^Ssou American Fern Journal 92(l):l-9 (2002) Rl ^OTANl (^AL May ^ 4 200^ Milton Tryon, Rolla Obituary: Jr ^^^Of/v (1916-2001) ^'^RARY ^ .1' #' r -t n i Gastony Gerald J. 47405-3700 Bloomington, IN Indiana University, Department of Biology, Barrington David S. VT 05404-0086 Vermont, Burlington, University of Department of Botany, Con ant David S. VT 05851 Lyndon Lyndonville, State College, Department of Natural Sciences, and one member American Fern Society since 1932 of of the Rolla Tryon, a was born on most eminent students of pteridophytes, twentieth century's the American His professor of history father, a August 1916 Chicago, Illinois. in 26, summer Chicago, maintained a cottage in University of and education the at home Chicago. RoUa's fascination addition his in to Indiana in Chesterton, boyhood from Ches- developed during forays that and with fern allies ferns Hodge Mexico December, 1972 and was was taken by Dr. Walter H. in City in The photograph Documentation. Hunt Botanical made by the Institute for available AMERICAN FERN VOLUME NUMBER 2 JOURNAL: 92 1 (2002J terton cottage into sand dune habitats along Lake Michigan in the northwest At of Indiana. the age of he piibUshed 18, his first paper, relating his obser- Osmunda on vations Dunes plants in the Indiana complete (see bibliography As below). was a boy, Rolla greatly influenced and by, in turn influenced, Deam Charles (author of the 1940 Flora of Indiana], advising Deam about fern species he had found dunes When Deam in the area. a doubting appeared at the cottage door one day asking meet and shown Rolla be to to these ferns in he was surprised was situ, to learn that Rolla not the adult of the family but mere boy Thus a of 14. began a productive friendship documented in Rolla's Deam correspondence with from May, 1935 penny to January, 1953. All the postcards and he Deam received from have been letters maintained carefully — now one in of Rolla's files, archived at Indiana University fascinating reading. boyhood him Rolla's insatiable appetite for ferns got into a bit of trouble at home, when however, his father learned that he had charged Bower's three volumes on The Ferns account to his Brentano's bookstore at in Chicago. Rolla built a solid academic superstructure on the foundation of these boy- Among hood experiences. his academic accomplishments were an A.A. degree and 1935 in a B.S. degree in 1937, both from the University of Chicago, and Ph.M. 1938 from a in the University of Wisconsin. In 1940 he earned an M.S. and in 1941 a Ph.D., both from Harvard University. During his days as a Har- vard he student, contracted malaria South in Carolina while collecting plants M Warfare Massachusetts Ph.D. botany with in another, time this in chemistry, so that he could earn a Dartmouth Wis Minnesota While in 1945. Wis in 1945 initiated not only happy and enduring a domestic partnership but also a research synergism whose productivity has nourished pteridologists through- out the world. became In 1947 Rolla Associate Professor Botany Washington in Univer- at and sity, St. Louis Assistant Curator of the herbarium of the Missouri Botanical Garden, positions he held 1957. During appointment, he and to this Alice were the original organizers of the Missouri Botanical Garden's annual Systematics Symposium, whose 48th meeting was held 12-13 October, 2001. This highly successful annual meeting has received continuous support from the National Science Foundation from second year its (1954) to the present (with the lapse From of a single 1946 1957 year). Rolla served and to as curator librarian of American the Fern and Society's library herbarium, responding members' to requests for loans of materials. That herbarium and was library subsequently entrusted Warren H. Wagner to at the University of Michigan. Following a year Research as Associate the University at of California, Berkeley, Rolla went to Gray Herbarium the Harvard of University as Associate Curator and Curator of Ferns in 1958 and became Curator of the Gray Herbarium in 1967. Rolla ROLLA MILTON TRYON, 3 JR. and Alice traveled the world extensively, attending international meetings, conducting field work, studying specimens at major herbaria in the Americas, Europe, and Africa, and conducting field courses on the ferns. In addition to many other services to professional societies, Rolla served for years as Asso- ciate Editor of Rhodora and the American Fern Journal, as Associate Editor of and Brittonia (1961-1964), as Editor-in-Chief of fli7odora (1977-1982), as Pres- New ident of the England Botanical Club and the American Fern Society. A framed photograph of his revered mentor, Charles A. Weatherby (see American Fern Journal 40[ll for a remarkable series of papers honoring this unusually respected and beloved botanist), was always prominently displayed own on Rolla's desk Harvard, undoubtedly inspiring his welcoming, patient, at who and supportive response to all entered his office seeking counsel. In 1970 New Rolla an annual England Fern Conference Harvard Forest. For initiated at which 20 years this provided a stimulating intellectual setting in students of became fern biology discussed and developed their ideas. In 1972 he Professor and of Biology Harvard University, holding both the Curatorship Professor- at He ship until his retirement in 1987. remained at Harvard as Professor Emer- when moved itus from 1987 to 1989 he to the University of South Florida in Tampa him as Adjunct Professor, bringing with his extensive library of fern and biogeographic To mark the retirements of Alice and Rolla Tryon literature. from Harvard, of 13 papers plus introduction was published in a festschrift honor Annals Missouri Botanical Garden 225-339. their in the of the (vol. 77: 1990). At the University of South Florida, Rolla and Alice helped found the Insti- endowed Systematic Botany and the Tryon Lecture Series that brings tute for known each several internationally botanists to the university year. In their on Tampa campus, he and Alice continued research-active the their pter- office idological work, as his following bibliography indicates. exceeds 100 and includes breadth Rolla Tryon's publication titles a great list of topics. Papers ranged from articles on pteridophytes for the 1943 Encyclo- pedia Britannica to a glossary of terms relating to the fern leaf, discussions of the history of pteridology and fern classification, a remembrance of his grad- mentor and counselor Charles A. Weatherby, discussions of the formali- uate ties of fern nomenclature, and many book reviews. His monographs and re- visions focused mostly on ferns but also included angiosperms Convolvulus among were and Elymus. Signal these his revisions oi Pteridium, Doryopteris, American Notholaena, and Cyatheaceae. group, the the Selaginella rupestris on biogeography began with Doryopteris in 1944, matured in His papers fern and continued his exposition of geographic speciation in Selaginella in 1971, phytogeography North and 1999 discussion of the of eastern to his Alice's American (honoring Ching Ren-Chang). Floristic and taxonomic notes on ferns and growth forms hybrids ranged from simple observations of to eluci- ferns dations of complex taxonomic and nomenclatural issues. For his 1955 publi- Washington cation on the taxonomy of cycads (coauthored with students in his University he was awarded the 1956 Robert Montgomery award of the class) Garden distinguished achievement in the world of palms Fairchild Tropical for AMERICAN FERN JOURNAL: VOLUME NUMBER 4 92 1 (2002) and cycads. At the time of his death, he had a book review in press in Rhodora and a paper in press in Bradea (coauthored with his former student Paulo Windisch). numerous In addition to his papers on ferns and other topics, Rolla no- is Among table for his books. these are two editions of his Ferns and Fern Allies Wisconsin 1953) and Ferns and Fern Minnesota of (1940, Allies of (1954, He renowned knowledge 1980). is for his of the ferns of Peru, expressed first in his 1964 Ferns of Peru (250 pages in the Contributions from the Gray Her- barium Harvard This treatment was updated and completed of University). in six subsequent parts entitled Pteridophyta of Peru between 1989 and 1994, mostly coauthored with Robert Stolze but with several portions contributed by other pteridological His monumental 1982 book Ferns and specialists. Al- lied Plants with Special Reference to Tropical America, coauthored with Alice Tryon, an encyclopedic is treatment of this subject that continues to stimulate new research, as does his treatment of Pteridaceae, with Alice Tryon and Karl Kramer, volume and in 1 of Families Genera of Vascular Plants edited by K. R Kramer and Green, S. and Rolla's kindly perceptive mentoring and his outstanding contributions knowledge to our of ferns is signaled by having the following four fern taxa named in his honor. Asplenium 1) tryonii CorrelL In describing this species, Donovan Correll name who (1961) said '*It is a pleasure to this species for Dr. Tryon, has always been most gracious in helping his fellow-workers with their never- Known ending problems in the study of ferns/' only from Chihuahua, Mex- was ico, this species further discussed and illustrated in Ferns and Fern '/ tryonorum The eminent Mexican Riba. Journal 90:112-118, 2000) stated that "this species is M taxonomy ognizes the close professional relationship between this highly productive now known research team. This tree fern species as Trichipteris tryono- is rum Tryon by (Riba) R. following transfer Rolla in his 1970 paper on the its classification of the Cyatheaceae. am my Nephelea name tryoniana Gastony. pleased *T to this species for mentor, M. Dr. Rolla Tryon, in recognition of his outstanding contribution the understanding and to of the systematics evolution of the family Cy- atheaceae" (Gastony, 1973). Subsequent research by Conant (Conant and Cooper-Driver, 1980; Conant, 1983) revealed that this tree fern species a is now reproductively stabilized diploid hybrid species regarded that as is Alsophila tryoniana (Gas Tryonella Pichi Sermolli 4) M many author of important papers on who, supported ferns, inter alia, the distinction of the present genus from Doryopteris, though without giving it ROLLA MILTON TRYON, JR. 5 Dory opt Kramer Among the doctoral graduate students he trained, Rolla counted the follow- ing (those with asterisks received their degrees from other institutions): Alice Ramon Tryon, *Karl Kramer, * Riba, Gerald Gastony, Lawrence Palkovic, F. David Barrington, David Conant, Paulo Windisch, James Hickey, *Robbin *R. He Moran, Sonia Sultan, and Galvin Sperling. also mentored Robert Stolze in Museum. taxonomic Cnemidaria Always his revision of the Field available at to his students, he modeled his supportive and insightful mentoring on his own experiences with graduate mentor, Charles Weatherby. For he has his this earned our love as well as our respect. His impact on his students, and their and students, their students incalculable. is 1978 M. was honorary membership Amer- In Rolla Tryon, elected to in the Jr. who ican Fern Society, a special category of membership for persons have made outstanding contributions to the study of ferns. In 1984 he received a Award Merit from the Botanical Society of America recognition of distin- **In guished achievement in and contributions to the advancement of botanical science. Pre-eminently knowledgeable in matters of taxonomy and nomencla- ture, this foremost pteridologist a perceptive student of phytogeography and is of the evolutionary impact of the selective process during plant migration." was he In addition to Rolla's botanical activities also highly skilled in run- many Knox He ning a family farm in County, Indiana for years. visited the farm, overseeing management, a few times each year. The extensive records its he managing and man- kept in crops livestock illustrate his practical ability in aging business as well as scientific data. On August 20, 2001, six days before his eighty-fifth birthday, Rolla Milton work Tryon, us continue our with the pteridophytes of the world, to left Jr., We and delight in them, without him. do this fortified by his writings, the to He echoes encouraging words, and his everlasting example. was the of his beloved husband of Alice Faber Tryon, the benefactor of countless students of many who knew him never an pteridophytes, including personally, inspiration many and and counselor collaborators coauthors, the advisor of doctoral to and and students, the teacher of innimierable undergraduates, our dear friend He He mentor. be deeply missed. already will is. Literature Cited A CoNANT, D. 1983. revision of the genus Alsophila (Cyatheaceae) in the Americas. Arnold S. J. 333-382. Arbor. 64: CoNANT, and Coofer-Driver. 1980. Autogamous allohomoploidy in Alsophila and Ne- D. G. S. new homoploid homosporous pbelea (Cyatheaceae): a hypothesis for speciation in ferns. Amer. 1269-1288. Bot. 67: J. Two Texas-Chihuahuan Wrightia 200-203. CuRRELL, D. 1961. ferns. 2: S. A Gastony, G. 1973. revision of the fern genus Nephelea. Contr. Gray Herb. 203: 81-148. J. Knobloch, W. and Correll. 1962. Ferns and fern allies of Chihuahua, Mexico. Texas D. S. I. Research Foundation, Renner, Texas. Webbia Fragmenta pteridologiae— 1-16. PiCHi Sermolli, R. E. G. 1974. V. 29: AMERICAN FERN VOLUME NUMBER JOURNAL: 6 92 1 (2002) New RiBA, R. 1967. taxa in the genus Alsophila. Rhodora 69: 65-68. Bibliography of Rolla M. Tryon, (1916-2001) Jr. Some Tryon, R. M., 1934. observations on Osmundas. Amer. Fern 24: 119. Jr. J. Tryon, R. M., 1936. Ferns of the dune region of Indiana. Amer. MidL Naturalist 425-429. 17: Jr. Tryon, R. M., 1936. Botrycbium dissecturn mid form a obliquum. Amer. Fern 26: 26-30. Jr. J. Tryon, R. M., 1938. The phenomenon of forking in ferns. Amer. Fern 28: 81-90. Jr. J. Tryon, R. M., 1938. Recent additions to the flora of Indiana. Proc. Indiana Acad. Sci. 47: 76- Jr. 77. Tryon, R. M., 1939. Notes on the ferns of Wisconsin. Amer. Fern 29. 1-9. Jr. J. Tryon, R. M., Jr. 1939. The varieties of Convolvulus spitharnaeus and of C. sepium. Rhodora 41: 415-423. Tryon, R. M., 1940. Notes on some Indiana plants. Proc. Indiana Acad. Sci. 49: 89-90. Jr. Tryon, R. M., N. C. Fassett, D. W. Dunlop, and M. E. Diemer. 1940. The ferns and fern allies Jr., of Wisconsin. Botany Dept., Univ. of Wisconsin, Madison. An Osmunda Tryon, R. M., 1940. hybrid. Amer. Fern 65-66. 30: Jr. J. A Tryon, R. M.. 1941. revision of the genus Pteridlum. Rhodora 43: 1-31, 37-67. Reprinted as Jr. Contr. Gray Herb. 134: 1-70. Tryon, R. M., Jr. 1942. Review of Roland, A. E., The ferns of Nova Scotio. Amer. Fern 32: 73- J. 74. A Tryon, new R. M., 1942. Dryopteris hybrid. Amer. Fern 81-85. 32: Jr. J. A Tryon, R. M., 1942. revision of the genus Doryopteris. Contr. Gray Herb. 143: 1-80. Jr. Tryon, R. M., Jr. 1942. Review of Brown, C. A. and D. S. Correll. The ferns and fern allies of Louisiana. Rhodora 484-485. 44: Tryon, R. M., Jr. 1943. Several articles on Pteridophyta for Encyclopedia Brittanica. Tryon, R. M., Jr. 1943-1947. Abstracts of the American Fern Journal for Biological Abstracts. Tryon, Dynamic R. M., 1944. phytogeography of Doryopteris. Amer. 470-473. Jr. Dot. 31: J. Tryon, R. M., and W. Moore. 1946. Notes on aquatic and prairie vegetation in southwestern Jr. J. Minnesota. Rhodora 113-116. 48: A Tryon, and W. Moore. R. M., Jr. 1946. preliminary checklist of the flowering plants, ferns J. and fern allies of Minnesota. Univ. of Minnesota, Minneapolis. A Tryon, new R. M., 1946. Doryopteris hybrid. Amer. Fern 48-49. Jr. 36: J. A Moore, W. and M. Tryon, new R. 1946. record of Isoetes melanopoda. Amer. Fern Jr. 36: J. J. 89-91. A Butters, F. K. and R. M. Tryon, Jr. 1948. fertile mutant of a Woodsia hybrid. Amer. Fern 35: J. 132. Tryon, R. M., 1948. Some Woodsias from the north shore of Lake Superior. Amer. Fern Jr. 38: J. 159-170. A Booher, L. E. and R. M. Tryon, 1948. study of Elynms in Minnesota. Rhodora 50: 80-91. Jr. — Tryon, R. M., 1950. Charles Alfred Weatherby teacher and counselor. Amer. Fern 9-10. 40: Jr. J. A Tryon, new R. M., Jr. 1950. erect species of tlie Selaginella rupestris group. Amer. Fern 40: J. 69-74. Tryon, R. M., Jr. 1951. Ferns of the Missouri Ozark region. Missouri Bot. Card. Bull. 39: 136-138. Tryon, R. M., Jr. 1951. Review of Manton, I., Problems of cytology and evolution in the Pterido- phyta, Ecology 769-770. 32: A Tryon, R. M., Jr. 1952. sketch of the history of fern classification. Ann. Missouri. Bot. Gard. 39: 255-262. Tryon, R. M., Jr., N. C. Fassett, D. W. Dunlop and M. E. Dfemer. 1953. The ferns and fern allies Wisconsin, Ed. Univ. Wisconsin Madison. of Press, 2. Tryon, R. M., Jr. 1954. The ferns and fern allies of Minnesota. Univ. Minnesota Press, Minneap- olis. Tryon, Barbour, R. M., Jr, E. E. Davis, H. Kidd, R. Long, C. Marvin, B. Mikula, and R. Mohlen- brock. 1955. Ancient seed plants: the cycads. Missouri. Bot. Gard. Bull. 43: 65-80. ROLLA MILTON TRYON, JR. 7 Tryon, R. M., Jr. 1955. Selaginella rupestris and its allies. Ann. Missouri. Bot. Card. 42: 1-99. A Kramer, K. U. and R. M. Tryon, 1955. new species of Doryopteris from Surinam. Ann. Jr. Missouri. Bot. Card. 42: 213-214. A Tryon, R. M., Jr. 1956. revision of the American species of Notlwlaena. Contr. Gray Herb. 179. 4-106. Tryon, R. M., Jr. 1957. Adlanturn in Peru: new species and combinations. Anier. Fern 47: 139- J. 143. KoBUSKi, C. E., C. V. Morton, M. Ovvnbey, and R. M. Tryon. 1958. Report of the committee for recommendations on desirable procedures in herbarium practice and ethics. Brittonia 10: 93-95. Tryon, R. M., Jr. and A. F. Tryon. 1959. Observations on cultivated ferns: the hardy species of tree ferns (Dicksonia and Cyatheaceae). Amer. Fern. 49: 129-142. Reprinted in Lasca Leaves J. 26-33. 1960. 10: Tryon, The R. M., 1960. ecology of Peruvian Amer. Fern 46-55. ferns. 50: Jr. J. A Tryon, some R. 1960. glossary of terms relating to the fern leaf. Taxon 9: 104-109. Reprinted in Russian translation In Bot. Jour. Akad. Nauk, U.S.S.R. 46: 736-739. 1961. A Tryon, Review Munz, R. M., 1960. of Pteridophyia in A., CaUfornia Amer. Fern Jr. P. flora. J. 157-159. 50: A Tryon, R. M., 1960. review of the genus Dennstaedtia in America. Contr, Gray Herb. 187: Jr. 23-52. New Tryon, R. M., 1960. species of ferns from Central and South America. Rhodora 1-10. 62: Jr. Tryon, R. M., 1961. Taxonomic fern notes. Rhodora 63: 7-88. Jr. I. Tryon, R. M., 1962. The fern genus Doryopteris in Santa Catarina and Rio Grande do Sul, Jr. Sellowia 51-59. Brazil. 14: Tryon, R. M., 1962, Review of Wherry, E. T., The Fern Guide. Amer. Fern 52: 89-91. Jr. J. A Tryon, R. M., 1962. note on Nepbrolepis cordifolia cv. Daffii. Amer. Fern 52: 153-155. Jr. J. Tryon, R. M., 1962. Taxonomic Fern Notes. PityTogromma (including Trismerio) and Ana- Jr. II. gramma. Contr. Gray Herb. 189. 52—76. Tryon, R. 1962. Taxonomic fern notes. Contr. Gray Herb. 191: 91-107. III. Tryon, R. M., 1962. Review of Knobloch, W. and D. S. Correll, Ferns and Fern Allies of Jr. I. Chihuahua. Rhodora 347-348. 64: A Tryon, R. 1962. commentary on superfluous names. Taxon 11: 116-120. Tryon, Nomenclatural Taxon 28-285. R. 1963. proposals. 12: Mem. Tryon, R. M. 1964. Evolution in the leaf of living ferns. Torrey Bot. Club 21: 73-85. Tryon, R. 1964. The ferns of Peru. Polypodiaceae (Dennstaedtieae to Oleandreae]. Contr. Gray Herb. 194: 1-253. Tryon, R. 1964. Taxonomic Fern Notes IV. Rhodora 66: 110-117. Pichi-Sermolli, R. E. G., F. Ballard, R. E. Holttum, H. Ito, F. M. Jarrett, A, C. Jermy, E. A. C. L. E. Schelpe, M.-L. Tardieu-Blot, and R. M. Tryon. 1965. Index Filicum Supplementum Quartum pro Annis 1934-1960. Internat. Bureau for Plant Taxonomy and Nomenclature, Utrecht, Netherlands. Tryon, R. 1965. Paraphyses in the ferns. Taxon 14: 213-218. A Tryon, R. and D. M. Britton. 1966. study of variation in the cytotypes of Dryopteris spinulosa. Rhodora 59-92. 68; New Tryon, R. 1967. Taxonomic fern notes V. combinations in Peruvian species of Thelypteris. Rhodora 5-8. 69: Tryon, R. 1967. Review of Hara. H., The flora of eastern Himalaya, Rhodora 69; 456-457. A compounds some BojiM, B. A. and R. M. Tryon. 1967. Phenolic in ferns. survey of ferns for I. cinnamic acid and benzoic acid derivatives. Canad. Bot. 45: 585-593. J. Tryon, R. M. and A. F Tryon. 1968. Edith Scamman (1882-1967J. Amer. Fern 58: 1-4. J. Tryon, R. 1968. Nomenclatural proposals. Taxon 17: 588-590. Ewan A Tryon, R. 1969. Pteridology, Pp. 97-102 in (ed.) short histor}' of botany in the United J. New Hafner Publishing Co., York. States. Tryon, R. 1969. Taxonomic problems in the geography of North American ferns. BioScience 19: 790-795. VOLUME NUMBER AMERICAN FERN JOURNAL: 8 92 (2002) 1 Manual Tryon, R. 1970. Review of Correll, D. S. and M. C. Johnston., of the vascular plants of Rhodora 533-535. Texas. 72: Tryon, R. 1970. The classification of the Cyatheaceae. Contr. Gray Herb. 200: 3-53. Tryon, R. 1970. Development and evolution of fern floras of oceanic islands. Biotropica 2: 76-84. Reprinted 1971 as Pp. 54-62 in W. L. Stern (ed.) Adaptive aspects of insular evolution. Washington State Univ. Press. Tryon, R. 1971. The American tree ferns allied to Spbaeropteris horrida. Rliodora 73; 1-19. Tryon. R. 1971. Ferns of the Andes and Amazon. Morris Arbor. Bull. 22: 7-13. The 89- Tryon, R. 1971. process of evolutionary migration in species of Selaginella. Brittonia 23: 100. Tryon, R. 1972. Endemic areas and geographic speciation in tropical American ferns. Biotropica 121-131. 4: — New Tryon, R. 1972. Taxonomic fern notes, VI species of American Cyatheaceae. Rhodora 74: 441-450. Tryon, R. M., and A. Tryon. 1973. Geography, spores and evolutionary relations in the F. Jr. cheilanthoid ferns. Bot. Linn. Soc. 67: Suppl. 1; 145-153. J. 65- Tryon, A. and R. Tryon. 1973. Thelypteris in northeastern North America. Amer. Fern 63: J. 76. Tryon, R., B. Voeller, A. Tryon, and R. Riba. 1973. Fern biology in Mexico (a class field program). BioScience 28-33. 23: Tryon, Widen, A. Huhtikangas, and M. Lounasmaa. 1973. Phloroglucinol derivatives in R., C.-J. Dryopteris parallelogramma and D. patulo. Phytochcmistry 12: 683-687. Tryon, A. and R. Tryon. 1974. Geographic patterns in temperate American ferns and some rela- tionships in Thelypteris. Amer. Fern. 64: 99-104. J. Tryon, R. 1975. The Benjamin D. Gilbert fern herbarium. Amer. Fern 65: 60. J. Tryon, R. M. and G. Gastony. 1975. The biogeography of endemism in the Cyatheaceae. Fern }. Gaz. 11: 73-79. Tryon, R. M. and D. S. Conant. 1975. The ferns of Brazilian Amazonia. Acta Amazon. 23-34. 5: A Tryon, R. 1976. revision of the genus Cyathea. Contr. Gray Herb. 206: 19-98. Gastony, G. and R. M. Tryon. 1976. Spore morphology in the Cyatheaceae. The genera II. J. Lophosoria, Metaxya, Spbaeropteris, Alsophila, and Nephelea. Amer. 738-758. Bot. 63: J. Tryon, R. M. 1977, Studies on the American Cyatheaceae and on the classification of the family. 2839-2842. Males. Bull. 30: Fl. Tkyon, M. and Vitale. 1977. Evidence antheridogen production and mediation R. G. for of a its mating system in natural populations of fern gametophytes. Bot. Linn. Soc. 74: 243-249. J. Tryon, R. 1979. Biogeography of the Antillean fern flora. Pp. 55-68 in D. Bramwell (ed.) Plants New and Academic islands. Press, York. Tryon, A. R. Tryon, and Badre. 1980. Classification, spores, and nomenclature of the marsh F., F. Rhodora 461-474. fern. 82: Tryon, R. 1980. Review of Steyermark, A. and O. Huber, Flora of Avilo. Amer. Fern 70: 79. J. J. Tryon, R. M,, 1980. The ferns and fern allies of Minnesota, Ed. 2. Univ. Minnesota Press, Jr. Minneapolis. Tryon, R. and A. Tkyon. 1981. Taxonomic and nomenclatural notes on ferns. Rhodora 83: 133- 137. Tryon, R. 1982. Review of Smith, A. R., Pteridopbytes in D, E. Breedlove (ed.) Flora of Chiapas Madrono Port 2. 29: 216. Tryon, R. and A. Tryon. 1982. Additional taxonomic and nomenclatural notes on ferns. Rho- F. dora 125-130. 84: Tryon, R. M. and A. Tryon. 1982. Ferns and allied plants with special reference to tropical F. New America. Springer-Verlag, York. An new Tryon, R. 1984. unusual Elaphoglossum from Peru. Amer. Fern 74: 108-110. J. Tryon, R. 1985. Fern speciation and biogeography. Proc. Roy. Soc. Edinburgh, B. 86: 353-360. Tryon, R. 1986. Dicksoniaceae. Lophosoriaceae. Metaxyaceae. Cyatheaceae. Pp. 1-59 in G. Harling and L. Andersson (eds.) Flora of Ecuador, No. 27. Swedish Research Councils, Stockholm. Tryon, R. 1986. Some new names and combinations in Pteridaceae. Amer. Fern 76: 184-186. J. ROLLA MILTON TRYON, 9 JR. Tryon, R. 1986. The biogeography of species, with special reference to ferns. But. Rev. [Lancaster] 117-156. 52: Tryon, R. 1989. Ptcridophytes. Pp. 327-338 in H. Lieth and M. A. Werger (eds.) Tropical rain J. Amsterdam. ecosystems. forest Elsevier, Ophioglossaceae-12. Cy- Tryon, R. M. and R. G. Stolze. 1989. Pteridophyta of Peru. Part 1. I. New 1-145. atheaceae. Fieldiana, Bot. Series 20: Dennstaed- Tryon, R. M. and R. G. Stolze. 1989. Pteridophyta of Peru. Part IT. 13. Pteridaceae-15. New 1-128. tiaceae. Fieldiana, Bot. Series 22: Tryon, M., A. Tryon, and K. U. Kramer. 1990. Pteridaceae. Pp: 230-256, in K. Kubitzki (ed.) R. F. and gymnosperms. The families and genera of vascular plants. Vol. 1. Pteridopbyics Vol. New Kramer and Green. Springer-Verlag, York. eds. K. U. S. P. Dryopteridaceae [with Tryon, M. and Stolze. 1991. Pteridophyta of Peru. Part IV. 17. R. R. G. New 1-176. collaboration of Mickel and R. C. Moran). Fieldiana, Bot. Series 27: T. J. Thelypteridaceae (con- Tryon, M. and Stolze. 1992. Pteridophyta of Peru. Part 16. R. R. G. III. New 1-80. tributed by A, R. Smith]. Fieldiana, Bot. Series 29: Aspleniaceae-21. Poly- Tryon, M. and Stolze. 1993. Pteridophyta of Peru. Part V. 18. R. R. G. New 1-190. podiaceae (with collaboration of B. Leon). Fieldiana, Bot. Series 32: Tryon, M. and Stolze. 1994. Pteridophyta of Peru. Part VI. 22. Marsiliaceae-28. Isoctaccae R. R. G. New 1-123. (with collaboration of R. Hickey and B. 01Igaard). Fieldiana, Bot. Series 34: J. Rhodora 335-343. Tryon, 1997. Systematic notes on Oleandra. 99: R. name ebeneum Taxon 339- Acrostichurn (Pteridaceae). 46: Tryon, R. 1997. Proposal to reject the 340. Churchill, H., R. Tryon, and D. S. Barrington. 1998. Development of the sorus in tree ferns: 1245-1252. Dicksoniaceae. Canad. Bot. 76: J. North Amer- Tryon, M. and A. Tryon. 1999. Observations on the phytogeography of eastern R. F. Pp 250-273 X-G Zang and K-H Shing Clung Memorial Volume. Institute (eds.). ican ferns. in Chinese Acad. Beijing. of Botany, Sci., 428-438. Tryon, 2000. Systematic notes on the Old World fern genus Oleandra. Rliodora 102: R. manual and Moran, Fern grower's revised Tryon, M. 2001. Review of Hoshizaki, B. R. C. R. J. Rhodora WiNDISCH Bradea 267-276. probable migration route and its present fern flora. 8: