n more or less lithified, very fossiliferous, turitellid c fragments (0.5 m) 7 A6 d Calcareous sand with traces of quartz and glauconite, bioturbated on its base (0.5 m) Calcareous sand with traces of quartz and glauconite, e more or less lithified, localy bioturbated (0.4 m) Calcareous sand with traces of quartz f and glauconite, very fosiliferous, bioturbated, bioclastic, molluscs (Cardium subporulosum) (0.6 m) m Calcareous sand with traces of quartz 8 a 1 and glauconite, more or less lithified, molluscs (Cardium subporulosum) (0.9 m) fig. 2. — The “Falunière” of Grignon section, after Guernet et al. (2012) for the section profile, lithologic units and sub-units and descriptions, Gély (1996) for the sequential units and Huyghe et al. (2012) for the correlation of sequential and lithologic units. Section profile modified from Sanders et al. 2015. 463 GEODIVERSITAS • 2018 • 40 (17) Poignant A. GERMANY CZECH rep. 22 this SLOVAKIA work FRANCE AUSTRIA 74 38 8 17-42 SLOVENIA HUNGARY 30 12-77 1-64 53 65-66 67-68 CROATIA ROMANIA 80 67 7-70 SPAIN 18-28 1-64 30 47-66 ITALY 40-52 ALBANIA 26 4-54 63 47 57 5 30 10 GGRreEeECcEe 30 13-30 30 36-79 MOROCCO ALGERIA 13 3 TUNISIA 200 km fig. 3. — Distribution of Chapmanina gassinensis Silvestri, 1931 in the Eocene of Europe and North Africa. The numbers correspond to the references indicated in the Appendix 1. GERMANY CZECH rep. SLOVAKIA FRANCE AUSTRIA 81 SLOVENIA HUNGARY 14-16 20-24 53 55 41 CROATIA ROMANIA SPAIN ITALY 33 82 ALBANIA ? GREECE 25 MOROCCO ALGERIA TUNISIA 200 km fig. 4. — Distribution of Chapmanina gassinensis Silvestri, 1931 in the Oligocene of Europe. The numbers correspond to the references indicated in the Appendix 1. 464 GEODIVERSITAS • 2018 • 40 (17) The foraminifer Chapmanina gassinensis in the Grignon “falunière” (Yvelines, Paris Basin). The species of the genus Chapmanina TURKEY 6 51 2 34 71-72 30 31 15 30 SYRIA IRAK 30 59 45 BANGLADESH 48 INDIA OMAN 27 1000 km TANZANIA fig. 5. — Distribution of Chapmanina Silvestri, 1931 in the Eocene of Asia. The numbers correspond to the references indicated in the Appendix 1. dle Miocene in Italia, France, Spain, Greece, Romania. samples have been studied. Le Calvez (1970) did not report But, I haven’t found any available mentions concerning it among the numerous listed species, but as it is very rare, the Miocene, the species galea Silvestri, 1923, being quite it is not surprising. Along my numerous examinations of dubious (see earlier). Grignon sediment, I also met one Hantkenina and Rober- Chapmanina gassinensis was emended by Frizzell in 1949 tina declivis (Reuss, 1863) which was only reported from and well studied by Fuchs (1969). Barbin & Decrouez the Oligocene. (1987) work is of greatest interest, because they described It is interesting to link the presence of Chapmanina gas- in detail the complex structure of the species and gave sinensis in the Paris Basin with the one of Fabiania cassis. numerous references concerning its distribution. Chap- Fabiania cassis is mentionned by Bignot (1968) in the manina gassinensis has been found in the Eocene and the upper Lutetian of Mantes (western Paris Basin), then by Oligocene associated with a typic fauna of Nummulites Le Calvez (1970) in the Lutetian of Paris Basin (Le Til- and other larger foraminifera: Nummulites fabianii, N. let drilling), where she saw only a few specimens (cf. also aturicus, N. vascus, Discocyclina, Asterodiscus, Assilina, Lin- Curry in Le Calvez 1970). Bignot (1968) remarked that F. derina, Halkyardia, Fabiania, Pellatispira, etc. in Eocene cassis, peri-mediterranean species (Neumann & Boulanger (Fleury et al. 1985; Sztrákos & du Fornel 2003); Num- 1955) went rather far to the North during the Eocene. It mulites vascus, N. intermedius, Neorotalia, Amphistegina, is the same for Chapmanina gassinensis, rather uncommon etc. in Oligocene (see Boulanger 1968; Cahuzac & Poign- species, only observed at Grignon until now. ant 2002). oTher ciTaTions of Chapmanina Gassinensis The species of The GriGnon “falunière” in france (Yvelines, souTh-WesTern paris Basin) Eocene (Fig. 3) The lutetian exposure of Grignon called “falunière” is well Aquitaine. The Eocene is well represented and has been known in the Paris Basin because of its abundance in vari- largely studied. Ch. gassinensis has been reported by many ous well-preserved organisms (Guernet et al. 2012; Sanders authors in numerous localities in the Lutetian, Bartonian, et al. 2015; Fig. 2). The two found specimens don’t allow Priabonian. For example: Halkyard (1918: pl. 8, fig. 7): any doubt about their systematical attribution as their Eocene, blue marl of the “Côte des Basques”, Biarritz illustrations show (Fig. 1). They have been observed only in (Pyrénées-Atlantiques), named Patellina (see above); Cuvil- one level without any precise localisation, although many lier (1951): Bartonian, Chapmanina sp. in Abatilles drill- 465 GEODIVERSITAS • 2018 • 40 (17) Poignant A. ing (Arcachon, south of Bordeaux) and upper Eocene of ciTaTions of Chapmanina Gassinensis Biarritz; Sacal & Debourle (1957: pl. 19, fig. 9): middle or cf. Gassinensis or Chapmanina sp. to upper Eocene of Pontonx (Landes); Magné & Mal- in oliGocene of europe (Fig. 4) moustier (1964): northern Aquitaine Paleogene; Veillon Chapmanina gassinensis is reported by Gauthier (1971), in (1964): lower and middle Eocene of Arcachon; Boulanger a drilling, in Lluchmajor, in Palma de Mallorca Island; it is (1968: pl. 32, fig. 64; pl. 33, fig. 66): Bartonian, Biarritz frequent and associated with Lepidocyclinas and Miogypsinas. and Saint-Cricq-du-Gave (Landes); Deloffre & Hamaoui But this suggests a reworking, since Chapmanina has never (1973: pl. 11, fig. 8): upper Eocene of Labatut (Landes); been observed in the Miocene and B. Cahuzac (pers. comm.) Poignant (1991): upper Eocene of Lac Mouriscot (near told me that reworking is frequent in Mallorca. Biarritz); Mathelin & Sztrákos (1993): Truncatulinoides In Italy, Cita & Scipolo (1961) observed Chapmanina gas- rohri zone and lower zone of Bartonian age; Sztrákos et al. sinensis in the Oligocene of Monte Baldo (North-Western (1998): Bartonian of Listrac Marls in northern Aquitaine; Italy), and these authors suggested a reworking; but later Sztrákos (2000: pl. 17, fig. 15): Bartonian-Priabonian of on Chapmanina gassinensis was found also in the Oligocene Adour valley; Sztrákos et al. (2010): very abundant in the of south Apennine and of Vicenza province (Zanfra 1965; upper Eocene of the “Acacias drilling”, near Arcachon. de Zanche 1967) suggesting that the occurrence range of this species spans the Eocene to the Oligocene. South-eastern France. Blondeau et al. (1968): Ch. gas- Di Carlo et al. (2010) mentioned Chapmanina sp. in the sinensis would be abundant in the Alpes-Maritimes, in Rupelian of Zakinthos Island. In Slovenia, Pavlovec et al. a transition zone between Lutetian and Upper Eocene; (1986) found Chapmanina sp. in the Oligocene at Bohring Sztrákos & du Fornel (2003): Bartonian-Priabonian of and Nova Stifta. Alpes-Maritimes and Provence. ciTaTions in TeThYs (MediTerranean doMain Oligocene (Fig. 4) and easTern TeThYs; Fig. 5) Aquitaine. Ch. gassinensis is rather rarely mentioned in the Chapmanines and above all Chapmanina gassinensis, sometimes Oligocene: Veillon (1964): “Les Acacias” drilling; Bou- Chapmanina sp. have been frequently cited out of Europa, in the langer (1968): Lower Oligocene, bottom of “Villa Belza” Eocene of the Mediterranean domain or further towards East cliff, in Biarritz; Poignant (1972, erroneously attributed (e.g. in Morocco, Algeria, Tunisia, Syria, Turkey, Oman, Qatar, to Priabonian), Cahuzac & Poignant (2002): Orx pond, Tanzania, Iran). In North-eastern India, North of Bangladesh Saint-André-de-Seignanx (Landes). (Matsumaru & Sarma, 2010): Chapmanina spp., and Num- Cavelier et al. (1981), in a publication on the Eocene- mulites atacicus, N. globulus, N. burdigalensis and Fabiania cas- Oligocene limit, thought that Chapmanina gassinensis sis are found in Bartonian. The section figured (Matsumaru & disappeared exactly at the Eocene-Oligocene limit. They Sarma 2010: pl. 3, fig. 12) does not allow to assert if it is really said: “At the Eocene/Oligocene boundary there was a a Chapmanina, but the associated microfauna is of a Bartonian general drop of temperature accompanied by a period of age. The presence of a tethysian microfauna in the East of India great drought which was associated with the disappearance corresponds to the persistence of the Tethys ocean which will close or regression of most of the tropical species, for example: at the end of Burdigalian-Ottnangien (Rögl 1999). We can also Discocyclina, Asterocyclina, Lockhartia, Fabiania, Chap- remark the existence of aquitanian faunas (Miogypsina, Lepidocy- manina gassinensis, etc.” (Cavelier et al. 1981: 226, 228). clina) in the Qom Basin in Iran (Rahaghi 1973). The rotation of This work invalidates this opinion as far as Chapmanina Africa and Arabia meeting the anatolian platform will separate gassinensis is concerned. the Mediterranean sea from the Indian Ocean (Rögl 1999). In Australia, in Adelaïde region, Murray Lindsay & McGow- ciTaTions of Chapmanina Gassinensis or cf. ran, 1986, described an Eocene tethysian fauna, with Halk- Gassinensis or Chapmanina sp. in eocene of europe yardia, Linderina, but they did not mention any Chapmanina (Fig. 3) which could be associated to these species. Chapmanina gassinensis has been pointed out in many In Australia too (off southern New South Wales), Quilty & European countries: Albania, Croatia, Germany, Greece, Packham (2006), described Chapmanina conjuncta, in the Hungaria, Italy, Spain, Thrace. Frequently named cf. Paleocene and they saw a clear tethysian influence (see earlier, gassinensis or Chapmanina sp. (for instance: Di Carlo Murray Lindsay & McGowran 1986). At that period, Tethys et al. (2010), it is without any doubt gassinensis, because was widely opened towards the Indian Ocean. This new species of the associated microfauna which is the same as in has been observed in a micritic limestone, found at a depth France. Silvestri cited the genus in Toscania (Silvestri of 1750 m. The specimens are observed in thin sections: 1904), then in Piemonte region (Silvestri 1905a; Gassino, “Material: seven thin sections”. Planktonic foraminifera spe- see earlier), in Umbria (Silvestri 1905b) and in Veneto cies are very badly preserved and are not easily identifiable as (Silvestri 1923), Romania (Kovacs & Arnaud-Vanneau well as calcareous nannoplancton. No Nummulites is identi- 2004), Spain (Molina et al. 1986) thought that it would fied. According to Quilty & Packham (2006), Chapmanina be characteristic of the Upper Eocene in Italy and Spain), conjuncta will differ from Chapmanina gassinensis by a larger Thrace (Less et al. 2011). marginal zone, but their illustration is not conclusive, speci- 466 GEODIVERSITAS • 2018 • 40 (17) The foraminifer Chapmanina gassinensis in the Grignon “falunière” (Yvelines, Paris Basin). The species of the genus Chapmanina mens being known only in thin section. Moreover, the authors REFERENCES say (Quilty & Packham 2006: 331): “In some respects the aGip Mineraria. 1959. — Microfacies italiane: dal carbonifero al species found here seems intermediate between Cymbalopora miocene medio. San Donato milanese, 35 p., 145 pls. van Hagenow, 1851, and Chapmanina”. Consequently, it akkiraz M. s., akGün f., Örcen s., Bruch a. a. & MosBruGGer.v. seems difficult to have a reliable opinion. The authors give a 2006. — Stratigraphic and Paleoenvironmental Significance of Late Paleocene-Early Eocene age to this fauna. Bartonian-Priabonian (Middle-Late Eocene) Microfossils from the Bascesme Formation, Denizli Province, Western Anatolia. Turkish Journal of Earth Sciences 15: 155-180. oTher ciTaTions allison a. 1953. — La geologia della structura della zona di Priolo Barbin & Decrouez (1987), according to a A. Blondeau (pers. (Siracusa). 57e reunione della Societa Geologica Italiana. Guida comm.), reported the presence of Chapmanina gassinensis in alle escurzioni, Palermo. Bollettino della Società geologica italiana the Eocene of Mexico, this is not very surprising since Salm- 71: 109-111. erón (1986) has reported the occurrence of Fabiania cassis, anania M. G., BoTTino c., MaTTeucci r. & piGnaTTi J. s. 2000. — A late Bartonian-early Priabonian benthonic foraminif- often associated with C. gassinensis, in the Upper Eocene of eral assemblage from Tornimparte near l’Aquila (Central Italy). Mexico. In the “Foz dos Amazonas Basin”, Central Brasil, de Geologica Romana 36: 147-161. Mello e Souza et al. (2003), observed Fabiania cassis in the auBouin J. & neuMann M. 1959. — Contribution à l’étude Eocene, associated with american microfaunas. Chapmanina stratigraphique et micropaléontologique de l’Éocène en Grèce. gassinensis could also be present. Besides, Fleury et al. 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Developments in Paleontology and Stratigraphy 9: since widespread in the whole Tethys, as well as Fabiania cassis 49-52. https://doi.org/10.1016/S0920-5446(08)70093-1 to which it appears very often associated. Its distribution is BarBin v. & decrouez d. 1987. — Le genre Chapmanina (foraminifère) : État des connaissances et distribution géographique. even worldwide since pointed out in the american continent Archives des Sciences 40 (2): 207-224. (Upper Eocene of Mexico, see earlier). BenedeTTi a. 2010. — Biostratigraphic remarks on the Caltavuturo We must notice that the extension of Chapmanina gassinensis Formation (Eocene-Oligocene) cropping out at Portella Colla towards the North does not seem to exceed some latitude. (Madonie Mounts, Sicily). Revue de Paléontologie de Genève 29 The Paris Basin could be its northernest distribution during (1): 197-216. BiGnoT G. 1968. — Nouvelle découverte de Fabiania cassis (Oppen- the Eocene. It had not citated by Kaasschieter (1961) in his heim) dans le Lutétien du Bassin de Paris. Colloque sur l’Éocène. important work on the Eocene of Belgium. Bavaria, where it Bureau de Recherches géologiques et minières n°58, Paris: 79-81. has been found (Darga 1990) is at a lower latitude. Blondeau a., Bodelle J., caMpredon r., lanTeauMe M. & Very abundant during the Eocene, it becomes rare at the neuMann M. 1968. — Répartition stratigraphique des grands Oligocene, wh ere Greece seems to be its easternest localisa- foraminifères de l’Éocène dans les Alpes-Maritimes (franco- italiennes et Basses-Alpes). Colloque sur l’Éocène. Bureau de tion. It does not occur in the Miocene. Recherches géologiques et minières n°3: 13-26. It would be interesting to study the Eocene specimens of Bonnefous J. & BisMuTh h. 1982. — Les faciès carbonatés de plate- Chapmanina gassinensis from India since associated to a Medi- forme de l’Éocène moyen et supérieur dans l’offshore tunisien nord- terranean microfauna: Nummulites, Fabiania. The presence of oriental et en mer pélagienne : implications paléogéographiques et the species on the american continent would ask for a work analyse micropaléontologique. Bulletin du Centre de Recherches et d’Exploration-Production, Elf-Aquitaine 6 (2): 337-403. of geographical and stratigraphical distribution, as also the BoulanGer d. 1968. — Révision du Nummulitique de la Chalosse, specimen found in New-Caledonia. du Béarn et du Bas-Adour (Landes et Basses-Pyrénées). Thèse de The new australian species, supposed Paleocene, would also la Faculté des Sciences de l’Université de Paris, 332 p., 41 pls. require further studies to confirm its age and its taxonomical BozorGnia f. (avec la collaBoraTion de BanafTi s.) 1964. — attribution. Microfacies and Microorganisms of Paleozoic through Tertiary Sediments of some parts of Iran. Natural iranian oil Company, Teheran-Iran, 22 p., 152 pls. cahuzac B. & poiGnanT a. 2002. — Associations de foraminifères Acknowledgements benthiques dans quelques gisements de l’Oligo-Miocène sud- I am deeply indebted at first to Annachiara Bartolini who aquitain. Revue de Micropaléontologie 47 (3): 221-256. https:// made optical photographs and carefully revised the manu- doi.org/10.1016/S0035-1598(02)90027-9 script and Alexandre Lethiers who realized the outline of the casTellarin a. & ciTa M. B. 1968. — Chapmanina gassinensis dans l’Éocène moyen de la coupe de Gallerie. Étude de quelques coupes carts of distribution of the species. Special thanks are due to priaboniennes dans le Monte Baldo (Province Verona et Trento, V. Barbin, H. Bismuth, B. Cahuzac, H. Salles-Astier for their Italie) et discussions des limites de l’étage. 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