2 ArachnologischeMitteilungen43:5-16 Nuremberg,July2012 On the spider species described by L. Koch in 1882 from the Balearic Islands (Araneae) Robert Bosnians&JohanVan Keer doi;10.5431/aramit4306 Abstract:ExaminationoftheL.Koch collection oftheZoological Museum in Berlin allowsusto proposethefol- lowing new synonyms and corr\b\naX\ons:Erigonemarina L.Koch, 1882 = Oedothoraxfuscus (Blackwall, 1834) n.syn.; Theridionelimatum L.Koch, 1882 =Enoplognathadiversa (Blackwall, 1859) n.syn.;LiocranumvariabilisWunderlich,2008 = ZorainornataL.Koch,1882n.syn.=Liocranuminornatumn.comb.;LycosaperspicaxL.Koch,1882=Arctosafulvolineata(Lucas, 1846)n.syn.;AlopecosellaRoewer,1960=ArctosaC.L.Koch,1847n.syn.;LycosasubhirsutaL.Koch,1882=Arctosalacustris (Simon, 1876) n.syn.;Philodromusvegetus L.Koch, 1882 = ThanatusvulgarisSimon, 1870 n.syn.;OzyptilabicuspisSimon, 1932=OzyptilafurculaLKoch,1882 n.syn.;Haplodrassusmaroccanus Denis,1956=DrassusparvulusL.Koch,1882 n.syn. =Haplodrassusparvicorpus(Roewer,1951)n.comb.(replacementnaroe);ZelotesruscinensisSimon,^9^4=Zelotessemirufa (L.Koch, 1882) n.syn.;PhlegrasimoniL.Koch, 1882 =PhlegrabresnieriLucas, 1846 n.syn.; Trochosulaconspersa (L.Koch, 1882),Lycormafraisnei(L.Koch,1882),Lycormainsulana(L.Koch,1882),Arctosamisella(L.Koch,1992) and Piratasimplex(L. Koch,1882)areallretransferredtotheiroriginalgenusLycosastat.rev.CheiracanthiumoccidentaleL.Koch,1882,Ozyptila furculaL.Koch,1882andZelotescallidus(Simon,1878)areredescribed. Keywords:BalearicIslands,L.Koch,redescriptions,spiders,synonymies In recent years, the number ofdescriptions ofnew Thereferencematerialmentionedinthepresentpaper species hasincreasedconsiderably,greatlyexpanding ispartofthecollectionofthefirstauthor. Specimens the knowledge of spiders. However, it is also very wereexaminedandillustratedusingaWildM5 ster- important to revise or redescribe species from the eomicroscope. Further details were studied using an 19^^ century. Some of these older descriptions are OlympusCH-2stereoscopicmicroscopewithadraw- incomplete and not accompanied by illustrations, ingtube.Leftstructuresaredepicted.Malepalpswere but others have adequate descriptionswith excellent detached and transferredtoglycerolforexamination figures. One ofthe papers that has been completely underthe microscope. Female genitaliawere excised forgottenis‘ZoologischeErgebnissevonExcursionen using sharpened needles. These were transferred to aufden Balearen. II. Arachniden und Myriapoden’ cloveoilforexaminationunderthemicroscope.Later, by L. Koch (1882). The author described 28 new palps and epigyneswere returned to 70% ethanol. species ofspiders in it, ofwhich only ten have been fully redescribed, which means that 18 ofthem are The following abbreviations are used in the text: left as poorlyknown species. CRB: Collection RobertBosnians; MNHNP:Museum nationald’Histoire naturelle,Paris; Materialandmethods ZMB: ZoologischesMuseum Berlin Type material ofnine species could be loaned from the BerlinMuseum (possible otherlocations oftype CommentsonthespeciesdescribedbyL. Koch material in London orViennawere not checked). If Many authors of the 19*^^ or the beginning of the the material was not present, the descriptions and 20^^^ century, for example Thorell and Strand, did figures were carefully examined and compared with not present figures at all, so these species cannot be similarorrelated species from the Mediterranean. recognized without examination of the type mate- rial. Koch (1882) reportedtwenty-eightnewspider species from the Balearic Islands. His descriptions RobertBOSMANS,UniversityofGhent,TerrestrialEcologyUnit, areveryaccurate andwere accompaniedbyexcellent Ledeganckstraat35,9000Gent,Belgium. E-Mail:[email protected] figures,which allowthe recognition ofdetails in the JohanVANKEER,Bormstraat203bus3,1880Kapelle-op-den-Bos, palpal and epigynal structures. Careful examination Belgium.E-Mail:[email protected] of the figures should have allowed identification submitted:14.12.2011;accepted:15.3.2012;onlineearly:25.8.201 of several species, especially when compared with 6 R.Bosnians&J.vanKeer other material from the Mediterranean region. It is cies(authors’personalobservations)andtheholotype therefore surprising that L. Koch’s paper has been ofDysderamordaxsN2&comparedwiththisspecies.In neglected by previous authors. Ofthe twenty-eight D. mordax^the rugosityofthe prosoma and the ster- described species, only ten have subsequently been numisdifferentandthecolourismoreburgundyred. studied. Out ofthese ten redescribed species, seven Spinationoftheholotypemaybeincomplete,butno appeared to be synonyms, and only Iberesia brauni, spinesareobservedonlegsIandII(probablylost),leg Xelotes semirufus and 7.,flagellans remain valid. The IVhas2basalspinesonthefemora,legsIII—IVhave ten species concerned are listed inTab. 1. 2pairsoflateralspinesand 1 pairofventralspineson the tibia, and several spines on the metatarsi. In D. Twelve of the eighteen remaining species have crocotatibiae III-IV have fewer spines. For the male received new names by preoccupation, were trans- palp we must rely on the figure ofKOCH (1882; see ferredtoothergeneraorweredeclared nominadubia. Fig. 1).The bulb is similarto thatofDysderacrocota., However, none ofthemwere redescribed.The other but more slender, narrowing terminally and more We species have neverbeen subsequentlycited,exceptin pointed. consideritavalidspecies,buttopotypic catalogues.These species are listed inTable 2. material is needed for a complete redescription. Distribution So far, the species is an endemic to Mallorca. Taxonomy FamilyDysderidae FamilyTheridiidae DysderamordaxL. Koch, 1882 (Fig. 17) Enoplognathadiversa (Blackwall, 1859) (Figs2-3) DysderamordaxL. Koch, 1882: 640,pi.20,fig.20. TheridionelimatumL.Koch,1882:630,pi.20,fig.8;new Typematerial synonymy. Holotype male ofDysdera mordax from Spain, Baleares, Typematerial Mallorca,Palma,beginningofMay, Schaufussleg. (ZMB Holotype female ofTheridion elimatum from Spain, Bal- 7905);examined,butbothpalps are absent. eares,Mallorca,PalmaRiera,22April, Schaufussleg.;not Comments examined,unavailablein ZMB. In absence ofthe palps, a complete newdiagnosis of Comments thisspeciescannotbegiven.InMallorca,Dysderacro- According to KOCH (1882), the prosoma of this cataC.L.Koch,1838isthecommonestDysderaspe- species isyellowishbrown, the abdomen greybrown Tab.1:Listofspeciesdescribed byL.Koch(1882)which havealreadybeen redescribed Kochsname Currentname Author(s) FamilyNemesiidae NemesiabrauniL. Koch, 1882 Iberesiabrauni(L. Koch, 1882) Decae&Cardoso(2006) FamilyTheridiidae Theridion mansuetum L. Koch, 1882 Enoplognathamandibularis(Lucas, 1846) Bosmans&VanKeer(1999) MetaschaufussiL. Koch, 1882 Enoplognathamordax(Thorell, 1875) Wunderlichin:Merrett& Snazell(1975) FamilyAraneidae SinganigrofasciataL. Koch, 1882 Hypsosingaalbovittata(Westring, 1851) Denis(1952) EpeiramimulaL. Koch, 1882 Neosconaadianta (Walckenaer, 1802) Roewer(1942) FamilyLycosidae LycosasubterraneaL. Koch, 1882 Arctosafuholineata(Lucas, 1846) LUGETTI&TONGIORGI(1965) PardosavenaticaL. Koch, 1882 PardosacribrataSimon, 1876 Wunderlich(1984) FamilyCorinnidae TrachelasflavipesL. Koch, 1882 Paratrachelasmaculatus(Thorell, 1875) BOSSELAERSetal.(2009) FamilyGnaphosidae ProsthesimaflagellansL. Koch, 1882 Zelotesßagellans(L. Koch, 1882) Senglet(2011) ProsthesimasemirufaL. Koch, 1882 Zelotessemirufus(L. Koch, 1882) Senglet(2011) SpidersdescribedbyL.KochfromtheBalearicIslands 7 Tab.2.Listofspeciesthathavereceived newnamesbypreoccupation,weretransferredtoothergenera,weredeclarednomina dubiaorwerenevermentionedagain in literature Koch’sname Currentname Action FamilyDysderidae DysderamordaxL. Koch, 1882 DysderamordaxL. Koch, 1882 None FamilyTheridiidae TheridionelimatumL. Koch, 1882 TheridionelimatumL. Koch, 1882 None FamilyLinyphiidae ErigonemarinaL. Koch, 1882 ErigonemarinaL. Koch, 1882 None FamilyZoridae ZorainornataL. Koch, 1882 ZorainornataL. Koch, 1882 Nomendubium (URONES 2005) FamilyMiturgidae CheiracanthiumoccidentaleL. Koch, 1882 CheiracanthiumoccidentaleL. Koch, 1882 None FamilyLycosidae LycosaperspicaxL. Koch, 1882 Alopecosellaperspicax(L. Koch, 1882) TransferbyROEWER(1955) LycosasubhirsutaL. Koch, 1882 LycosasubhirsutellaRoewer, 1955,but Replacementname,later transferredbacktoLycosasubhirsutain transferredback Roewer(1960) LycosaconspersaL. Koch, 1882 Trochosulaconspersa(L. Koch, 1882) TransferbyROEWER(1955) LycosafraisseiL. Koch, 1882 Lycormafraissei(L. Koch, 1882) TransferbyROEWER(1955) LycosainsulanaL. Koch, 1882 Lycormainsulana(L. Koch, 1882) TransferbyROEWER(1955) LycosamisellaL. Koch, 1882 Arctosamisella(L. Koch, 1882) TransferbyROEWER(1955) LycosasimplexL. Koch, 1882 Piratasimplex(L. Koch, 1882) TransferbyROEWER(1955) PardosatenuipesL. Koch, 1882 PardosatenuipesL. Koch, 1882 None FamilyPhilodromidae PhilodromusvegetusL. Koch, 1882 PhilodromusvegetusL. Koch, 1882 Nomendubium (BRAUN 1965) FamilyThomisidae OzyptilafurculaL. Koch, 1882 OzyptilafurculaL. Koch, 1882 None FamilyGnaphosidae DrassusparvulusL. Koch, 1882 DrassodesparvicorpusRoewer, 1955 Replacementname ProsthesimasemirufaL. Koch, 1882 Zelotessemirufus(L. Koch, 1882) Redescription(SenGLET 2011) ProsthesimaplumigeraL. Koch, 1882 Zelotesplumigerif. Koch, 1882) TransferbyROEWER(1954) FamilySalticidae PhlegrasimoniL. Koch, 1882 PhlegrasimoniL. Koch, 1882 None withadarkfoliumwithwhitespotsandborderedwith Koch, 1882is therefore consideredajuniorsynonym white, ventrallywith two parallel white stripes, legs ofEnoplognatha diversa (Blackwall, 1859). It should yellowish brown. The description ofsuch a folium benotedherethatanother TheridionspeciesinKoch’s pointsclearlyinthedirectionofEnoplognathaspecies paper{Theridion mansueturri) appeared tobe ajunior ofthe diversaor mandibularisgroup. Koch’s figure of synonym Enoplognathamandibularis(Lucas,1846), & theepigyneprovidesfurtherdetailsthatpointclearly see BOSMANS VAN KEER (1999). in the directionofE. diversa(compare Figs2-3),the Distribution most common Enoplognatha species in Spain (BOS- Mediterranean,fromtheIberianPeninsulaandMo- & MANS Van KEER 1999). Theridion elimatum L. rocco in thewest, to Crete in the east. 8 /?.Bosnians&J.vanKeer Liocranum variabilis Wunderlich, 2008: 506, figs 42-46 (descr. male, female); newsynonymy. Typematerial Holotype female of Zora inornata from Spain, Baleares, Mallorca, Miramare, IV.1866, Schaufuss leg. (ZMB 7920); ex- amined. Comparativematerialexamined SPAIN. Caceres: Plasencia, 1 female of Liocranum majus Simon, 1878, IV.1990, P. Pootleg. (CRB). Comments From Koch’s drawing of the epigyne (1882, fig. 19) it is evident that this species does not belong in the genus Zora.The figure shows alarge,anterior pocket and two smaller lateral pockets (Fig. 6). Examination ofthe specimen shows it to belong in the Liocranidae Figs.1-10:Fig.1.MalepalpofDysderamordaxL.Koch,1882inL.Koch(1882). and that it is identical to Liocranum Figs.2-3:Enoplognathadiversa(Blackwall,1859)2.EpigyneofTheridionelimatumin Koch(1882);3.EpigyneofEnoplognathadiversa\n Bosmans&Van Keep(1999). variabilis^ onlyrecently described from Figs.4-5.Oedothoraxfuscus(Blackwall,1834).4.EpigyneofErigonemarinain Koch MallorcabyWUNDERLICH (2008),see (1882).5.EpigyneofOedothoraxfuscusin Locket&Millidge(1953).Figs.6-7: Fig.7.Thislatterspeciesthusbecomesa Liocranuminornatum(L.Koch,1882).6.EpigyneofZorainornatain Koch(1882); juniorsynonym.Liocranuminornatumis 7.EpigyneofLiocranumvariabilisinWunderlich(2008).Figs.8-10:Cheiracanthium occidentaleL.Koch,1882.8.EpigyneofCheiracanthiumoccidentalein Koch(1882).9. closelyrelatedtoL. majusSimon, 1878, Vulva,ventralview;10.Idem,dorsalview. recentlyredescribedbyLEDOUX(2008). This author confirmed the synonymy FamilyLinyphiidae ofL. majuswith Liocranum apertum Denis, 1954, L. Oedothoraxßiscus (Blackwall, 1834) (Figs.4-5) pallidulum Simon, 1878 and L. segmentatum Simon, Erigone marina L. Koch, 1882: 629, pi. 20, fig. 7 (descr. 1878. According to WUNDERLICH (2008), the female);newsynonymy. Typematerial helmet-like structure is largerinL. major. Distribution Holotype female ofErigone marina from Spain, Baleares, Mallorca, Soller, mid-April, Schaufuss leg. (ZMB 7915); So far, the species is an endemic to Mallorca. examined. Comments FamilyMiturgidae Examination ofthe holotype clearly shows that this CheiracanthiumoccidentaleL. Koch, 1882 (Figs. 8-10) species is identical to Oedothoraxfuscus (Blackwall, CheiracanthiumoccidentaleL.Koch,1882:637,pi.20,fig. 1834)andthusErigonemarinahtcomtsajuniorsyno- 16 (descr.female). Typematerial nym ofthat species. Koch’s drawing ofthe epigyne HolotypefemaleofCheiracanthiumoccidentalefrom Spain, (Fig. 4) is somewhatconfusingbutshows clearlythe Baleares, Minorca, Mahon, 18.V.1866, Schaufuss leg. twotypicalcurvedlateralfoldsasinFig.5 takenfrom (ZMB 7929);examined. Locket 5cMillidge (1953). Comments Distribution Koch’s figure 16 (1882) clearly shows a Cheira- Europe, North Africa, Azores, European part of canthium-Xik.t epigyne (Fig. 8), but does not give Russia. enough diagnostic characters for identification. A redescription ofthe epigyne and vulva is therefore FamilyLiocranidae given here. According to the author, the species is Liocranum inornatum (L. Koch, 1882) new combina- related to Cheiracanthium letochae L. Koch, 1876 (= tion (Figs 6-7) C. elegansTXcovtWy 1875).The epigyne has a median Zora inornata L. Koch, 1882: 639, pi. 20, fig. 19 (descr. depressionwith aninteriormargin andiswiderthan female). SpidersdescribedbyLKochfromtheBalearicIslands 9 long.The spermathecae are relativelysmall,only 1/3 Comparativematerialexamined ofthewidthofthedepression.Thecopulatoryopen- SPAIN. Caceres: Plasencia(N 40T’52”E 6°5’18”),525m, ingsaresituatedantero-laterallyofthedepressionand 1 male 1 female,IV.1990,P.Pootleg. (CRB). Comments the sperm ducts make three coils to the elongated spermathecae (Figs 9-10). In the literature, no spe- The holotype male has only one palp left with a cies with such avulva could be found and hence the tegular apophysis as inArctosa lacustris. The female species is consideredvalid. hasanintactepigynewithabroadmedianseptum,as Distribution clearlyshowninKoch’sfigure29(1882),correspond- Onlyknown from the type locality. ingwellwiththeepigyneof.^.lacustris(compareFigs 13, 15 with Figs 14, 16 (taken from KNÜLLE 1959). FamilyLycosidae Lycosasubhirsutaand its replacement nameAlopecosa Arctosafulvolineata(Lucas, 1846) (Figs 11-12) subhirsutellathus becomejuniorsynonyms oiArctosa LycosaperspicaxL. Koch, 1882:658,pi.21,fig. 32 (descr. lacustris. female);newsynonymy. Distribution Alopecosellaperspicax, Roewer, 1955:225. Mediterranean, CanaryIslands. Typematerial Holotype female ofLycosaperspicaxfrom Spain, Baleares, LycosaconspersaL.Koch, 1882stat.rev.(Fig. 17) Mallorca, Soller valley, mid-April, Schaufuss leg. (ZMB LycosaconspersaL. Koch, 1882: 661 pi.21,fig. 33 (descr. 7910);examined. female). Comparativematerialexamined Trochosulaconspersa', Roewer, 1955: 304. FRANCE.Aude: Gruissan, N. les Pujots (N 43°06’30” E Typematerial 3°3’28”), 3m, 1 female, litter in salt marsh, 1.IV.1980, R. Type series oftwo females ofLycosaconspersafrom Spain, Bosmansleg. (CRB). Baleares, Mallorca, Ses Prat de SanJordi, end April, and Comments Soller, mid-May, Schaufuss leg. (ZMB 7912); examined. The holotype female has an intact epigyne and on The two females belong to different species. One ofthe examination it can immediately be recognised as femalesis,Arctosafuholineata,theotheronehasanepigyne Arctosafuholineata. Lycosaperspicax thus becomes a correspondingtotheoriginaldrawingbyKOCH(1882)and junior synonym. Koch’s figure is sketchy (Fig. 7) but is selectedhere as thelectotype. Comments shows the typical antero-median, triangular septum ofiA.fuholineata(Fig. 8,takenfrom KNÜLLE 1959). This species has not been mentioned since the Roewer (1955) created the new genusAlopecosella originaldescription,withtheexceptionofR0EWER‘S forthisspecies.SincethetypeofthegermsAlopecosella (1955) transfer to the genus Trochosula,without any is here transferred to Arctosa, the genus Alopecosella justification. KOCH’s figure 33 (1882) ofthe epigyne becomes ajuniorsynonym oiArctosa. onlyother resembles the epigyne of Hogna radiata (Latreille, speciesofthegermsAlopecosella,A.pelusiaca(Audouin, 1817), a widespread species in the Mediterranean. 1826) has to be returned to Alopecosa, where it was Like manylarge lycosids in the Mediterranean, this placed byCAPORIACCO (1936). speciescomplexisinneedofrevisionandaconclusion Distribution about synonymyhas to be postponed.Materialfrom Western Europe, Iberian Peninsula, Italy, south of Mallorca is needed to resolve the situation. This is France and the Maghreb. also the case for three otherLycosa species described byL.KochfromMallorca(seebelow):Lycosafraissei, Arctosalacustris(Simon, 1876) (Figs 13-16) L. insulana and L. simplex.They are all large species Lycosa subhirsuta L. Koch, 1882: 653, pi. 21, figs 28-29 and Koch’s figures 33-36 all show the same type of (descr. male,female);newsynonymy. epigyne(seeFigs 17-20).ROEWER(1955)transferred Alopecosa subhirsutella Roewer, 1955: 221 (replacement these species (without furtherjustification) to three name);newsynonymy. different genera: Trochosula, Lycorma and Pirata. To Lycosa subhirsuta', Roewer, 1960: 874 (transferred back, facilitatefuturestudies,weconsideritbettertoreturn withoutarguments). themtotheiroriginalgenus.Topotypicmaterialmay Typematerial help solve the identityofthese problematic species. Lectotype male and paralectotype female ofLycosa sub- Distribution Mhiirrsaumtaarf,roMmayS,paSicnh,auBfaluesasrelseg,.S(olZlMerBva7l9l1ey1,);miedx-aAmpirnield.and The type localityonMallorca. 10 R.Bosnians&J.vanKeer LycosafraisseiL.Koch,1882stat.rev. (Fig. 18) LycosafraisseiL. Koch, 1882: 666, pL 21, fig. 36 (descr. male). Lycormafraissei\ Roewer, 1955: 265 (transfer). Typematerial Holotype male ofLycosafraissei from Spain, Baleares,Mallorca, Fraisse leg.; notexamined,unavailablein ZMB. Comments A mm large lycosid of 19 total length. The type material is not available. See comments under Lycosaconspersa. Distribution The type localityon Mallorca. Lycosa insulana L. Koch, 1882 stat. rev. (Fig. 19) Lycosa insulana L. Koch, 1882: 664,pi.21,fig.35 (descr.female). Lycorma insulana\ Roewer, 1955: 265 (transfer). Typematerial HolotypefemaleofLycosainsulanafrom nSpoatienx,aBmailneeadre,su,nMaavlaliloarbclae,iFnraZisMsBe.leg.; Fig/iscn./cK1uos1cm-sh2(3(S.1iF8mi8og2ns).,;111128-.716E2):p.iA1rg3cy.tnoMseaaolffuel4vrpocalfio\snpöe/aiot/a/fv'(LoLy/cu/>oc7saeafsl,srufl1bh8ii4nr6sK)un.tüa1l\1ln.eEK(po1ic9gh5y9n()1.e8F8oi2fg)sL;yc1o4s.aMpearlsAepricctpaoaxslap Comments ofArctosalacustrisin Knülle(1959);15.EpigyneofLycosasubhirsutain Koch(1882); A large lycosid of 16 mm total 16.EpigyneofArctosalacustrisin Knülle(1959).Fig.17.EpigyneofLycosaconspersain length.Thetypematerialcouldnot Koch(1882).Fig.18.EpigyneofLycosafraisei\n Koch(1882).Fig.19.EpigyneofLycosa insulana'm Koch (1882).Fig.20.Epigyneoffycosos/mp/eA'in Koch(1882).Fig.21.Male be examined. See comments under palpofPardosatenuipesin Koch(1882).Figs22-23.ThanatusvulgarisSimon,1870.22. Lycosaconspersa. EpigyneofPhilodromusvegetusin Koch(1882).23.EpigyneofThanatusvulgarisinSzita Distribution &Samü(2000). The type localityon Mallorca. Piratasimplex'., Roewer, 1955:284. LycosamisellaL. Koch, 1882 nomendubium Typematerial Lycosa misellaL. Koch, 1882: 660 (descr.juvenile). Holotype male ofLycosa simplex Spain, Baleares, Palma Arctosa misella\ Roewer, 1955: 226 (transfer). citymoats,beginMay,and SesPratdeJordi,beginningof Typematerial May, Schaufussleg.; notexamined,unavailablein ZMB. Comments Juvenile holotype ofLycosa misella from Spain, Baleares, SesPratdeJordi,endApril,Schaufussleg.,notexamined, The holotype could not be examined. According unavailablein ZMB. to the original description by KOCH (1882), it is a Comments large species (23 mm) suggesting a lycosid genus The holotype is ajuvenile specimen and its status is encompassinglargerspecies,suchasHogna^Lycorma unclear.Lycosa misellais therefore a nomen dubium. orLycosa. KOCH’s (1882) figure ofthe epigyne does Distribution not look like a Pirata epigyne at all and the transfer The type localityonMallorca. proposedbyROEWER(1955)seemsarandomchoice. See also comments underLycosaconspersa. LycosasimplexL. Koch, 1882 stat.rev. (Fig.20) Distribution LycosasimplexL. Koch, 1882: 663, pi. 21, fig. 34 (descr. The type localityonMallorca. female). SpidersdescribedbyLKochfromtheBalearicIslands 11 PardosatenuipesL. Koch, 1882 (Fig.21) but Koch’s figure 21 (1882) allows a positive iden- PardosatenuipesL. Koch, 1882:649,pi.21,fig.24(descr. tification with respect to the tegular apophysis of male). specimenscollectedbyusinSpainandNorthAfrica. Pardosopstenuipes\ Roewer, 1955: 197. The tegulum has an oblique banana-shaped tegular Pardosatenuipes\Tongiorgi, 1966: 351. apophysis with two postero-median concavities Typematerial (compare Figs 24 and 25). Such an apophysis exists Holotype male ofPardosa tenuipes from Spain, Baleares, in three Ozyptilaspecies occurring in this region: O. Ses PratdeJordi,mid-May, Schaufussleg.;notexamined, unavailablein ZMB. pauxilla Simon, 1870, O.perplexa Simon, 1875 and Comments O. bicuspis Simon, 1932. Koch’s species also has two Roewer(1955) createdthegenus Pardosopsforthis postero-median incisions, and these are absent in O. and other species, but TONGIORGI (1966) syno- pauxillaand O.perplexa. considerO.furculaand O. nymised it with Pardosa. The holotype male could biscuspisdoesamespeciesand O.bicuspisSimon,1932 notbe examined. KOCH’s (1882) figure 24 offers no becomesajuniorsynonymofO.furculaL.Koch,1882. indication of its affinities (see Fig. 21). Topotypic Ozyptilapauxilla and O.perplexaxv'iW be redescribed materialis needed to clarifyits systematic position. in a separate paper. Distribution Distribution The type localityonMallorca. The species is currently known from the south of France and from Spain. FamilyPhilodromidae Family Gnaphosidae Thanatusvulgaris Simon, 1870 (Figs22-23) Philodromus vegetus L. Koch, 1882: 645, pi. 20, fig. 22; Haplodrassusparvicorpus(Roewer,1951)newcombina- newsynonymy. tion (Figs27-28) Typematerial Drassusparvulus L. Koch, 1882: 632, pi. 20, fig. 10-11 Holotype female ofPhilodromus vegetus from Spain, Bal- (descr.male). Drassusparvicorpus Roewer, 1951: 443 (replacement eares,Mallorca,Miramare,beginMay, Schaufussleg.;not examined,unavailableinZMB. name). Comments Haplodrassus maroccanus Denis, 1956: 196, fig. 4-6; new Figure22inKOCH(1882) showsaroundedepigynal synonymy (here removed from the synonymy ofH. dalmatensis(L. Koch, 1866)). plate with some concentric circles, typical for the Typematerial epigyne ofthe common Thanatus vulgaris (compare Holotype male ofDrassusparvulus from Spain, Baleares, Figs 22 and 23).This species has no less than eight Mallorca, Riera near Palma, 22.IV.1882, Schaufuss leg. synonymsintheMediterraneanregion (LEVY 1977) (ZMB 7913);examined. and there is no doubt that Philodromusvegetusis yet Type series ofHaplodrassus maroccanus from Morocco, Pr. another one. Taroudant,OuledTeima {- Houara), accordingto DENIS Distribution (1956) composed of1 male 3 subadult males, 1 female 3 Circum-mediterranean. subadult females, 21.11.1954 and 1 subadult female from Amzou; notexamined,notfoundin theMNHNP. Comparativematerialexamined FamilyThomisidae SPAIN. Cadiz:Tarifa (N 36°0’50” E 5°36’25”), 2 males 5 Ozyptilafurcula L. Koch, 1882 (Figs24-26) females,IV.1992,P. Pootleg. (CRB). OzyptilafurculaL. Koch, 1882: 648,pi.21,fig.23. Comments Oznyeptwilsaybniocnuyspmiys.S)\.mon,1932:873,figs1186-1187,1208; Drassusparvulus was described by KoCH (1882). Typematerial Being preoccupiedbyDrassusparvulus Lucas, 1846, Holotype male of Ozyptilafurcula from Spain, Baleares, Roewer (1951) offered Drassusparvicorpus as a Mallorca, Ses Prat de SanJordi, end ofApril, Schaufuss replacement name.The species has neverbeen cited leg.;notexamined,unavailablein ZMB. since. Comparativematerialexamined AccordingtoKOCH(1882),thisspeciesisrelated SPAIN.Jaen: RiberaBaja (N 37°26’58”E 3°50’5”), 870m, toDrassusminusculus(=Haplodrassusdalmatensis)and 1 male,litterinPopulusforest,6.IV.1997, R.Bosmansleg. differsbythemorearchedprosomawithablackmar- (CRB). PM gin,andthe dispositionofthe eyes,withthe not Comments touchingandtheAM.separatedbynearlytheirdiam- The type material of this species is not available, eter.ThesecharactersaretoovariableinHaplodrassus 12 R.Bosnians&J.vanKeer to distinguish species, but Kochs figure 10 (1882) 6°4’12”), 270m, 4 males 2 females, pitfalls, 15.V1T23. shows amalepalpwithalarge,subterminaltoothon VIII.1996,U.WStengeleleg.(CRB);Talavan,FincaelBaldio thetegularapophysis,andhisfigure 11 showsapalpal (N39°43’12” 6T9’4”),370m,10males2females,pitfalls, tibiawithanapophysis aslongaswide (Fig.27).The 10.VIT5.IX.1996W,U.Stengeleleg.(CRB).Granada:Santa malepalps oftheholotype malearebothpresentand Fe (N 37T1’31” 3°45’17”), 700m, 1 female,litterinir- ingoodcondition (Fig.28). Examinationshowsthat rMiaglaatgead:PoCpouilnu,salongR9i.oVGIrIaL1n9d9e1N,R3.6B°4o1s’m2a9n”sWle4g.°4(8C’R22B”))., theyare identical to the palps ofHaplodrassusmaroc- 110m, 1 female,stonesingrassland, 15.VII.1991, R. Bos- canus^describedfromMoroccobyDENIS (1956) and Wmansleg.(CRB).Murcia:PuertoLumbreras(N37°29’37” occurring all over the south-western Mediterranean 1°51’9”),530m, 1 female,understone, 14.VII.1991, R. (unpublished data). In H. dalmatensis^ the tegular Bosmansleg.(CRB).-ALGERIA.Oran:MerselHadjad apophysishasonlyasmallsubterminaltooth,andthe (N35°47’52”W0°9’51”),2males,litteringarden,IX.1988, tibial apophysis is shorter than wide; thus there are R.Bosmansleg.(WCRB).-MOROCCO.Essaouira:Ounara sufficientdiagnosticcharacterstoseparatethespecies. E.(N31°32’33” 9°30’47”),250m,1 male,stonesingar- HaplodrassusmaroccanusDenis,1956becomesajunior ganiersteppe, 8.VII.1999, R.Bosmansleg. (CRB). Comments synonymofH.parvicorpus(Roewer, 1951 n.comb.). The discovery of large series of Zelotes callidus in The synonymyofH. maroccanuswithH. dalmatensisy proposedbyLEVY(2004)isrejected.Thespecieswill central Spain - including males and females - al- lows us to resolve a complicated case ofsynonymy. be fullyredescribed in afurtherpaper. The males from Central Spain could be identified Distribution as Zelotes callidus, based on the original figures of Mallorca andMorocco. Simon (1878, fig. 23; 1914, figs 297-299); a species originallydescribed from Corsica. Characteristic are Zelotescallidus(Simon, 1878) (Figs29-35) the two anterior teeth in the bulb, compare Figs 32- Prosthesima callida Simon, 1878: 91 (descr. male, non female = Z. caucasius). 33). The females did not correspond with Simons ProsthesimasemirufaL.Koch,1882:636,pi.20,fig15(de- figuresofZ.callidus,whichcorroboratestheobserva- scr. female);newsynonymy. tion ofSenglet (2004) that the females described Zelotescallidus-,Simon,1882:37(descr.male,nonfemale); as Z. callidushySiMON (1878,fig.24; 1914,fig.354), Simon, 1914:219 (descr.male,nonfemale). JEZEQUEL(1962,fig.29) andLedOUX(1972,fig. 1) Zelotes ruscinensis Simon, 1914: 157, 169, fig. 259, 346 all illustrate the epigyne orvulva ofZelotes caucasius (descr. male, female); Senglet, 2004: 104, figs 47-50 (L. Koch, 1866). (descr. male,female);newsynonymy. The females from Central Spain couldbe identi- Zelotes circumspectus-, Denis 1935: 117 (descr. female); fied as Zelotes semirufus (L. Koch, 1882) described misidentification. ZeloteslugensDenis 1941: 162 (correctionofthecitation from Menorca. KOCHs figure of Zelotes semirufus of Z. circumspectus from 1935); Di Franco 1997: 258 (Fig. 29) shows an epigynewith a median plate that (synonymybySenGLET2004). is slightly longer than wide, with an open posterior Zelotes adolescentulus Denis 1952: 118 (synonymy by margin, and comparable position of the rounded Senglet2004). spermathecae (Fig. 30). Zelotes massiliensis Soyer, 1967: 278 (synonymyby SEN- Perhaps the mismatching ofthe sexes ofZ. cal- GLET2004). lidus confused Simon in 1878, because in 1914, he Zelotessemirufus-,Senglet,2011:518,figs2-17,74(syno- described the species again from the south ofFrance nymwith Z. ruscinensis). as Z. ruscinensis. Recently, this latter species was re- Typematerial described by Senglet (2004);who further pointed Holotypemale ofProsthesima(= Zelotepcallidafrom Cor- sica,Ajaccio (MNHNP,notexamined). out that Zelotes adolescentulus Denis, 1952, Z. lugens Holotype female ofProsthesima (= Zelotes) semirufa from Denis, 1941 and Z massiliensis Soyer, 1967 are its Spain,Baleares,Menorca,Braun leg.; notexamined,una- juniorsynonyms.Allofthesetaxanowenterintothe vailablein ZMB. synonymyofZelotescallidus. HolotypefemaleofZeloteslugensfrom France,Var,Vallon Distribution dePortCros (MNHNP,examined). Recorded from Morocco, Algeria, Spain, Portugal, Holotype male of Zelotes adolescentulus from Morocco, France and Italy. DRENSKY (1915) cited the spe- Skhirat(MNHNP,examined). cies erroneouslyfrom Bulgaria; itwas in fact Zelotes Comparativematerialexamined W caucasius. SPAIN. Caceres: Torrejon el Rubio (N 39°46’15” . SpidersdescribedbyLKochfromtheBalearicIslands 13 Zelotesplumiger (L. Koch, 1882) (Fig.36) ProsthesimaplumigeraL.Koch, 1882: 630,pi.20,figs 12, 13. Typematerial HolotypemaleofProsthesima(=ZeloteP) plumigera from Spain, Baleares, Mal- lorca, Ses Prat de San Jordi, end of April, Schaufuss leg.; not examined, unavailablein ZMB. Comments According to KOCH (1882), this mm species measures 5 and the prosoma is brownish black, the abdomenyellowishbrown,thelegs reddishbrownwithredfemoraand thespinneretsyellowishbrown.The prosoma is shiny, and coveredwith long,plumosehairs,hencepresum- ably the specific name 'plumiger Kochsfigure (Fig.36) showsapalp with thebulb taperingto the top, a terminaltooth (the embolus ?) and aretrolateral,curvedapophysis(ter- minalapophysis?).Theaffinitiesof the species are presentlyunknown, but most probably it does not be- longinthegenus Zelotes. Gnaphosa artaensis Wunderlich, 2011 from Mallorcahasaverysimilarpalpbut there are not enough elements to consider it conspecific with Zelotes Figs.24-38.Figs.24-26:OzyptilafurculaL.Koch,1882.24.MalepalpofOzyptHafurculain plumiger. Further investigation on Koch (1882);25.Malepalp,ventralview;26.Idem,lateralview.Figs.27-28.Wr7/?/o- the island could reveal the identity drassusparvicorpus{Roewer,1951).27.MalepalpofDrassusparvulus\n Koch(1882).28. ofKoch’s species. MalepalpofHaplodrassusparvicorpus,specimenfromTarifa.Figs.29-35.Zelotescallidus (Simon,1878).29.EpigyneofZelotessemirufus\n Koch (1882);30.EpigyneofZelotes Distribution ra/Z/r/us,specimenfromTalavan;31.Vulva;32.MalepalpofZelotescallidus\oSimon Onlyknownfromthe typelocality. (1914,fig.298);33.Malepalp,detail;34.Malepalp,ventralview;35.Malepalp, lateralview;Fig.36.MalepalpofZelotesplumigerLKoch,1882in L.Koch(1882).Figs. Family Salticidae 37-38Phlegrabresnieri(Lucas,1846).37.MalepalpofPhlegrasimoniln L.Koch(1882); 38.MalepalpofPhlegrabresnieriin Metzner(1999). Phlegrabresnieri(Lucas,1846) (Figs 37-38) Comments PhlegrasimoniL. Koch, 1882: 667, pi. 21, fig. 37 (descr. female);newsynonymy. The holotype female could be examined.The dorsal Typematerial stripes on the prosoma and abdomen have disap- Holotype female ofPhlegra simoni from Spain, Baleares, peared,buttheclypeusis denselycoveredwithwhite Minorca, Riera near Palma, 30 May 1872, Schaufuss leg. hairs.The long, threadlike embolus is clearlyvisible (ZMB 7932);examined. and its placement in the genus Phlegra is thus con- Comparativematerialexamined firmed. According to KOCH (1882), Phlegra simoni GREECE.Attiki:EnoeeruinsnearMarathon(N38°9’13” differsfromP.bresnieriinthemalepalpaltibiawhich E23°56’33”),180m,3males,stonesinmaquis,19.IV.2000, bearsblackhairs. Examinationofsome specimensof R. Bosmansleg. (CRB). P.bresnierifrom our collection showthat the femora 14 R.Bosnians&J.vanKeer andpatellae havewhite hairs,butonlyblackones on fully studied and three species could be recognized. the tibiae (cfr. SiMON, 1876: 121: “Patte-machoire, Ozyptilafurcula was recognized and redescribed. au moins la patella et le femur, garnie en dessus de Philodromusvegetusand Theridionelimatumwererec- polls blancs”). As the palps are otherwise identical ognizedasjuniorsynonyms ofThanatusvulgarisand (Figs 37, 38), the two species are here synonymised. Enoplognatha diversa respectively. Lycosafraissei, L. Distribution insulana^L.simplexandPardosatenuipescouldnotbe Southern Europe to Azerbaijan, Ivory Coast, Tan- recognizedandareleftintheiroriginalgenus.Finally zania. Lycosa misella is declared a nomen nudum, since the descriptionwas based on ajuvenile specimen. Conclusions Taxonomic studies by earlier arachnologists are of Acknowledgements variablequality,butuntilthereisprooftothecontrary, Benjamin Nitsche andJason Dunlop (ZMB) are thanked theyallhavetobeconsideredvaluable. Someauthors fortheloanofthematerialdescribedbyL. Kochfromthe working on Mediterranean spiders in the past - for Baleares.AntonyRussell-Smithisthankedforafinalreview instance (but not exclusively) LuCAS (e.g. 1846), O. ofthepaperandHeikoMetznerandJörgWunderlichfor P.-CAMBRIDGE(e.g. 1872,1876) and KULCZYNSKI permissiontouse theirfigures. (e g. 1908, 1911) - presented detailed descriptions References accompanied by excellent figures ofthe general ap- pearanceandsexualorgans,makingidentificationvery BOSMANS R. &J. Van Keer (1999): The genus Eno- plognatha Pavesi, 1880 in the Mediterranean region easilypossible.Inmostcases,however,examinationof (AraneaeiTheridiidae).-BulletinoftheBritisharach- the type material is necessaryto come to a definitive nologicalSociety11:209-241 cdeosnccrliupstiioonn.sObtuhteprreasuetnhtoerds ngoavfeigluornegs,avtearlyl;dleitkaeilfeodr BosAsleBlEaReDrIs(J.2,00C9.):UrOonnetsh,eJM.eAd.iBtearrrraineenatnosspe&cieJs.Mo.f instance PAVESI (e.g. 1880, 1884) andthe numerous Trachelinae (Araneae, Corinnidae) with a revision of papers byTHORELL (e.g. 1875) and STRAND (e.g. TrachelasL.Koch,1872ontheIberianPeninsula.-Jour- 1906, 1908). Here, recognition ofthe species is not nalofArachnology37: 15-38-doi: 10.1636/A08-33.1 possible without examining the original type mate- Braun R. (1965): Beitrag zu einer Revision derpaläark- rial. However, even ifthese species are not instantly tischenArtenderPhilodromus«z/r^o/z^r-Gruppe(Arach., recognizable,theirnamesremainvaliduntilhasbeen Araneae). I. Morphologisch-systematischer Teil. - statedinapublicationthatthetypesarenotavailable; Senckenbergianabiologica46: 369-428 like for instance many types ofStrand destroyed in CAPORIACCO L.di(1936):Aracnidifezzanesiraccoltidal the lastWorldWar. profG. Scorteccinel 1934-XII.(MissionedeUaR. So- In the case ofthe study by L. KOCH (1882) on cietageografica).-AttidellaSocietäitalianadiscienze naturali 75: 67-93 gthoeodspiddrearwsinogfst,heinBamlaeanryiccaIsselsanmdsa,ktihnegpiadpeenrtiifnicclautdieosn DECAEA.&PCardoso(2006):Iberesia,anewgenusof trapdoor spiders (Araneae,Nemesiidae) from Portugal possible. Only ten ofthe 28 described species were & Spain.-RevistaIbericadeAracnologfa 12: 3-11 redescribedpreviously,resultinginthreevalid names and seven synonyms. Type material of ten of the DendiesPJo.r(t1-9C35r)o:sA(dVdairt)i.o-nsAnänlaalfeausnedearlaacShoncoileotgeiqdu’eHidsetofiirlee remaining 18 species could be examined by us. Five naturelledeToulon 19: 114-122 ofthese ninespecies namesarevalid: Cheiracanthium Denis J. (1952): Quelques araignees de file de Cabrera occcidentale,Dysdera mordax^Liocranum inornatum n. et remarques sur la fauna des Baleares. - Bulletin de comb,andHaplodrassusparvicorpisn.comb.Fourout ITnstitut royal de Sciences naturelles de Belgique ofthe nine species appeared to be synonyms ofspe- 28(25): 1-4 ciesdescribedearlier.Erigonemarina^Lycosaperspicax, Denis (1956): Spiders collected in French Morocco by J. Lycosa subhirsuta and Phlegra simoni are the junior the Durham Colleges Expedition Club.-Proceedings synonyms of Oedothoraxfuscus^ Arctosafuholineata, oftheZoologicalSocietyofLondon126:275-281-doi: Arctosa lacustris and Phlegrafasciata respectively. Fi- 10.1111/j.l096-3642.1956.tb00437.x nally, Trochosulaconspersaisretransferedtoitsoriginal DRENSKY P. (1915): Araneides nouveaux ou peu connus genus,Lycosa^where it awaits further study. de Bulgarie. - Spisanie na bulgarskata Akademiya na Of the eight species whose types could not Naikite. 12: 141-176 be examined, the drawings of L. Koch were care-