© The Author, 2018. Journal compilation © Australian Museum, Sydney, 2018 Records of the Australian Museum (2018) Vol. 70, issue number 1, pp. 1–111. ISSN 0067-1975 (print), ISSN 2201-4349 (online) https://doi.org/10.3853/j.2201-4349.70.2018.1645 urn:lsid:zoobank.org:pub:82A58B37-13FE-4EA8-AFF2-E954CDBEFD69 Iorgu Petrescu orcid.org/0000-0002-7654-4675 On the Family Nannastacidae (Crustacea, Cumacea) from the Australian Museum Collection Iorgu Petrescu “Grigore Antipa” National Museum of Natural History, Kiseleff 1, Bucharest 011341, Romania [email protected] Abstract. 108 species and 10 genera of Nannastacidae were identified in collections of the Australian Museum from samples around Australia; 48 are described as new species: Campylaspides stanescui, Campylaspis adami, C. adelae, C. aureliani, C. berentsae, C. chisamerai, C. cursaruae, C. dumitrumurariui, C. elenaionuti, C. gabrielamircea, C. georgetae, C. gherasimi, C. guerragarciai, C. hangiuae, C. heardi, C. keablei, C. lowryi, C. marinescui, C. matacheae, C. mioarae, C. oanae, C. oanalexandru, C. oneai, C. panai, C. paucai, C. popai, C. radui, C. roccatagliatai, C. sienkiewiczi, C. stanae, C. udrescui, C. vasilescui, C. wilsoni, Nannastacus papadopoli, Procampylaspis capraruae, P. corberai, Scherocumella weinbergae, Scherocumella wittmanni, Schizocuma antipai, Schizocuma bacescui, Schizotrema dumitriui, Schizotrema leswatlingi, Schizotrema radui, Schizotrema zimmeri, Styloptocuma anae, Styloptocuma angelae, Styloptocuma aurorae, and Styloptocuma halei. An additional taxon is recognized as Styloptocuma sp. but remains undescribed. Additionally, two genera— Campylaspides Fage, 1929, and Schizocuma Băcescu, 1972—and the following 15 species are newly recorded from Australia: Cumella bunakenensis Petrescu, 1995, Cumella indosinica Zimmer, 1952, Nannastacus antipai Petrescu, 1995, N. gamoi Băcescu, 1992, N. mitreai Petrescu, 1995, N. nyctagineus Gamô, 1962, N. stebbingi defformis Fage, 1945, N. wisseni Petrescu, 1997, Procampylaspis andamanensis Watling & Angsupanich, 2002, P. bispinosa Ledoyer, 1988, Schizotrema bidens Fage, 1945, Schizotrema bifrons Calman, 1911, Schizotrema depressum Calman, 1911, Schizotrema sakaii Gâmo, 1964, and Schizotrema sordidum Calman, 1911. Keywords. Crustacea; Cumacea; Nannastacidae; taxonomy; new species; new records. Petrescu, Iorgu. 2018. On the family Nannastacidae (Crustacea, Cumacea) from the Australian Museum collection. Records of the Australian Museum 70(1): 1–111. https://doi.org/10.3853/j.2201-4349.70.2018.1645 2 Records of the Australian Museum (2018) Vol. 70 Contents Introduction ...............................................................................................................................................................4 Material and methods ..................................................................................................................................................4 Genus Campylaspides Fage, 1929 ..............................................................................................................................4 Key to species of Campylaspides from Australian waters ....................................................................................4 Campylaspides stanescui sp. nov. ..................................................................................................................4 Genus Campylaspis Sars, 1865 ...................................................................................................................................5 Key to species of Campylaspis from Australian waters .......................................................................................5 Campylaspis adami sp. nov. ..........................................................................................................................8 Campylaspis adelae sp. nov. .........................................................................................................................8 Campylaspis anae Petrescu, 2006 ..................................................................................................................9 Campylaspis angelae Petrescu, 2006 .............................................................................................................9 Campylaspis aspera Hale, 1945 .....................................................................................................................9 Campylaspis aureliani sp. nov. .....................................................................................................................9 Campylaspis australiensis Petrescu, 2006 ...................................................................................................10 Campylaspis berentsae sp. nov. ..................................................................................................................10 Campylaspis chisamerai sp. nov. ................................................................................................................10 Campylaspis cursaruae sp. nov. ...................................................................................................................11 Campylaspis dumitrumurariui sp. nov. .......................................................................................................12 Campylaspis echinata Hale, 1945 ................................................................................................................12 Campylaspis edenensis Petrescu, 2006 ........................................................................................................12 Campylaspis elenaionuti sp. nov. ................................................................................................................12 Campylaspis gabrielamircea sp. nov. .........................................................................................................13 Campylaspis georgetae sp. nov. ...................................................................................................................14 Campylaspis gherasimi sp. nov. ..................................................................................................................15 Campylaspis guerragarciai sp. nov. ...........................................................................................................15 Campylaspis hangiuae sp. nov. ...................................................................................................................16 Campylaspis heardi sp. nov. ........................................................................................................................16 Campylaspis johnstoni Hale, 1937 ...............................................................................................................17 Campylaspis keablei sp. nov. ......................................................................................................................17 Campylaspis latidactyla Hale, 1945 ............................................................................................................19 Campylaspis lowryi sp. nov. ........................................................................................................................19 Campylaspis marinescui sp. nov. ................................................................................................................19 Campylaspis matacheae sp. nov. ................................................................................................................20 Campylaspis minor Hale, 1945 ....................................................................................................................21 Campylaspis mioarae sp. nov. ....................................................................................................................21 Campylaspis nowrae Petrescu, 2006 ...........................................................................................................21 Campylaspis oanae sp. nov. ........................................................................................................................22 Campylaspis oanalexandru sp. nov. ............................................................................................................22 Campylaspis oneai sp. nov. .........................................................................................................................23 Campylaspis panai sp. nov. .........................................................................................................................23 Campylaspis paucai sp. nov. .......................................................................................................................24 Campylaspis pileus (Foxon, 1932) ...............................................................................................................25 Campylaspis popai sp. nov. .........................................................................................................................25 Campylaspis pustulosa Hale, 1945 ..............................................................................................................25 Campylaspis radui sp. nov. ..........................................................................................................................25 Campylaspis rectangulata Petrescu, 2006 ...................................................................................................26 Campylaspis roccatagliatai sp. nov. ...........................................................................................................26 Campylaspis roscida Hale, 1945 ..................................................................................................................27 Campylaspis sienkiewiczi sp. nov. ..............................................................................................................27 Campylaspis similis Hale, 1945 ...................................................................................................................28 Campylaspis stanae sp. nov. .......................................................................................................................28 Campylaspis tasmaniensis Petrescu, 2006 ...................................................................................................28 Campylaspis thetidis Hale, 1945 ..................................................................................................................28 Campylaspis thompsoni Hale, 1945 .............................................................................................................28 Campylaspis trisulcata Petrescu, 2006 ........................................................................................................29 Campylaspis udrescui sp. nov. ....................................................................................................................29 Campylaspis uniplicata Hale, 1945 .............................................................................................................29 Campylaspis unisulcata Hale, 1945 .............................................................................................................29 Campylaspis vasilescui sp. nov. ...................................................................................................................30 Campylaspis wardi Băcescu, 1992...............................................................................................................30 Campylaspis wilsoni sp. nov. .......................................................................................................................31 Campylaspis sp. ...........................................................................................................................................32 Genus Cumella Sars, 1865 ........................................................................................................................................32 Key to species of Cumella from Australian waters ............................................................................................32 Subgenus Cumella Sars, 1865 ............................................................................................................................33 Cumella (Cumella) bunakenensis Petrescu, 1995 ........................................................................................33 Cumella (Cumella) cana Hale, 1945 ............................................................................................................33 Petrescu: New Cumacea in the Australian Museum 3 Subgenus Cumewingia Băcescu, 1971 ...............................................................................................................33 Cumella (Cumewingia) hispida Calman, 1911 ............................................................................................33 Cumella (Cumewingia) indosinica Zimmer, 1952 .......................................................................................33 Genus Nannastacus Bate, 1865 ................................................................................................................................35 Key to species of Nannastacus from Australian waters (present study) ............................................................34 Nannastacus antipai Petrescu, 1995 ............................................................................................................35 Nannastacus gamoi Băcescu, 1992 ..............................................................................................................35 Nannastacus gibbosus Calman, 1911...........................................................................................................35 Nannastacus hanseni Calman, 1905 ............................................................................................................35 Nannastacus inconstans Hale, 1945 ............................................................................................................35 Nannastacus inflatus Hale, 1945 ..................................................................................................................36 Nannastacus johnstoni Hale, 1945 ...............................................................................................................37 Nannastacus mitreai Petrescu, 1995 ............................................................................................................37 Nannastacus nyctagineus Gamô, 1962 ........................................................................................................37 Nannastacus papadopoli sp. nov. ................................................................................................................37 Nannastacus stebbingi defformis Fage, 1945 ..............................................................................................37 Nannastacus subinflatus Hale, 1945 ............................................................................................................37 Nannastacus wisseni Petrescu, 1997 ............................................................................................................37 Genus Procampylaspis Bonnier, 1896 ......................................................................................................................38 Key to species of Procampylaspis from Australian waters ................................................................................38 Procampylaspis andamanensis Watling & Angsupanich, 2002 ...................................................................38 Procampylaspis australiensis Petrescu, 2006 ..............................................................................................38 Procampylaspis bispinosa Ledoyer, 1988 ....................................................................................................38 Procampylaspis capraruae sp. nov. ............................................................................................................38 Procampylaspis corberai sp. nov. ...............................................................................................................39 Procampylaspis poorei Petrescu, 2006 ........................................................................................................39 Procampylaspis sordida Hale, 1945 ............................................................................................................39 Genus Scherocumella Watling, 1991 ........................................................................................................................40 Key to species of Scherocumella from Australian waters ..................................................................................40 Scherocumella clavata (Hale, 1945) ............................................................................................................40 Scherocumella lima (Hale, 1936) .................................................................................................................40 Scherocumella nasuta (Zimmer, 1914) ........................................................................................................40 Scherocumella nichollsi (Hale, 1949) ..........................................................................................................40 Scherocumella sheardi (Hale, 1945) ............................................................................................................40 Scherocumella vieta (Hale, 1949) ................................................................................................................41 Scherocumella weinbergae sp. nov. .............................................................................................................41 Scherocumella wittmanni sp. nov. ...............................................................................................................41 Genus Schizocuma Băcescu, 1972 ............................................................................................................................42 Key to species of Schizocuma from Australian waters ......................................................................................42 Schizocuma antipai sp. nov. ........................................................................................................................42 Schizocuma bacescui sp. nov. .....................................................................................................................43 Genus Schizotrema Calman, 1911 ............................................................................................................................44 Key to species of Schizotrema from Australian waters ......................................................................................44 Schizotrema aculeatum Hale, 1936 ..............................................................................................................44 Schizotrema bidens Fage, 1945 ....................................................................................................................45 Schizotrema bifrons Calman, 1911 ...............................................................................................................45 Schizotrema depressum Calman, 1911 .........................................................................................................45 Schizotrema dumitriui sp. nov. .....................................................................................................................45 Schizotrema leopardinum Hale, 1949 ..........................................................................................................45 Schizotrema leswatlingi sp. nov. .................................................................................................................46 Schizotrema nudum Tafe & Greenwood, 1996 ............................................................................................46 Schizotrema radui sp. nov. ..........................................................................................................................46 Schizotrema sakaii Gamô, 1964 ...................................................................................................................47 Schizotrema sordidum Calman, 1911 ...........................................................................................................47 Schizotrema zimmeri sp. nov. .......................................................................................................................47 Genus Styloptocuma Băcescu & Muradian, 1974 .....................................................................................................48 Key to species of Styloptocuma from Australian waters ....................................................................................48 Styloptocuma anae sp. nov. ..........................................................................................................................48 Styloptocuma angelae sp. nov. .....................................................................................................................49 Styloptocuma aurorae sp. nov. .....................................................................................................................49 Styloptocuma halei sp. nov. ..........................................................................................................................50 Styloptocuma sp. ..........................................................................................................................................50 Genus Vemacumella Petrescu, 2001 .........................................................................................................................51 Vemacumella bacescui Petrescu, 2006 .........................................................................................................51 Figures 1–58 .............................................................................................................................................................52 References ............................................................................................................................................................110 Supplementary data (collection data) .......................................................................................................................111 4 Records of the Australian Museum (2018) Vol. 70 Introduction (MGAB) with CUM prefix. Holotype collection data are given in full here. Specimen collection data for all material The most important studies on Australian Cumacea were examined are dervied from the AM database (EMu) and made by the famous specialist, Herbert M. Hale, who dealt given, in full, in a supplementary dataset uploaded at figshare with 90% of Australian fauna between 1936 and 1949. More and permanently and freely (CC BY 4.0) available via DOI recently, Tafe & Greenwood (1996), Gerken (2013) and (see Petrescu [2018] in the list of references). Distribution Petrescu (2004a,b; 2006) brought new data on Bodotriidae, data in the present work is abbridged and summarized; it is Diastylidae, Nannastacidae and Ceratocumatidae. The first derived from EMu-output May 2017 (Petrescu, 2018).The species known from this family from Australia, however, Australian states and territories are abbreviated as follows: were Cumella (Cumella) cyclaspoides, Cumella (C.) gibba, Queensland, QLD; New South Wales, NSW; Victoria, VIC; C. (C.) michaelseni, Scherocumella nasuta, S. n. camelus, Tasmania, TAS; South Australia, SA; Western Australia, S. n. nasutus, described by Zimmer (1914) from Shark WA; and the NT, NT. The Arafura Sea lies on the Sahul Bay, Western Australia and also Cumella (C.) hispida Shelf between New Guinea (Indonesian West Papua) and Calman, 1911. Foxon (1932) described Campylaspis northern Australia; the Coral Sea lies off the Great Barrier pileus from the Great Barrier Reef. Hale added 32 species Reef, between northeastern Australia, Papua New Guinea of Nannastacidae to the Australian list, 29 being new to and several South Pacific countries; the Tasman Sea lies science. Băcescu (1991) described Campylaspis wardi between Australia and New Zealand. and Tafe and Greenwood, 1996 added a new species of Schizotrema. Stoddart & Lowry (2003) included 38 species Genus Campylaspides Fage, 1929 and 2 subspecies of Nannastacidae in their list of Cumacea from Australia. Petrescu (2006) described 29 new species, Diagnosis. Rudimentary eye lobe. Dactylus of maxilliped increasing the number of species to 70. 2 like a trident. Female with 3 exopods and male with 5 Material from 404 samples was sorted up to family range exopods. and partially identified in Australian Museum and finally in the “Grigore Antipa” National Museum of Natural History, Campylaspides stanescui sp. nov. Bucharest, Romania. One hundred eight species and 10 genera of Nannastacidae were identified in these collections, Fig. 1 49 species are described as new, one genus, Schizocuma Băcescu, and other 15 species are added to Australian waters. Holotype subadult ♀, 3.19 mm, P.88258, Tasman Sea, Lord A total of 133 species of Nannastacidae is known now from Howe Rise, -27.983° 162.859°, 1250 m, epibenthic sled, waters of Australia. coarse sediment with pumice, 6 May 1989, J.K. Lowry & party, RV Franklin, FR0589-28, in AM. Material and methods Etymology. The species is named in honour of my colleague, The material was identified to species in the Australian Dr Mihai Stănescu, specialist in Lepidoptera, as a sign of Museum, Sydney. Drawings and text were compiled at sincere friendship and appreciation of his work. “Grigore Antipa” Museum, Bucharest. Drawings were made Diagnosis. Carapace almost 0.5 body length, carpus of 2nd with microscopy and camera lucida. Body length is measured maxilliped with medial apophysis, long terminal spines; between the tip of the pseudorostrum lobes and the posterior maxilliped 3 basis 0.5 entire length; dactylus 4.1 propodus margin of the last pleonite. length; uropodal peduncle 1.86 pleonite 6 length, 1.09 Specimens are registered in the Australian Museum, endopod length. Sydney (AM). All AM register numbers are with “P.” prefix and representative specimens are in the “Grigore Antipa” Description. Carapace 0.49 body length; 1.9 as long as National Museum of Natural History, Bucharest, Romania high, 1.25 as long as wide; pseudorostrum 0.22 carapace Key to species of Campylaspides from Australian waters 1 Maxilliped 3 with spines ........................................................................................................... 2 —— Maxilliped 3 without spines ...................................................................................................... 5 2 Carapace integument with spines or mamelons ........................................................................ 3 —— Carapace integument smooth ................................................ C. diva Mühlenhardt-Siegel, 2005 3 Carapace integument with mamelons ...................................................... C. grandis Fage, 1929 —— Carapace with spines ................................................................................................................. 4 4 Maxilliped 2 carpus with spines .......................... C. abyssotrucidatus Watling & Gerken, 1999 —— Maxilliped 2 carpus with setae ......................................................... C. echinata Ledoyer, 1988 5 Maxilliped 2 with spines ........................................................................................................... 6 —— Maxilliped 2 with 1 spine and 1 apophysis ................................................ C. stanescui sp. nov. 6 Maxilliped 2 carpus with 2 spines, 1 much longer ........................... C. canariensis Jones, 1973 —— Maxilliped 2 carpus with 2 short spines ................................................. C. spinifer Jones, 1973 Petrescu: New Cumacea in the Australian Museum 5 length, 3.4 ocular lobe length; ocular lobe without eyelobe; dactylus fused with terminal robust seta (Fig. 1 J). —Uropod marked anterior notch; few simple setae on anterior half (Fig. peduncle 1.86 pleonite 6 length, 1.09 endopod length, with 1 A,B). —Antenna 1 peduncle article 1 1.1 rest of article’s serrated margins; exopod 0.9 endopod length, 1 subterminal length; article 2 1.3 article 3 length; main flagellum 2.5 article simple and 1 terminal microserrate seta; endopod with short 3 length; accessory flagellum with 1 article; aesthetascs 2 terminal microserrate seta, 0.3 endopod length, 2 simple main flagellum length (Fig. 1 C). —Maxilliped 2 basis with setae medially (Fig. 1 K). 2 simple setae; ischium fused with merus; carpus 1.6 merus Remarks. The female is known for only two species of the length, second longest article, with 1 spine and 1 large genus, Campylaspides grandis Fage (1929) from the Atlantic apophysis medially, 1 simple and 1 plumose seta; propodus Ocean (Morocco) and C. stanescui sp. nov. from the Pacific 0.64 carpus length, with 1 robust spine; dactylus with 3 Ocean (Australia). The main shared character is the aspect terminal, very long, spines (Fig. 1 D). —Maxilliped 3 basis of the propodus and dactylus of maxilliped 2. It differs 0.5 entire length, with 6 plumose setae, setulated lateraly; from this species with: carapace without spiny integument, ischium 0.5 merus length, serrated medially; merus with 2 the carpus of maxilliped 2 with medial apophysis instead plumose and 4 simple setae; carpus 0.75 merus length, with of setae (unique character within the known species of the 1 plumose seta; propodus 0.6 carpus length, with 2 pappose genus), maxilliped 2 with a large apophysis on the carpus setae; dactylus 0.5 propodus length, with 3 terminal simple instead of 3 strong teeth, uropod with longer exopod, and setae; with exopod (Fig. 1 E). —Pereopod 1 basis 0.43 endopod with 2 instead of 4 medial setae. The nearest known entire length, 1 pappose and 2 plumose setae; ischium with congener, C. echinata Ledoyer, 1988 from the Indian Ocean plumose seta on medial margin; merus 4 ischium length, (Comoro Islands), shares a similar aspect of the propodus with 4 simple setae, 2 much longer setae on medial margin, and dactylus of maxilliped 2. The new species is close to 1 plumose and 1 pappose seta; carpus 1.5 merus length, with other species of the genus, having a similar propodus and serrated medial margin, with 5 simple, plumose and pappose dactylus of maxilliped 2, pereopod 2 with long and slender setae; propodus 0.8 carpus length, setulated medially, with 5 dactylus, namely C. diva Mühlenhardt-Siegel, 2005 from the simple, plumose and pappose setae; dactylus 0.7 propodus Angola Basin, C. spinifera Jones, 1973 from Recife, western length, with 3 simple subterminal and 3 terminal setae Atlantic, and C. abyssotrucidatus Watling & Gerken, 1999, shorter than dactylus; with exopod (Fig. 1 F). —Pereopod from Brazil. 2 (Fig. 1 G) basis 0.31 entire length, with 5 simple setae; ischium with 1 pappose seta; merus 3.7 ischium length, with Distribution. Tasman Sea (between Australia and New 1 pappose seta; carpus 1.8 merus and ischium length, with Zealand), Lord Howe Rise, at 1250 m depth. This is the first 1 pappose and 2 simple setae; propodus 0.3 carpus length; record of the genus Campylaspides from waters off Australia. dactylus 4.1 propodus length, with 4 short simple setae; with exopod (Fig. 1 G). —Pereopod 3 (Fig. 1 H) basis 0.5 entire Genus Campylaspis Sars, 1865 length, with 1 simple seta; ischium with 2 plumose setae; merus 1.8 ischium length, with 1 plumose seta; carpus 2.1 Diagnosis. Carapace longer than 0.4 body length; maxilliped merus length, with 1 annulate seta; propodus 0.52 carpus 1 with 3 articles; maxilliped 2 dactylus ending in 3 diverging length, with 1 annulate seta; dactylus fused with terminal spines; pereopod 2 dactylus with usual or digitiform robust seta (Fig. 1 H). —Pereopod 4 basis 0.4 entire length; extremity; female maxilliped 3 and pereopods 1 and 2 and ischium and merus with 1 plumose seta; carpus 1.3 ischium male maxilliped 3 and pereopods 1-4 with exopods. and merus length, with 1 annulate seta; propodus 0.5 carpus length, with 1 annulate seta; dactylus fused with terminal Remarks. Thirty-five species were previously known from robust seta (Fig. I). —Pereopod 5 basis 0.38 entire length; Australia (Petrescu, 2006), and 31 new species are added carpus as long as ischium and merus length, with 1 annulate herein. A total of 53 species were found in the Australian seta; propodus 0.43 carpus length, with 1 annulate seta; Museum collection. Key to species of Campylaspis from Australian waters 1 Eye lenses present ..................................................................................................................... 2 —— Eye lenses absent ..................................................................................................................... 17 2 Carapace with tubercles, spines, carinae or a lateral furrow ..................................................... 3 —— Carapace smooth, without tubercles, spines, carinae or a lateral furrow ......................................................................................................... C. aureliani sp. nov. 3 Uropodal peduncle 3 as long as its endopod ....................................... C. thompsoni Hale, 1945 —— Uropodal peduncle less than 3 as long as its endopod .............................................................. 4 4 Carapace with 2 sulci .................................................................................... C. wilsoni sp. nov. —— Carapace with 1 sulcus .............................................................................................................. 5 5 Maxilliped 3 medial margin of merus to propodus serrate ....................................................... 6 —— Maxilliped 3 medial margin of merus to propodus smooth .................................................... 16 6 Maxilliped 3 large ..................................................................................................................... 7 —— Maxilliped 3 slender ................................................................................................................ 14 6 Records of the Australian Museum (2018) Vol. 70 7 Carapace lateral sulcus with a transverse ridge ......................................... C. berentsae sp. nov. —— Carapace lateral sulcus without transverse ridge ...................................................................... 8 8 Carapace with circular sulcus .......................................................................... C. lowryi sp. nov. —— Carapace with normal sulcus ..................................................................................................... 9 9 Pereopod 2 dactylus with tapering tip ..................................................................................... 10 —— Pereopod 2 dactylus without tapering tip ................................................................................ 11 10 Pereopod 2 dactylus 3.22 propodus length ............................... Campylaspis similis Hale, 1945 —— Pereopod 2 dactylus 2.33 propodus length ..................................................... C. adelae sp. nov. 11 Pereopod 2 with acute dactylus ............................................................................................... 12 —— Pereopod 2 with not acute dactylus ..................................................... C. latidactyla Hale, 1945 12 Uropodal endopod with 5 medial setae .................................................... C. roscida Hale, 1945 —— Uropodal endopod with 3 medial setae ...................................................... C. minor Hale, 1945 14 Pereopod 2 with long dactylus ................................................................................................ 15 —— Pereopod 2 with short dactylus ............................................................. C. wardi Băcescu, 1991 15 Maxilliped 3 merus with serrate outer margin ....................................... C. echinata Hale, 1945 —— Maxilliped 3 merus with smooth outer margin ..................................... C. pustulosa Hale, 1945 16 Carapace with a lateral carina .............................................................. C. uniplicata Hale, 1945 —— Carapace without lateral carina ........................................................... C. unisulcata Hale, 1945 17 Carapace with lateral sulcus .................................................................................................... 18 —— Carapace without lateral sulcus ............................................................................................... 43 18 Lateral sulcus with transverse ridge ........................................................................................ 19 —— Lateral sulcus without transverse ridge ................................................................................... 26 19 Carapace with small tubercles ................................................................................................. 20 —— Carapace without tubercles ..................................................................................................... 23 20 Uropodal peduncle with longitudinal ridge ....................................... C. angelae Petrescu, 2006 —— Uropodal peduncle without longitudinal ridge ........................................................................ 20 21 Pereopod 2 dactylus with pedunculate tip ....................................... C. gabrielamirceai sp. nov. —— Pereopod 2 dactylus without pedunculate tip .......................................................................... 21 22 Maxilliped 2 with long dactylus ................................................ C. tasmaniensis Petrescu, 2006 —— Maxilliped 2 with short dactylus ............................................................. C. chisamerai sp. nov. 23 Carapace with 3 lateral sulci ........................................................... C. trisulcata Petrescu, 2006 —— Carapace with 1 sulcus ............................................................................................................ 24 24 Carapace with dorsal translucent lenses ......................................... C. edenensis Petrescu, 2006 —— Carapace without dorsal translucent lenses ............................................................................. 25 25 Pereopod 2 dactylus with long terminal seta ............................................. C. gherasimi sp. nov. —— Pereopod 2 dactylus with short terminal seta ................................................... C. oneai sp. nov. 26 Carapace with small tubercles and spines ............................................................................... 27 —— Carapace without tubercles and spines .................................................................................... 30 27 Carapace with 2 lateral sulci ................................................................................................... 28 —— Carapace with 1 sulcus ............................................................................................................ 29 28 Uropod peduncle with 8 medial setae ...................................................... C. matacheae sp. nov. —— Uropod peduncle with 9 longer medial setae and 16 shorter ones .................................................................................................................. C. oanae sp. nov. 29 Pereopod 2 dactylus propodus length ....................................... C. australiensis Petrescu, 2006 —— Pereopod 2 dactylus propodus length ................................................ C. guerragarciai sp. nov. Petrescu: New Cumacea in the Australian Museum 7 30 Carapace with 2 lateral sulci ................................................................................................... 31 —— Carapace with 1 lateral sulcus ................................................................................................. 33 31 Pereopod 2 dactylus 3.8 propodus length .................................................... C. udrescui sp. nov. —— Pereopod 2 dactylus shorter 3.8 propodus length .................................................................... 32 32 Pereopods 3–5 dactylus fused with terminal robust seta ........................... C. georgetae sp. nov. —— Pereopods 3–5 dactylus not fused with terminal seta ............................. C. sienkiewiczi sp. nov. 33 Pereopods 3–5 with short and robust articles ..................................... C. roccatagliatai sp. nov. —— Pereopods 3–5 with long and slender articles ......................................................................... 34 34 Carapace with long pseudorostrum ....................................................... C. pileus (Foxon, 1932) —— Carapace with short pseudorostrum ........................................................................................ 35 35 Carapace with circular sulcus ........................................................................... C. radui sp. nov. —— Carapace with normal sulcus ................................................................................................... 36 36 Carapace with dorsal transverse ridges .............................................. C. nowrae Petrescu, 2006 —— Carapace without dorsal ridges ............................................................................................... 37 37 Pereopod 2 dactylus with terminal tapering tip ....................................................................... 38 —— Pereopod 2 dactylus without terminal tapering tip .................................................................. 39 38 Maxilliped 3 merus longer than 1.7 ischium length ................................................................ 40 —— Maxilliped 3 merus 1.7 ischium length ......................................................... C. keablei sp. nov. 39 Maxilliped 3 with robust articles ............................................................... C. vasilescui sp. nov. —— Maxilliped 3 with slender articles .................................................................. C. stanae sp. nov. 40 Lateral sulcus as long as carapace length ................................................................................ 41 —— Lateral sulcus shorter than carapace length ............................................................................. 42 41 Uropodal peduncle with longitudinal crest ................................................ C. hangiuae sp. nov. —— Uropodal peduncle without longitudinal crest ...................................... C. johnstoni Hale, 1937 42 Maxilliped 3 merus with a medial process ................................ C. rectangulata Petrescu, 2006 —— Maxilliped 3 merus without medial process ............................................................................ 44 43 Maxilliped merus 4 ischium length ........................................................ C. anae Petrescu, 2006 —— Maxilliped 3 merus 2.6 ischium length .......................................... C. dumitrumurariui sp. nov. 44 Carapace with tubercled integument ....................................................................................... 45 —— Carapace with smooth integument .......................................................................................... 52 45 Pereopod 2 dactylus with tapering tip ..................................................................................... 46 —— Pereopod 2 dactylus without tapering tip ................................................................................ 47 46 Uropodal endopod with 6 setae on medial margin ....................................... C. mioarae sp. nov. —— Uropodal endopod without any seta ......................................................... C. thetidis Hale, 1945 47 Pleonite 5 with a transverse constriction ................................................... C. aspera Hale, 1945 —— Pleonite 5 without transverse constriction ............................................................................... 48 48 Maxilliped 3 carpus, 2nd longest article, with special medial setae ................................................................................................................ C. heardi sp. nov. —— Maxilliped 3 carpus shorter than merus .................................................................................. 49 49 Pereopod 2 dactylus with long terminal seta ........................................................................... 50 —— Pereopod 2 dactylus with short terminal seta ................................................. C. paucai sp. nov. 50 Uropodal endopod with 5 medial setae ........................................................... C. adami sp. nov. —— Uropodal endopod with less than 5 medial setae .................................................................... 51 51 Maxilliped 3 merus with a medial process .............................................. C. marinescui sp. nov. —— Maxilliped 3 merus without medial process ..................................................... C. panai sp. nov. 8 Records of the Australian Museum (2018) Vol. 70 52 Pereopod 2 dactylus with pappose setae ................................................. C. elenaionuti sp. nov. —— Pereopod 2 dactylus with simple setae .................................................................................... 53 53 Carapace with prominent anteroventral corner ................................... C. oanalexandru sp. nov. —— Carapace with hardly visible anteroventral corner .................................................................. 54 54 Uropodal endopod with 7 medial setae ............................................................ C. popai sp. nov. —— Uropodal endopod with 4 medial setae ..................................................... C. cursaruae sp. nov. Campylaspis adami sp. nov. 1 simple and 1 annulate seta; propodus 0.3 carpus length, with 1 annulate seta; dactylus fused with terminal robust Fig. 2 seta (Fig. 2 H). —Uropod peduncle 2.65 pleonite 6 length, with serrated margins, 2.57 endopod length; exopod as Holotype subadult ♀, 4.0 mm, P.64741, Australia, NSW, long as endopod, with 3 simple, 1 subterminal simple and east of Port Jackson, -33.87° 151.38°, 80 m, epibenthic sled, 1 terminal seta 0.77 exopod length; endopod with serrated shelly mud, 11 Dec 1980, R.T. Springthorpe, FRV Kapala, margins, with 5 simple and 1 terminal robust simple seta K80-20-11, in AM. 0.66 endopod length (Fig. 2 I). Etymology. The species is dedicated to my colleague Dr Remarks. Campylaspis adami sp. nov. is closely related to Costică Adam, specialist in Malophaga, as a sign of highly C. roscida Hale, 1945 with the integument of the carapace appreciation of his work. having with tiny tubercles, and being opaque in the new Diagnosis. Carapace with small tubercles, eyeless ocular species (glassy in Hale’s), and the uropodal endopod with lobe; robust maxilliped 3 basis 0.68 rest of article’s length; 5 setae. The new species differs in the basis of maxilliped merus to carpus with teeth medially; pereopod 1 basis 0.68 3 lacking setae on medial margin, pereopod 1 with more pereopod length; pereopod 2 basis 0.7 pereopod length, numerous setae, and a longer uropodal peduncle. dactylus 2.8 propodus length, with terminal simple seta; Distribution. Australia: NSW—east of Port Jackson, at 80 uropodal peduncle 2.65 pleonite 6, 2.57 endopod length. m depth. Description. Carapace 0.4 entire length, with small tubercles; 1.38 as long as high; 1.5 as long as wide; Campylaspis adelae sp. nov. pseudorostrum 0.2 carapace length; small antennal notch; Fig. 3 smooth ventral margin (Fig. 2 A, B). —Maxilliped 3 basis 0.68 rest of article’s length, setose medially, 1 pappose seta; Holotype adult ♂, 3.46 mm, P.64692, Australia, NSW, Jervis basis to propodus serrated medially; ischium 0.28 basis Bay, off Moona Moona Creek, -35.0483° 150.6867°, 8 m, length; merus 2.4 ischium length, second longest article, airlift, sediment, 17 Dec 1982, P.B. Berents, NSW 262, in with 9 setulae, 3 teeth and 1 pappose seta; carpus 0.22 merus AM. Paratype: 1♂, P.65553, Australia, WA, North West length, with 2 setulae and 1 pappose seta; propodus 0.94 Shelf, -19.94° 117.8983°, 44 m, 25 Jun 1983, CSIRO North- carpus length, with 4 pappose setae; dactylus 0.4 propodus West Shelf project, FRV Soela, 03-B2-S | 0383 B02 S, in AM. length, with 3 simple setae; with exopod (Fig. 2 C). — Pereopod 1 basis 0.62 rest of article’s length, with 2 pappose Etymology. The species is dedicated in honour of my setae; ischium 0.14 basis length, with 1 tooth and 1 pappose beloved mother, Adela, as a sign of gratitude and love for her seta; merus, 2nd longest article of pereopod, 3.6 ischium devotion shown to me during my life and especially during length, with 7 pappose and 3 simple setae; carpus 0.83 my medical problems. merus length, with 6 simple and 4 pappose setae; propodus Diagnosis. Carapace 0.4 body length; ocular lobe with 3 0.76 carpus length, with 4 simple and 4 pappose setae; large lenses; pereopod 2 dactylus 2.2 propodus length, with dactylus 0.52 propodus length, with 6 simple setae; with digitiform tip; uropodal peduncle 2.5 pleonite 6 length, with exopod (Fig. 2 D). —Pereopod 2 basis 0.7 rest of article’s 5 plumose, 2.46 endopod length; endopod 1.4 exopod length, length, with 2 plumose setae; ischium 0.1 basis length; endopod with 9 microserrate setae. merus 3 ischium length, with 1 simple and 2 pappose setae; carpus 1.6 merus length, with 2 simple and 1 plumose seta; Description. Carapace 0.4 body length; 1.8 as long as high; propodus 0.4 carpus length, with 1 simple seta medially; large lateral sulcus, 0.85 of carapace length, almost straight dactylus 2.8 propodus length, with 8 simple; with exopod dorsally; ocular lobe with 3 large lenses; pseudorostrum (Fig. 2 E). —Pereopod 3 basis 1.8 rest of article’s length; 0.38 carapace length (Fig. 3 A). —Maxilliped 3 with large merus 1.14 ischium length, with 1 plumose seta; carpus 1.1 basis, 0.68 rest of article’s length, with 5 plumose setae; merus length, with 1 plumose and 1 annulate seta; propodus merus 2.7 ischium length, with 3 plumose setae; carpus 0.4 carpus length, with 1 annulate seta; dactylus fused with 0.37 merus length, with 1 plumose seta; propodus 1.1 carpus terminal robust seta (Fig. 2 F). —Pereopod 4 basis 0.54 rest length; dactylus 0.33 propodus length, with 4 terminal of article’s length; ischium with 1 simple seta; merus 1.6 simple setae; with exopod (Fig. 3 B). — Pereopod 1 basis ischium length, with 1 plumose medially; carpus 2.5 merus 0.8 rest of article’s length, with 2 plumose setae; merus, 2nd length, with 1 simple and 1 annulate seta; propodus 0.17 longest article, 2.25 ischium length, with 4 plumose and carpus length, with 1 annulate seta; dactylus fused with 2 simple setae; carpus 0.8 merus length, with 2 plumose terminal robust seta (Fig. 2 G). —Pereopod 5 basis 0.16 setae; propodus 0.7 carpus length; dactylus 0.8 propodus rest of article’s length; merus 1.5 ischium length; ischium length, with 4 terminal simple setae; with exopod (Fig. 3 and merus with 1 simple seta; carpus 3.3 merus length, with C). —Pereopod 2 basis 0.7 rest of article’s length, with 2 Petrescu: New Cumacea in the Australian Museum 9 plumose setae; merus 0.4 ischium length, with 2 plumose Campylaspis aureliani sp. nov. setae; carpus 1.5 merus length, with 1 simple and 4 plumose setae; dactylus 2.2 propodus length, with 11 simple setae and Fig. 4 digitiform tip; with exopod (Fig. 3 D). —Pereopods 3, 4 with Holotype subadult ♂, P.65051, Australia, NSW, Bass Point, decreasing basis and increasing carpus; basis to merus with -34.6° 150.9°, 35–40 m, Smith-McIntyre Grab, 3–18 Jan 1 simple seta, carpus with 2 annulate setae, propodus with 1991, The Ecology Lab for RMI/Pioneer Project, 4-217, 1 annulate seta; dactylus with 1 terminal simple seta; with in AM. exopods (Figs 3 E, F). —Pereopod 5 basis 0.6 pereopod 4 basis; basis to merus with 1 simple seta; merus 2 ischium Etymology. The species is dedicated in honour of Dr length; carpus 1.2 merus length, with 2 annulate setae; Aurelian Popescu, palaeontologist, specialist in fossil propodus 0.16 carpus length, with 1 annulate seta; dactylus mammals from the Oltenia Museum, Museum of Natural 0.6 propodus length with 1 terminal simple seta (Fig. 3 Sciences, Craiova, as a sign of deep respect and friendship. G). —Uropod peduncle 2.5 pleonite 6 length, 2.5 endopod Diagnosis. Carapace, 0.5 body length, small eyelobe with length with 5 plumose setae and serrate margins; endopod 3 lenses; maxilliped 3, robust; merus 2.7 ischium length; 1.4 exopod length; exopod with 2 simple setae; endopod pereopod 2 dactylus 3.2 propodus length, with tapering end; with 9 microserrate and 1 terminal simple seta (Fig. 3 H). uropodal peduncle 2.3 pleonite 6 length, 2.3 endopod length; Remarks. Campylaspis adelae sp. nov. is closely related exopod 0.8 endopod length; endopod with 4 setae. to C. minor Hale, 1945, but with maxilliped 3 with 3 vs. Description. Carapace 0.5 body length; smooth integum ent, 2 plumose setae on the basis; no setae on the ischium and 2 as long as high, 1.25 as long as wide; pseudorostrum 0.25 merus vs. numerous short setae; dactylus with 4 vs. 2 terminal carapace length; small antennal notch; smooth ventral margin; setae on dactylus; pereopod 1 with 2 plumose setae on basis small eyelobe with 3 lenses; siphon 0.25 carapace length vs. none, dactylus with 3 simple terminal short setae vs. 2 (Fig. 4 A,B). —Antenna 1 with robust articles; peduncle plumose long and 1 simple short seta; pereopod 2 with 2 setae article 1 0.8 rest of articles; article 2 1.3 article 3 length; main on basis vs. none, dactylus with simple setae vs. plumose flagellum 1.4 peduncle article 3 length; accessory flagellum setae; 1.5 longer uropodal peduncle. with 1 article; aesthetascs 1.6 main flagellum length (Fig. 4 Distribution. Australia: NSW—off Moona Moona Creek, C). —Maxilliped 3 robust, large basis, 0.9 rest of article’s Jervis Bay, at 8 m depth. length, with 1 simple and 4 plumose setae; ischium 0.16 basis length, with 1 simple seta; merus 2.7 ischium length, with 1 Campylaspis anae Petrescu, 2006 plumose and 7 simple setae; carpus 0.59 merus length, with 2 simple and 1 pappose seta; propodus as long as carpus, Campylaspis anae Petrescu, 2006: 131, figs 1, 2. with 1 plumose and 3 pappose setae; dactylus 0.46 propodus Material examined: 1♂, P.88254. length, with 1 terminal simple seta; with exopod (Fig. 4 D). —Pereopod 1 basis 0.68 rest of article’s length, with 1 Distribution. Arafura Sea and Australia: WA—North West pappose, 1 simple and 1 plumose seta; merus 2.4 ischium Shelf; NT; and VIC—Gabo Island, from 1.5–210 m depth length, with 8 pappose and 2 simple setae; carpus 0.9 merus (Petrescu, 2006). The species is now reported from NSW— length, with 4 pappose and 3 simple setae; propodus 0.7 east of Port Jackson, at 132–135 m depth. carpus length, with 4 pappose and 2 simple setae; dactylus 0.6 propodus length, with 4 simple setae; with exopod (Fig. Campylaspis angelae Petrescu, 2006 4 E). —Pereopod 2 basis 0.57 rest of article’s length, with 1 simple and 2 pappose setae; ischium with 1 pappose seta; Campylaspis angelae Petrescu, 2006: 132–133, figs 3,4. merus 2.8 ischium length, with 2 pappose and 1 simple seta; Material examined: 1♀, P.88253. carpus 2 merus length, with 1 pappose and 2 simple setae; dactylus 3.2 propodus length, with 8 simple setae, with Distribution. Australia: NSW—shelf break off Nowra, at digitiform tip; with exopod (Fig. 4 F). —Pereopod 3 basis 204 m (Petrescu, 2006), and now from east of Long Reef 1.73 rest of article’s length, with 1 plumose seta; ischium Point, at 176 m depth. with 2 simple setae; merus 1.4 ischium length, with 1 simple seta; carpus 1.1 merus length, with 1 annulate seta; propodus Campylaspis aspera Hale, 1945 0.54 carpus length, with 1 annulate seta; dactylus fused with Campylaspis aspera Hale, 1945: 209, figs 45, 46.—Stoddart terminal robust seta; with exopod (Fig. 4 G). —Pereopod 4 & Lowry, 2003: 412. basis 1.5 rest of article’s length; ischium 0.6 merus length, with 1 simple seta; carpus as long as merus, with 1 simple Material examined: 1♀, P.64651; 1♀, P.88227; 1♀, and 1 annulate seta; propodus 0.6 carpus length, with 1 P.64705; 1♀, 1♂, P.64739; 3♀♀, P.88225; 1♀, P.88226; annulate seta; dactylus fused with 1 terminal robust seta; with 5♂♂, P.64742; 2♀♀, P.64743; 1♂, P.88228; 1 subadult ♂, exopod (Fig. 4 H). —Pereopod 5 basis 0.8 rest of article’s P.65041; 1♀, P.65049; 1♀, P.65050; 1♀, P.88224; 1 subadult length; ischium 0.6 merus length, with 1 simple seta; merus ♂, P.65549; 1♀, P.65559; 1♀, P.65561. with 1 simple seta; carpus 1.2 merus length, with 1 simple Distribution. Australia: NSW—Bass Point, Barrenjoey and 1 annulate seta; propodus 0.58 carpus length, with 1 Headland, Broken Bay, Long Reef Point, Merimbula, Port annulate seta; dactylus fused with terminal robust seta (Fig. Jackson, Providential Head, at 56–1115 m; VIC—Bass Strait, 4 I). —Uropod peduncle 2.3 pleonite 6 length, 2.3 endopod Gabo Island, at 118–131 m; and WA—North West Shelf, at length; exopod 0.8 endopod length, with 4 simple setae, 1 81–84 m depth (Petrescu, 2018). robust terminal seta 0.6 exopod length; endopod, 1.2 exopod 10 Records of the Australian Museum (2018) Vol. 70 length, with 2 subterminal robust and 4 microserrate setae, terminal simple seta, with 1 plumose and 5 simple setae; 1 terminal microserrate seta 0.51 endopod length (Fig. 4 J). with exopod (Fig. 5 E). —Pereopod 1 basis 0.48 pereopod length, with 4 plumose setae; basis to propodus with serrate Remarks. The new species is related to other species with margins; ischium with 1 plumose seta; merus 3. 6 ischium smooth carapace and eyelobe with lenses, C. thompsoni length, with 4 simple and 3 plumose setae; carpus 0.8 merus Hale, 1945 from Australia, C. amblyoda Gamô, 1960 from length, with 2 simple and 5 plumose setae; propodus 0.5 Japan and SE Asia, C. kiiensis Gamô, 1960 and C. totzkei carpus length, with 2 simple and 3 plumose setae; dactylus Mühlenhardt-Siegel, 2000 from Sri Lanka; only the new 0.5 propodus length, with 5 simple setae; with exopod species and C. totzkei have no dorsal keel on the carapace; (Fig. 5 F). —Pereopod 2 basis 0.37 pereopod length, with only C. aureliani has pereopod 2 dactylus with a tapering 3 plumose setae; merus 6.5 ischium length, with 2 plumose end, and the uropodal endopod has 5 setae as in C. totzkei setae; carpus 2 merus length, with 1 simple and 6 plumose (4 or 7 in the remaining species). setae; dactylus 3 propodus length, with 9 simple setae; Distribution. Australia: NSW—Bass Point, at 40 m depth. with exopod (Fig. 5 G). —Pereopods 3–5 with decreasing basis and increasing carpus; pereopods 3 and 4 basis to carpus with serrate margins; pereopod 3 with 7 plumose, Campylaspis australiensis Petrescu, 2006 3 simple and 2 annulate setae; pereopod 4 with 6 plumose, Campylaspis australiensis Petrescu, 2006: 133–134, figs 5, 6. 3 simple and 2 annulate setae; pereopod 5 with 4 simple and 2 annulate setae; dactylus with 1 terminal simple seta Material examined: 1♂, P.88198. (Fig. 5 H-J). —Uropod peduncle 1.8 pleonite 6 length, 2.3 Distribution. Australia: TAS—off Freycinet Peninsula, and endopod length; peduncle and rami with serrate margins; off Flinders Island, at 350–720 m depth. exopod 0.8 endopod length, with 2 simple setae; endopod with 4 microserrate setae (Fig. 5 K). Campylaspis berentsae sp. nov. Remarks. Campylaspis berentsae sp. nov. is similar to C. tasmaniensis Petrescu, 2006, with a tuberculate carapace and Fig. 5 lateral depression. It differs in the shorter pseudorostrum, eye Holotype subadult ♀, 5.04 mm, P.65546, Australia, WA, lobe with lenses, maxilliped 3 with shorter terminal setae, North West Shelf, -19.983° 117.856°, 42 m, epibenthic pereopod 1 with serrate margins of the articles, pereopod sled, 18 Feb 1983, CSIRO North-West Shelf project, FRV 2 with a shorter dactylus, and more robust uropods with Soela, 01-B1-S | S01-83-B1, in AM. Paratypes: 1 subadult shorter peduncles and rami. The new species also resembles ♂, P.63964; 1♀, P.63966; 1♀, P.63967; 1♀, (dissected) C. microsulcata Gerken, 2012 from New Zealand with 1 MGAB CUM 1649. similar sulcus, but has a transverse ridge on the anterior part. Etymology. The species is dedicated to Dr Penny Berents, Distribution. Australia: WA—North West Shelf; at 42–54 former Collection Manager of Marine Invertebrate m depth. collections at the Australian Museum, as a sign of high gratitude and respect for all her generous help she offered Campylaspis chisamerai sp. nov. to me during my stay in Sydney in 2003. Fig. 6 Diagnosis. Carapace lateral sulcus with 1 transverse ridge behind the basis of ocular lobe, its upper and inner margins Holotype adult ♂, 3.55 mm, P.64735, Australia, NSW, with rows of tubercles; pereonite 5 with to pleonite 4 and northeast of Long Reef, -33.68° 151.88°, 366 m, 5 Dec 1977, 6 with doubled carina, unique on pleonite 5; maxilliped 2 FRV Kapala, K77-23-03, in AM. carpus with 1 seta; uropodal peduncle 1.8 pleonite 6. Etymology. The species is named in honour of my colleague, Description. Carapace 0.56 entire body length, 2 lateral Dr Gabriel Chişamera, reknown specialist in birds and small ridges delimiting lateral sulcus, 1 transverse ridge on mammals, as a sign of sincere friendship. sulcus, behind posterior extremity of frontal lobe, its upper Diagnosis. Carapace, 0.48 entire body length, covered with and inner margins with rows of tubercles, 1 pair of small numerous tubercles placed either side of mid-dorsal ridge; tubercle on basis of frontal lobe; eyelobe with 3 lenses; maxilliped 2 with 2 small tubercles on carpus, 1 tooth on unique carina on basis of frontal lobe; pseudorostrum 0.38 distal margin of propodus, short dactylus; pereopod 2 with carapace length (Fig. 5 A,B). Double carina on pereonite 5 digitiform dactylus; uropodal peduncle 2.1 pleonite 6 length, and pleonites 1–4, 6, 1 carina on pleonite 5. —Antenna 1 equal rami. peduncle article 1 0.8 rest of article’s length; article 2 1.1 article 3 length; main flagellum 0.9 article 3 length, with Description. Carapace, 0.48 body length, 1.86 as long as 3 articles; accessory flagellum with 1 article; aesthetascs high, 1.58 as long as wide; lateral sulcus without posterior 1.6 main flagellum length (Fig. 5 C). —Maxilliped 2 basis limit, transverse ridge in anterior part; carapace covered with fused with ischium, with 1 long plumose seta; merus with numerous tubercles placed on each side of the mid-dorsal 1 long plumose seta; carpus 1.25 merus length, with 1 tooth ridge; small antennal notch; smooth ventral margin; very and 1 simple seta; propodus 2nd longest article, 1.3 carpus small eyelobe without lenses; pseudorostrum 0.2 carapace length with 1 tooth and 1 short simple seta longer than the length (Fig. 6 A,B). —Pleonites each with a mid-dorsal ridge, 3 terminal teeth of dactylus (Fig. 5 D). —Maxilliped 3 basis lateral ridge on each side of pleonites 1–5, pair of small 1.1 rest of article’s length, with 5 plumose setae; merus, 3 tubercles on each side of pleonites 5 (Fig. 6 B). —Antenna carpus length, with serrate margins, 1 long plumose seta; 1 peduncle article 1 1.25 rest of article’s length; article 2 carpus as long as propodus, strongly serrate margins, with 1.1 article 3 length; main flagellum 2.7 article 3 length; 3 plumose setae; dactylus 0.5 propodus length, with long accessory flagellum with 1 article; aesthetascs 0.3 main