Russian Entomol. J. 14(4):251–262 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2005 Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus Clairville, 1806 (Coleoptera: Dytiscidae) Çàìåòêè î íîìåíêëàòóðå, òàêñîíîìèè è ôàóíèñòèêå ðÿäà âèäîâ èç ðîäà Hydroporus Clairville, 1806 (Coleoptera: Dytiscidae) H. Fery1, P.N. Petrov2 Õ. Ôåðè1, Ï.Í. Ïåòðîâ 2 1 Räuschstr. 73, D-13509 Berlin, Germany. e-mail: [email protected] 1 Ройшштрассе 73, D-13509 Берлин, Германия. 2 Moscow South-West Gymnasium 1543, ul. 26 Bakinskikh Komissarov 3-5, 119571 Moscow, Russia. e-mail: [email protected] 2 Московская гимназия на Юго-Западе №1543, ул. 26 Бакинских комиссаров 3-5, 119571 Москва, Россия. KEY WORDS: Coleoptera, Dytiscidae, Hydroporus, lectotypes, new synonyms, first records. КЛЮЧЕВЫЕ СЛОВА: Coleoptera, Dytiscidae,H ydroporus, лектотипы, синонимы, первые указания. ABSTRACT. Several species of Hydroporus Clair- Wehncke, 1876 syn.n. Обозначены лектотипы следу- ville, 1806, chiefly members of the Hydroporus pla- ющих таксонов: Hydroporus corsicus Wehncke, 1872, nus-group, are revised at least in part, and information Hydroporus cyprius Régimbart, 1878, Hydroporus dis- on the taxonomy and the distribution of these and cretus ponticus Zaitzev, 1927, Hydroporus distinguen- some other species of the genus is given. The follow- dus var. estrellensis Schaufuss, 1882, Hydroporus lim- ing new synonymies are proposed: Hydroporus trans- batus Aubé, 1838, Hydroporus brucki Wehncke, 1875, grediens Gschwendtner, 1923 = Hydroporus discretus Hydroporus analis Aubé, 1838, Hydroporus decipiens ponticus Zaitzev, 1927 syn.n.; Hydroporus goldschmidti Sharp, 1878, Hydroporus habelmanni Wehncke, 1876и Gschwendtner, 1923 = Hydroporus goldschmidti var. Hydroporus bicolor G. Müller, 1933. Следующие виды recidivus Gschwendtner, 1923 syn.n., and Hydroporus впервые указаны для ряда территорий : H. tessellatus (Drapiez, 1819) = Hydroporus habelmanni transgrediens — Азербайджан, Армения, Иран, Турк- Wehncke, 1876 syn.n. Lectotypes of the following taxa менистан и Турция, H. pubescens (Gyllenhal, 1808) — are designated: Hydroporus corsicus Wehncke, 1872, Грузия и азиатская часть России (Урал) и Hydro- Hydroporus cyprius Régimbart, 1878, Hydroporus dis- porus marginatus (Duftschmid, 1805) —И ран и Казах- cretus ponticus Zaitzev, 1927, Hydroporus distinguen- стан. dus var. estrellensis Schaufuss, 1882, Hydroporus lim- batus Aubé, 1838, Hydroporus brucki Wehncke, 1875, Introduction Hydroporus analis Aubé, 1838, Hydroporus decipiens Sharp, 1878, Hydroporus habelmanni Wehncke, 1876, The present authors have been dealing with mem- and Hydroporus bicolor G. Müller, 1933. Several first bers of the genus Hydroporus for several years and records are given:H . transgrediens from Armenia, Azer- have studied a lot of material from a number of muse- baijan, Iran, Turkey, and Turkmenistan, H. pubescens ums and private collections. These studies allow a new (Gyllenhal, 1808) from Georgia and Asian Russia (the valuation of some taxa, and we take this opportunity to Urals), and Hydroporus marginatus (Duftschmid, 1805) propose several new synonymies and designate some from Iran and Kazakhstan. lectotypes. Further data of interest, such as on the distribution of these and some other taxa, have also РЕЗЮМЕ. Ревизованы (по крайней мере, частич- been included, as well as several first records. но) некоторые виды из рода Hydroporus Clairville, 1806, большинство из которых относятся к группе Hydroporus planus. Приведены сведения по таксо- Material and methods номии и распространению этих, а также некоторых других видов данного рода. Установлена синонимия The following acronyms are used for the collections следующих названий: Hydroporus transgrediens where material is located: Gschwendtner, 1923 = Hydroporus discretus ponticus BMNHBritish Museum of Natural History, London, Zaitzev, 1927 syn.n.; Hydroporus goldschmidti Great Britain (R. Booth) Gschwendtner, 1923 = Hydroporus goldschmidti var. CHF coll. H. Fery, Berlin, Germany; property of the recidivus Gschwendtner, 1923 syn.n. и Hydroporus NMW tessellatus (Drapiez, 1819) = Hydroporus habelmanni CJH coll. J. Hájek, Praha, Czech Republic 252 H. Fery, P.N. Petrov CJS coll. J. Št’astný, Liberec, Czech Republic (lectotype designation of H. discretus ); Nilsson, 2001: 156, CGW coll. G. Wewalka, Vienna, Austria 2003: 60. CKE coll. Ö.K. Erman, Erzurum, Turkey MATERIAL. Type material : Lectotype of Hydroporus CLH coll. L. Hendrich, Berlin, Germany discretus designated by Balke & Fery [1993: 95], stored in the MNHN. DEUMDepartment of Entomology, Moscow Lomono- Lectotype of Hydroporus corsicus (by present designa- sov State University, Russia (V.Y. Savitsky) tion): #, a very small square label without any text, “Haag” [= IRSN Institut Royal des Sciences Naturelles de Bel- collector?, hw Wehncke], “Corsica” [hw Wehncke], “Muséum gique, Brussels, Belgium (M. Cludts, K. De- Paris, ex coll. R. Oberthur, ex Wehncke”, “Lectotype, Hydro- porus corsicus Wehncke, 1872, des. H. Fery 2005” [red, printed]; sender) placed in the drawer below a label “ corsicus mihi” [hw MNB Museum für Naturkunde, Humboldt-Universäitt, Wehncke] (MNHN). Paralectotype: 1 $, “Habelm” [= collector Berlin, Germany (F. Hieke, M. Uhlig, B. Jaeger) Habelmann, hw Wehncke], “Muséum Paris, ex coll. R. Oberthur, MCTRMuseo Civico di Storia Naturale, Trieste, Italy ex Wehncke”, and the respective red paralectotype label (G. Alberti, A. Colla) (MNHN). Notes: The paralectotype is assumed to belong to the syntype series, since it is prepared in the same manner as MNHNMus éum National d’Histoire Naturelle, Paris, the lectotype. Wehncke [1872: 163] compared his new taxon France (H. Perrin) with Hydroporus longulus Mulsant & Rey, 1861, a species MRTOMuseo Regionale di Scienze Naturali, Torino, which does not belong to the planus-group, but to the Italy (M. Daccordi) Hydroporus longulus-group. And, indeed, the two types and some other specimens studied from Corsica are somewhat NMB Naturhistorisches Museum Basel, Switzerland elongate and at first glance resemble H. longulus. (M. Brancucci) Lectotype of Hydroporus cyprius (by present designation): NMW Naturhistorisches Museum Wien, Austria, Vi- #, “Chypre” [green, hw?], “ cyprius Régimb.” [hw Régimbart], enna (M. Jäch) “Muséum Paris, Coll Maurice Régimbart, 1908”, “Lectotype, OLML Oberösterreichisches Landesmuseum, Linz, Hydroporus cyprius Régimbart, 1878, des. H. Fery 2005” [red, printed] (MNHN). Paralectotypes: 7 exs., one specimen with Austria (F. Gusenleitner, C. Reitstätter) first and second labels as in the lectotype, the rest only with SMNS Staatliches Museum für Naturkunde Stuttgart, the first label; coll. Sedillot (MNHN). Each paralectotype is Germany (W. Schawaller) provided with its respective red label. ZISP Zoological Institute of the Russian Academy of Additional material examined : France, Corsica: 2 exs., “Corse”, “Muséum Paris, ex coll. R. Oberthur”; 4 exs., “Vizzavo- Sciences, St. Petersburg, Russia (A.G. Kirejtshuk) na, Corse”, “ Hydroporus Corsicus ([1]895)”, “Corse, Damry”, ZSM Zoologische Staatssammlung, München, Ger- “Muséum Paris, ex coll. R. Oberthur”; 2 exs., “Rouisseau, Zonza”, many (M. Baehr, M. Kühbandner) “Hydroporus Corsicus”, “Corse, Damry”, “Muséum Paris, ex coll. The following abbreviations are used in the text: TL R. Oberthur”; 1 ex., “Corsicus”, “Corse, Damry”, “Muséum Paris, (total length), MW (maximum width), hw (handwriting), ex coll. R. Oberthur” (all in MNHN). 1 #, “F 37 (2B) Morosaglia, (Corte), Bocca di Riscamone, 570 m. 09°15E and hw? (handwriting of an unknown person). 42°26N. 01.10.[19]91. leg. GER [= Gerecke]”; 1 #, “Bastia, For the ink drawings, male genitalia were studied Corsika, V.1907, F. Hopp”; 1 $, “10.7.[19]86 Corse, Pinito, Porto, wet. The terminology denoting genitalia orientation fol- Quellmoos, Fery leg.”; Cyprus: 7 exs., “31.3.[19]96 Zypern, nr lows Miller & Nilsson [2003]. All lectotype designa- Kannaviou, N Pafos, Bach, Fery leg.”; 3 exs., “10.4.[19]96 Zypern, nr Kannaviou, N Pafos, Ezousa river, Fery leg.”; 1 $, “4.4.[19]96 tions are made in order to support the stability of the Zypern (Lim.), Mandria, SW Platres, Bach, Fery leg.”; 5 exs., nomenclature (cf. Article 74.7.3 of the ICZN [1999], and “8.4.[19]96 Zypern (La.), Vavatsinia, W Lefkara, Bach, Fery leg.” Anonymous [2003]). (all in CHF). Specifications of localities are taken from standard DESCRIPTIVE NOTES. Specimens from Corsica coincide sources, such as Stiehlers Handatlas (Gotha, 1928/30), more or less with central European ones. Those from Cyprus often have the elytra somewhat more brownish; in addition, we Microsoft Encarta World Atlas 2000, and The Times have observed that the gonocoxosterna are slightly longer and Atlas of the World, Comprehensive Edition (London, the gonocoxae more rounded apically. With respect to the high 1997). We have adopted the style of the latter source variability of H. discretus, however, these observations seem for the presentation of the names and co-ordinates of not to justify a treatment as proper species or subspecies. See most localities. The authors’ remarks are given in square also the ‘Discussion’ at the end of the section about Hydro- brackets. porus transgrediens Gschwendtner, 1923. The male genitalia of a specimen from Bavaria, Germany, are given in Figs 1–3. Results Hydroporus transgrediens Gschwendtner, 1923 Gschwendtner, 1923: 109 ( H. pubescens transgrediens ) Hydroporus planus-group Type locality — Transcaspien, Neu-Saratow Hydroporus discretus = Hydroporus discretus ponticus Zaitzev, 1927: syn.n. (Type Fairmaire & Brisout de Barneville, 1859 locality — Armenia, Yerevan distr., Dara i ag [=Tsaghkad- zor], ca. 40 km NNE Yerevan) Fairmaire & Brisout de Barneville in Fairmaire, 1859: 28 Zimmermann, 1931: 137 ( pubescens transgrediens ); Guignot, Type locality — France, Marly 1932: 372 ( pubescens var. transgrediens); Zaitzev, 1951: 56 = Hydroporus corsicus Wehncke, 1872: 163 (Type locality — (pubescens transgrediens ), 1953: 165 ( pubescens transgredi- France, Corsica) ens), 169 (discretus ponticus ), 1972: 174 ( pubescens transgre- = Hydroporus cyprius Régimbart, 1878: 352 (Type locality — diens), 179 (discretus ponticus ); Wewalka, 1992: 55 (lectotype Cyprus) designation of H. pubescens transgrediens ); Ádám, 1996: 59 Guignot, 1959: 393 (synonymy of H. corsicus ); Miller et al., (ponticus); Nilsson, 2001: 156 ( discretus ponticus ), 158; Nils- 1997: 27 (synonymy of H. cyprius); Balke & Fery, 1993: 95 son, 2003: 61 ( discretus [= ponticus]), 65. Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus (Dytiscidae) 253 1 2 3 4 5 6 7 8 9 10 11 12 Figs 1–12. Male genitalia of Hydroporus spp.: 1–3 — H. discretus; 4–6 — H. transgrediens; 7–9 — H. goldschmidti; 10–12 — H. planus; 1–2, 4–5, 7–8, 10–11 — penis; 3, 6, 9, 12 — paramere; 1, 4, 7, 10 — ventral view; 2–3, 5–6, 8–9, 11–12 — lateral view. Ðèñ. 1–12. Ãåíèòàëèè ñàìöîâ Hydroporus spp.: 1–3 — H. discretus; 4–6 — H. transgrediens; 7–9 — H. goldschmidti; 10– 12 — H. planus; 1–2, 4–5, 7–8, 10–11 — ïåíèñ; 3, 6, 9, 12 — ïàðàìåðà; 1, 4, 7, 10 — ñíèçó; 2–3, 5–6, 8–9, 11–12 — ñáîêó. MATERIAL. Type material : Lectotype of Hydroporus gical Institute, Russian Academy of Sciences, St. Petersburg” pubescens transgrediens: #, “Transcaspia., Neu-Saratow”* [prin- [yellow], “Lectotype, Hydroporus discretus ponticus Zaitzev, ted], “Type, Gschw” [greyish brown label, hw Gschwendtner], 1927, des. H. Fery 2005” [red, printed] (ZISP). Paralectotypes: “Coll. Gschwendtner” [printed], “ Hydroporus pubescens trans- Armenia: 14 exs., the same label data as the lectotype; two grediens Gschw” [hw Gschwendtner?], “Type” [red, hw?], “Lec- specimens considerably damaged by dermestids; Georgia: 3 totypus, Hydroporus transgrediens Gschw, det. G. Wewalka ##, 1 $, “Baraleti [ca. 41.32N 43.29E], prov. Achalk., VI–VII. [19]91” [red, hw Wewalka] (OLML). Paralectotype: 1 #, first [19]16.”; 1 ex., “Ordzhoch [not found on any map], doroga [= three labels as in the lectotype, “ pubescens v. transgrediens road], 28.6.[19]09, Bat. ob. [= Batumi oblast]” [Cyrillic], consid- Gsch., det. Gschwendtner” [hw Gschwendtner], “Type” [red, erably damaged by dermestids; Turkey: 1 #, “Bajburt vic., distr. hw?], “Paralectotypus, Hydroporus transgrediens , det. G. We- Erzerum, 18.VII.[19]17”; 1 $, “Kesalar, distr. Erzerum, walka [19]91” [red, hw Wewalka] (OLML). 29.VII.[19]17”, somewhat damaged; 1 ex., “Sadach, 7.VII.[19]17, Lectotype of Hydroporus discretus ponticus (by present E...itter [? illegible]” [hw?], considerably damaged by dermestids; designation): #, “DaraÈiÈag [= Darachichag = Tsaghkadzor, ca. according to Zaitzev [(1927: 17)] this locality is in “Trapezunt 40.32N 44.45E] & vic, distr. Erivan, 24.VIII.[19]12.”, “Zoolo- [= Trabzon] Province”; Ukraine: 3 exs., “Krim. Jalta district, Tauschan-Basar, 15.6.[19]07. W. Pliginskiy” [Cyrillic], one spec- * We have not found a locality named “Neu-Saratow” [= New imen considerably damaged by dermestids; 1 $, “Krim. Jalta Saratov = Novyy Saratov] on any map. There exists the district, Tauschan-Basar, 23.6.[19]07. W. Pliginskiy” [Cyrillic]; 1 village Novosaratovka [which can be roughly translated #, “Krim. Angarsk-pass, puddle near road, 27.6.[19]07. W. from Russian as “New Saratov Village”] (40.36N 45.35E) in Pliginskiy” [Cyrillic]; 1 ex. “Krim”, considerably damaged by Azerbaijan, a country that, however, does not belong to dermestids; 1 #, “Sebastopol, Krim 22.3.[19]06, W. Pliginskiy”; “Transcaspia”. Possibly, “Transkaspien [= Transcapia]” is a 1 $, “Sebastopol, Krim 8.4.[19]06, W. Pliginskiy”; 1 $, “Sebasto- misspelling of “Transcaucasia”, and Novosaratovka might pol, Krim 26.V.[19]06, W. Pliginskiy”; 1 $, “Tauria 89, Aluschta be the correct type locality, a solution which we assume to be [ca. 44.41N 34.31E]” (ZISP). 1 #, “Sebastopol, Krim, Inkerman the most probable. Nilsson [2003: 65] provides as distribu- [suburb of Sevastopol], 6.3.[19]10 W. Pliginskiy.” [Cyrillic in tion area “TM” [= Turkmenistan], which is certainly due to part], “334”, “Hydroporus sp. n.” [hw Reitter], “Reitter det.” the statement in Wewalka [1992: 54] who gave “Turk- [printed], “H. planus var. aut n. sp., V. Pliginski. coll.” [hw Reitter meniya” as distribution area. in part], “Reitter det.” [printed], “ Hydroporus discretus ponticus 254 H. Fery, P.N. Petrov Zaitz., Shaverdo H. det. 2000”; 1 $, “Krim, Jalta Kreis, village emerak, 23.09.2001, Erman, Ö.K. leg.” (CKE); 7 exs., “18.9.2000 [?] Saki., 8.VIII.[19]13[?] W. Pliginskiy” [Cyrillic in part], “ Hydro- (TR) prov. Erzurum, Yedigöller, nr Uzunkavak, ca. 3000 m, ca. porus discretus ponticus Zaitz., Shaverdo H. det. 2000”; 1 #, 20 km E spir, Ö.K. Erman leg.” (CHF); 9 exs., “18.4.2000 (TR) “Tauria, village Saki, Yepatoria distr., 7.8.[19]12 W. Pliginski” Erzurum, ca. 28 km SW Erzurum, N pass, ca. 1800 m, ponds [Cyrillic in part], “H. planus var. aut n. sp., V. Pliginski. coll.” [hw on meadow, Fery leg.” (CHF); 3 exs., “18.4.2000 (TR) Erzurum, Reitter in part], “Reitter det.” [printed], “ Hydroporus discretus ca. 43 km SW Erzurum, S pass, ca. 2000 m, ponds on meadow, ponticus Zaitz., Shaverdo H. det. 2000”; 1 $, “Tauria, village Fery leg.” (CHF); 1 ex., “Turkey 20.VI.2003, Erzurum vil. [= Saki, Yepatoria distr., 26.8.[19]12 W. Pliginski” [Cyrillic in part], province] (ca 15 km W), Ilica env. (pools; sweeping), (39°55’N, “H. planus var. aut n. sp.?” [hw Reitter], “Reitter det.” [printed], 41°06’E; 1770 m), Jiøí Hájek & Josef Hotový leg.” (CJH); 1 ex., “Hydroporus discretus ponticus Zaitz., Shaverdo H. det. 2000”; “TR 5.6. 1987, Van – Baskale 2600 m, Güzeldere P. [= pass], leg. 1 #, “Saki, Krim 18.8.[19]10, W. Pliginski” [Cyrillic in part], Jäch (62)” (NMW); 2 exs., “Turkey 28.–29.VI.2003, Van vil. [= “Hydroporus nigrita” [hw Reitter], “Reitter det.” [printed], “T.k. province] (ca 40 km SW), Geva º env. (stream), (38°16’N, nadkrylya bez shagrenirovannogo fona, to eto discretus, v 43°03’E; 1880 m), Jiøí Hájek & Josef Hotový leg.” (CJH); 2 exs., spisok voshyol kak H. nigrita, N 388 yeshcho raz poslat na “Turkey 27.–28.VI.2003, Van vil. [= province] (ca 45 km SE), proverku, W. Pliguinskii [= Since the elytra are without Guselsu – Ho ºap (stream), (38°18’N, 43°47’E; 2070 m), Ji øí reticulate (shagreen) surface, this is discretus, was included in Hájek & Josef Hotový leg.” (CJH); 1 $, “Umg. [= environs] the checklist as H. nigrita, send No. 388 to be checked again]” Kayabo az, se Geva º, 2100–2200 m, 31.VII.1985” *, “Südost- [“W. Pliguinskii” printed in Latin; rest in Cyrillic, hw Pliginski?]. Anatolien, Heinz leg.” (NMB); 5 exs., “Turkey 17.–18.VI.2003, Unknown localities: 1 ex., “st. Passanaur, dist. Dušet, VII.[19]13”, K. [= Karaman] Mara º (ca 25 km SW), Fatih env. (stream), specimen almost totally destroyed by dermestids; 2 exs., (37°26’N, 36°41’E; 1115 m), Ji øí Hájek & Josef Hotový leg.” “Suram, 6.VI.[19]17, Ahnger”, one specimen considerably da- (CJH); 2 exs., “TR – Isparta, Gelendost, 9–VI–[19]81 G. Sama” maged by dermestids (all in ZISP). Each paralectotype is (CHF); 1 #, “TR – Antalya 24.5.[19]91, n Altinyaka 1100 m, leg. provided with its respective red label and the yellow label of Schödl (32)” (NMW); 1 ex., “Turkei 21.6.[19]97, Göreme, lgt Jiøí the Zoological Institute of the Russian Academy of Sciences. Hájek”, “Hydroporus discretus ponticus Zaitzev, Jiøí Hájek det. Additional material examined: Ukraine: 2 ##, “Tauria 2001” (CJH). 5 exs., “Simav Dag [ca. 39.07N 28.30E], 1978, [= Crimea]” [blue label]; 1 ex., “Tauria 89, Mshatka [= Yuzhnoye, 100/1500 m 16.VIII.”, “W Anatolien, leg. Casale” (NMB); 3 ca. 44.24N 33.48E]” [hw?], considerably damaged by dermestids exs., “TR 15.6. 1987, 80 km w. Istanbul, leg. M. Jäch (79)” (all in ZISP); 10 exs., “Crimea (Ukraine), env. Morskoje [ca. (NMW). Iran (first record): 1 #, “Danavar-dagh, s.e. [= south- 44.50N 34.48E], 24.04.1994” (NMB, CHF). Russia: 10 exs., east of] Ushnuiyeh, 1600 m, 10.VIII.1969”, “Azerbaijan (Iran), “Kaukasus, Dagestan, Kurush, 2200–2400 m, 3.–6.7.1991, Heinz leg.” (NMB). 21 exs., “15.8.1998 Iran, Kohkiluyeh & Martens, Schawaller et al.” (SMNS, CHF). 1 #, “USSR, Boyer Ahmad, 7 km SW Yasuj, Elmi & Fery leg.”, “Cheshmeh Sar- Caucasus, Krasnaja Poljana, 4.–8.7.1988, Dr. L. Danék lgt.” e-Ab Taveh, brook and pools (with stagnant water) (#2106)”; (CJS). Georgia: 2 exs., “Kaukasus, Tiflis”, one specimen consid- 4 exs., “15.8.1998 Iran, Fars, 17 km SE Sepidan (= Ardakan), erably damaged by dermestids (ZISP). Armenia (formally a Shesh Pir”, “ditch with running water, Elmi & Fery leg. first record under the name H. transgrediens ): We have (#2108)”; 10 exs., “18.4.2001 Iran, Fars, 17 km SE Sepidan (= studied a large amount of specimens collected by Shaverdo and Ardakan), Shesh Pir”, “ditch with running water, Elmi & Fery Schillhammer in May 2001. The respective records are listed in leg.”; 168 exs., “16.8.1998 Iran, Fars prov., Bamoo-Nat. Park, Shaverdo [2003] under the name H. discretus ponticus and shall spring Cheshmeh-ye-Ghanbari, Elmi & Fery leg. (#2109)”; 9 not be repeated here. Azerbaijan (first record): 6 exs., “Disar exs., “16.8.1998 Iran, Fars, Nat. P. Bamoo, Darreh-Bishen nr Ordubad, Nachitshewanski region, Znojko [collector] spring-stream system, Elmi & Fery leg. (#2113)” (all in CHF). 22.VII.[1]933”, “Hydroporus discretus ponticus Zaitz., Shaverdo Turkmenistan (first record, except that from the doubtful H. det. 2000” (ZISP); Turkey (formally a first record from locus typicus — see above): 1 $, “Turkmenia, Bolshoi [= big] Turkey under the name H. transgrediens, however known from Balkhan [mountain range, ca. 200 km E Krasnovodsk, ca. 40N this country under the name H. discretus ponticus since Zaitzev 55E], 10.VII.[19]40 leg. Starostin”; 1 #, “Turkmenia, Mary [1927]): 4 exs., “Turquie: Artvin, SW Artvin, 1900 m, 9.VI.1986”, station [ca. 37.34N 61.48E; Mary = ancient Merw], leg “Besuchet – Löbl, Burckhardt” (CHF); 3 exs., “TR – Artvin (47), Starostin” [Cyrillic]; 1 #, “Aj-Dere, W Kopet-dag, Iljitshev Artvin – Savsac, Schödl 4.6.[19]89” (CHF, further specimens in [collector], 30.IV.[1]952”; 1 #, “Kara-kala env., Turkmen., NMW); 1 #, “Turkey, Artvin prov., 8 km E avºat, Karagöl Steinberg [collector] 26.IV.[1]953” [Cyrillic] (all in ZISP). 1 #, Sahara Nat. Park, 1930–2170 m, 4.–5.VII.2004, Ji øí Hájek & 3 $$, “Turkmenia, 20.05.[19]87., mountain range Kopet Dag, Jan Rù iÈka leg.”, “41°13.5’N 042°27.1’E – 41°13.5’N 042°28.0’E, Schlucht [= gorge] Kuru Chudan, Wolkow D.A. leg.” (DEUM). pasture / wet coniferous forest, (dominant Picea, Abies)” (CJH). Three of the specimens from Turkmenistan are with rather 1 $, “ca. 13 km S Posof [ca. 41.30N 42.41E, in Ardahan prov.], yellowish spots near the base of elytra. 2000 m 18.VII.1983”, “Anatolien, W. Heinz” (NMB). 1 #, “TR DESCRIPTIVE NOTES. For detailed descriptions of – Rize, kisdere, Anzer Köyü Yaylasi, Göllerin Alt ndaki Dere, H.transgrediens see Gschwendtner [1923: 109] and Wewalka Brikinti, 29.6.[20]01, Erman, Ö.K. leg.” (CKE). 1 #, 2 $$, “TR – [1992: 55]. The species externally is similar to Hydroporus Rize, kisdere, Sivrikaya Köyü, Köprü, Yanndaki Ilca, 29.6.2001, pubescens (Gyllenhal, 1808); it has, however, the last ab- Erman, Ö.K. leg.”; specimens rather small, TL ca. 3.3 mm (CKE, dominal sternum distinctly reticulated, while that of H. pube- CHF). 2 exs., “Turcia 3.IV.2000, Bolu vil. [= province], 10 km scens is smooth. It is equally similar to H. discretus, howev- E Gerede, J. Hájek & M. Mikát leg.”, “ Hydroporus discretus ponticus Zaitzev, Ji øí Hájek det. 2001” (CJH); 1 #, 1 $, er, on average slightly longer (TL 3.4–4.0 mm; H. discretus: “18.7.1992 TR Erzurum, MT Palandöken, stream on road TL 3.25–3.5 mm, reaching 3.8 mm in some southern Europe- Erzurum – Çam, 2300 m, Toledo leg.” (CHF); 1 #, “21.7.1997 an specimens), and the colour of the upper surface is dark (TR) prov. Erzurum, Tortum – Uzundere road, ca. 5 km N brown and not black as it is at least in central and northern Tortum, artesian well under bridge, Ö.K. Erman leg.” (CHF); 6 European populations of H. discretus. In most specimens, exs., “26.4.2000 (TR) Erzurum, ca. 10 km S Tortum, ca. 2 km the elytra are lighter near the base; in a few specimens from N Güzelyayla pass, ponds on meadow, Fery leg.” (CHF); 8 exs., Iran and Turkmenistan we have even observed distinct yel- “26.4.2000 (TR) Erzurum, ca. 14 km S Tortum, ca. 2 km S lowish spots. The male genitalia are given in Figs 4–6. In Güzelyayla pass, ponds on meadow, Fery leg.” (CHF); 1 #, 2 $$, lateral view, the penis is slightly more curved in apical third “TR – Erzurum, Oltu, Güryaprak Köyü Gölleri, 22.9.2001, Erman, Ö.K. Leg.” (CKE, CHF); 1 #, 1 $, “TR – Erzurum, Oltu, and lacks the convexity of the corrugated membrane in the Çarzof Köyü Gölleri, 22.9.2001, Erman, Ö.K. Leg.” (CKE); 1 #, 2 $$, “TR – Erzurum, Oltu, Göçedere, 22.9.2001, Erman, Ö.K. * This locality is either located in Siirt Province (ca. 37.53N Leg.” (CKE); 1 $, “TR – Erzurum, enkaya, Turnab Köyü, 41.59E) or in Van Province (ca. 38.09N 43.09E). Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus (Dytiscidae) 255 middle part of the ventral surface, which is so characteristic Gschw” [greyish brown, hw Gschwendtner], “Coll. Gschwendt- of H. discretus (cf. Fig. 2). ner” [printed], “Goldschmidti Gschw., det. Gschwendtner” [hw DISCUSSION. Although H. transgrediens can be distin- Gschwendtner?], “Type” [red, hw?], “Paralectotypus #, Hydro- guished from “normal” central European H. discretus by its porus goldschmidti Gschw, det. G. Wewalka [19]91” [red, hw Wewalka] (OLML); 1 $ (strongly matt), “Issyk - kul, Ton-Fluss.” larger total length and differences in the shape of the penis, [hw?], “Type, Gschw” [greyish brown, hw Gschwendtner], “Coll. we cannot exclude the possibility that future investigations Gschwendtner” [printed], “Goldschmidti Gschw., det. Gschwendt- will reveal that the species belongs to a difficult complex of ner” [hw Gschwendtner?], “Type” [red, hw?], “Paralectotypus #, species, varieties and forms around H. discretus (see, e.g., Hydroporus goldschmidti Gschw, det. G. Wewalka [19]91” [red, Nilsson [2001: 156]), which is distributed over an enormous hw Wewalka] (OLML); 2 $$ (weakly matt), “Issyk - kul, Ton- area reaching from the Canaries, North Africa; most of Eu- Fluß.” [hw Gschwendtner], “Cotype, Gschw” [greyish brown, hw rope; large territories in Russia; the Near, Middle, and Far Gschwendtner], “Paralectotypus $, Hydroporus goldschmidti East; and China. Not only the two species and the varieties Gschw., det. G. Wewalka [19]91” [red, hw Wewalka]; one and forms mentioned above must be included into such stu- specimen with additional “7211, Goldschmidti” [hw?] (CGW). Zimmermann [1931: 131] studied at least a part of the syntype dies, but also other taxa which at present are treated as valid series. It is assumed that he kept this specimen for his species, such as Hydroporus errans Sharp, 1882 and Hydro- collection. Wewalka [1992: 54] gave for “Bagratsch-Kul”: porus feryi Wewalka, 1992. “China, Sinkiang-Uighur, Bagratsch-Kul or Po-ssu-teng Hu”. Hydroporus goldschmidti Gschwendtner, 1923 Today this lake is called “Bosten Hu”; it is in Xinjian province, its co-ordinates are ca. 42N 87E [H. Schütze in litt.]. Gschwendtner, 1923: 101 Lectotype of Hydroporus goldschmidti var. recidivus: $, Type locality — Kyrgyzstan, Issyk-Kul, Ton [= Tong] river* abdomen without inner parts, “Thian shan, Aksu-Musart, coll. = Hydroporus goldschmidti var. recidivus Gschwendtner, 1923: Winkler” [printed], “Type, Gschw” [greyish brown, hw Gschwendt- 103 syn.n. (Type locality — China, Xinjian, Tian Shan, be- ner], “Goldschmidti var recidivus Gschw” [hw Gschwendtner], tween Aksu and Musart pass**) “Coll. Gschwendtner” [printed], “Type” [red, hw?], “Lectotypus $, Hydroporus recidivus Gschw, det. G. Wewalka [19]91” [red, Zimmermann, 1931: 130, 131 ( goldschmidti var. recidivus); hw Wewalka] (OLML). Paralectotypes: 1 $, abdomen absent, Zaitzev, 1953: 164 (including goldschmidti var. recidivus), first two labels as in the lectotype except “Cotype” instead of 1972: 173 (including goldschmidti var. recidivus); V.B. Gue- “Type”, “Hydroporus Goldschmidti var. recidivus Gschw” [hw orguiev, 1963: 216 (as possible synonym of H. glasunovi Gschwendtner], “Paralectotypus $, Hydroporus goldschmidti Zaitzev, 1905); Wewalka, 1992: 54 (lectotype designation of Gschw, det. G. Wewalka [19]91” [red, hw Wewalka] (CGW); 1 H. goldschmidti ), 58 (lectotype designation of H. goldschmid- #, first two and fourth label as in the lectotype, third label as ti var. recidivus); Nilsson, 1995: 53, 2001: 157, 158 ( recidi- in the female paralectotype, “ Hydroporus ? goldschmidti vus), 2003: 61, 64 ( recidivus). Gschw, det. G. Wewalka [19]91” [hw Wewalka], “Paralectotype, MATERIAL. Type material : Lectotype of Hydroporus Hydroporus recidivus Gschwendtner, 1923, Fery 2000” [red] goldschmidti: #, “Jssyk - Kul, Ton-Fluss.” [printed], “Type, (OLML). Notes: This male became automatically a paralecto- Gschw” [greyish brown, hw Gschwendtner], “Coll. Gschwendt- type when Wewalka [1992: 58] selected the lectotype from the ner” [printed], “ Hydroporus Goldschmidti # Gschw.” [hw syntype series. This is why we have provided this specimen with Gschwendtner], “Type” [red, hw?], “Lectotypus #, Hydroporus the respective red label, although Wewalka refrained to do so, goldschmidti Gschw, det. G. Wewalka [19]91” [red, hw Wewal- because he believed — correctly — this specimen to belong to ka] (OLML). Paralectotypes: Kyrgyzstan: 1 $ (strongly matt), H. goldschmidti, not to H. recidivus. “Jssyk - Kul, Ton-Fluss.” [printed], “Type, Gschw” [greyish brown, Additional material examined: Uzbekistan: 1 #, 2 $$, hw Gschwendtner], “Coll. Gschwendtner” [printed], “ Hydro- “Uzbekistan, Tashkent Province, Yakka-tut near Burchmulla, porus Goldschmidti $ Gschw.” [hw Gschwendtner], “Type” [red, 26.7.–3.8.2004, 41°38’N 70°03’E, Hendrich leg. Loc. 1a”; see hw?], “Paralectotypus $, Hydroporus goldschmidti Gschw, det. Hendrich & Hendrich [2005: 430] (CLH). Kyrgyzstan: 1 $, G. Wewalka [19]91” [red, hw Wewalka]; 1 $ (weakly matt), “Kirghizia: Issyk - Kul bank S Chon-Urjukty [ca. 42.43N 77.50E], “Jssyk - kul, Ton-Flu ß.” [hw Gschwendtner], “Type, Gschw” 19.–23.VI.1993 1600 m, leg. Schawaller”; 1 $, “Kirghizia: Chatkal- [greyish brown, hw Gschwendtner], “Coll. Gschwendtner” skij Alatau, Sary-Celek [near Arkit, ca. 41.47N 71.56E], 27.– [printed], “Hydroporus Goldschmidti $ intersex. Gschw.” [hw 31.V.1993, 1400–1600 m, leg. Schawaller” (all in SMNS). 2 ##, Gschwendtner], “Type” [red, hw?], “Paralectotypus $, Hydro- 3 $$ (all smooth), “Kyrgyzstan: Sandalasch G.K. [= Gebirgs-Kette porus goldschmidti Gschw., det. G. Wewalka [19]91” [red, hw = mountain range], Chatkal riv., ca. 2000 m, 5.VII.1997, leg. Dolin Wewalka] (all in OLML). China: 1 #, “Ost-Turkestan, Bagratsch- (K – 9/97)” [ca. 41.50N 70.55E, Dolin in litt.]; 3 ##, 6 $$ (all Kul” [printed], “Cotype, Gschw” [greyish brown, hw Gschwendt- smooth), “Kyrgyzstan: Sandalasch G.K. [= mountain range], Chap- ner], “Hydroporus Goldschmidti Gschw.” [hw Gschwendtner], chama Pass env., 1600 m, 6.VII.1997, leg. Dolin (K – 10/97)” [ca. “Type” [red, hw?], “Paralectotypus #, Hydroporus goldschmidti 41.40N 70.50E, Dolin in litt.]; most probably the Chap-Chyma Gschw, det. G. Wewalka [19]91” [red, hw Wewalka] (OLML); 1 Pass, ca. 41.32N 70.48E [see Schütze & Kleinfeld 1997: 94]; 1 $, $ (strongly matt), “Ost-Turkestan, Bagratsch-Kul” [printed], “Kyrgyzstan, Ferganski range, nr. Bergut pass [not found on any “Type!” [hw Zimmermann], “Samml. A. Zimmermann”, “Coty- map], 2015 m, 24.6.1996, leg. Dolin (n 37)”; specimen with pus, Hydroporus Goldschmidti Gschw., Zool. Staatssammlung pronotum totally reticulated, but elytra smooth (all in NMW); 4 München” [red, hw? non-authorised curatorial designation], ##, 3 $$, “Kyrgyzstan, W Tien Shan, Chatkal gorge, Stab [not “Paralectotype, Hydroporus goldschmidti Gschwendtner, 1923, found on any map] env., 29.VII.1998, 1800 m, leg. Dolin (K – 9/ Fery 2004” [red] (ZSM). Notes: Judging by the original 98)” (NMW); 3 ##, “Kyrgyzstan, Tshatkal valley, Shtab 2000 m, description, Gschwendtner did not know females of this 28.07.[19]99, V. Dolin” (CJH); 2 ##, “Kyrgyzstan, Mt. Rg., species with smooth elytra. Most probably, he used the term Tshatkal, 2000, Shtab, 28 \2906[19]98” (CJS); 13 ##, 1 $ “intersex” in the meaning of “reticulation of the upper surface (smooth), “28.7.1999 Kyrgyzstan, Tschatkal mountain range, nr between that of strongly reticulated females and that of Stab, Bach und Pfützen, Dolin leg.”; found together with Hydro- smooth males”. 1 #, “Issyk-Kul, Ton-Fluss.” [hw?], “Type, porus glasunovi Zaitzev, 1905 (NMW). China: 1 $, “Thian-Shan, Musart” [printed], “Hydroporus goldschmidti var. recidivus Gschw., *According to Sch ütze & Kleinfeld [1997: 143], the Tong det. G. Wewalka [19]72” [hw Wewalka], “ Hydroporus recidivus river rises ca. at 41.52N 77.02E and flows into the southern Gschw, det. G. Wewalka [19]91” [hw Wewalka]; 2 ##, 5 $$ (all shores of the Issyk-Kul near 42.11N 76.55E $$ matt), “Bagratsch-Kul” [some labels printed, others hw **This locality should have about the following co-ordinates: Wewalka], the males with “ Hydroporus # goldschmidti Gschw, 42.30N 80.40E det. G. Wewalka [19]91” [hw Wewalka], the females with similar 256 H. Fery, P.N. Petrov labels, but with the female sex symbol or without any such symbol 6.[18]67” [hw Schaufuss], “Coll. L.W. Schaufu ß” [printed], (all in CGW). 1 #, 1 $ (matt), “Ost-Turkestan, Bagratsch-Kul” [hw “distinguend. var estrellensis, Estrella Schauf.” [hw Schaufuss], Wewalka], “Locus typicus aus der Typenserie [= locus typicus of “Zool. Mus. Berlin”, “Lectotype, Hydroporus distinguendus the type series], G. Wewalka” [hw Wewalka in part], “Hydroporus var. estrellensis Schaufuss, 1882, des. H. Fery 2005” [red, goldschmidti Gschw., det. G. Wewalka [19]72” [hw Wewalka in printed] (MNB). Paralectotypes: 1 $, the specimen is part]; specimens with head brownish, but not as light as in the provided with copies of lectotype labels; 1 $, “Estrella, 18/ types of H. recidivus (MRTO). Inexact localities: 1 #, “Turke- 667” [hw Schaufuss], “Coll. L.W. Schaufu ß” [printed], “Zool. stan”; 1 $ (smooth) “Samarkand” (all in NMW). Both specimens Mus. Berlin”; 1 #, “Type” [printed], “ Hydr. disting. var are rather old and have the head as brownish as in the types of estrellensis Schauf, 18/6 1867 Estrella” [hw?], “Zool. Mus. H. recidivus. Samarkand is certainly used in the sense of the 19th Berlin”; 1 #, “Type” [printed], “ Hydr. disting. var estrellensis, century Russian Empire area of Samarkand which enclosed today’s Schauf. I. 18/6 1867 Estrella” [hw?], “Zool. Mus. Berlin” (all southern Uzbekistan and northern Tajikistan (see Schütze & in MNB). Each paralectotype is provided with its respective Kleinfeld [1997: 80]). Definitely, this specimen has not been red label. Notes: Lectotype and the first paralectotype collected in China. originally were glued onto the same card; they have been DESCRIPTIVE NOTES. Hydroporus goldschmidt iresem- separated by us and each glued onto its own card. The bles externallyH ydroporus planus (Fabricius, 1781), and shares lectotype lacks both hind tarsi and the last two right mid with this species the reticulated last abdominal sternum. It has, tarsomeres. however, the sides of the pronotum — at least the lateral rim — Additional material examined : Latvia: 1 $, “Kurland, in most specimens brownish, the elytra of more reddish brown, Sundur, Aug. 1917, leg. Salchert” (MNB). Georgia (first and the total length on average smaller. The lighter spots record): 7 exs., “SSSR–Caucasus, Sno [ca. 90 km N Tbilisi, behind the base of elytra are often rather distinct, whilst the ca. 42.35N 44.38E] 1800 m, 26.6.[19]57 Øíha” (CHF). elytral base of H. planus is only rarely considerably lightened. Turkey: 10 exs., “Uludag, 1850 m, 2.VIII.1984”, “Türkey Body length and shape are strongly variable, as well as the [sic!], R. Kinzelbach et al.”, “84/23” (NMB, CHF). Specimens extension of the pronotal reticulation, and cannot be used to with rather light elytra; rim of pronotum in most specimens separate the two taxa as it has been done by Wewalka [1992] brownish. Azerbaijan: 130 exs., “17.5.1999 Azerbajdzhan: who constructed, however, his key on the base of only a few Talysch [= Talysh, mountains in southern Azerbaijan], Hir- kan-Wald, Aurora Umg. [= environs], 600 m, Dolin leg.”; 2 examined specimens. On the other hand, H. goldschmidti and ##, 2 $$, “17.5.1999 Azerbajdzhan: Talysch, Hirkan-Wald, H. planus can be easily separated by the shape of the male Parakent Umg., 20–180 m, Dolin leg.” 6 exs., “25.5.1999 genitalia (cf. Figs 7–9 and 10–12). Azerbajdzhan: Talysch, Suwant (Plateau), 1760 m, Dolin MEASUREMENTS. TL: 3.3–4.2 mm, MW: 1.7–2.1 mm, leg.”; 30 exs., “Azerbajdzhan: Talysch, Zuwant plateau, 1500 TL/MW: 1.9–2.1 (lectotype of H. goldschmidti: TL: 3.9 mm, m, Gosmoljan env. [ca. 38.40N 48.21E], 27.5.2000, leg. Dolin MW: 2.0 mm; lectotype of H. recidivus: TL: 4.0 mm, MW: (AT 5)”; 1 #, 3 $$, “31.5.1999 Azerbajdzhan: Talysch, Astara 2.1 mm). Umg. [= environs; ca. 38.28N 48.52E], Bach und Fluss, Dolin DISCUSSION. The area of distribution of the original leg.” (all in NMW); 1 $, “Palekesh env. [ca. 38.30N 48.37E], type series of H. goldschmidti is in the Tian Shan mountain 13.–15.V.1992”, “Aserbaidschan, A. Schamaev” (NMB); 1 $, range, and reaches from eastern Kyrgyzstan to north-west- “Azerbaijan Talysch Mts., km 14–18, Rte Lenkoran–Lerik, ern China (Xinjian). The locus typicus of H. recidivus is 30.IV.–9.V.2001, T. Lackner”, “Coll. Hendrich, Berlin”; 1 $, within H. goldschmidti’s distribution area. Furthermore, one “Azerbaijan Talysch Mts., Lerik mountains, 1700m, 1.– paralectotype of H. recidivus is very similar to H. gold- 3.V.2001, T. Lackner”, “Coll. Hendrich, Berlin” (all in CLH); Iran: 1 #, “Iran: Gilan, 20 km E Khalkhál, 1900 m, 13.– schmidti; in particular, the penis coincides absolutely with 16.V.2000, leg. J. Rejsek” (SMNS); 1 #, “Iran, centr. Gilan, that of H. goldschmidti (see also Wewalka [1992: 58]). In Masuleh, (37.10N/48.58E), 55 km WSW Rasht, J. Rejsek addition, we have not found significant differences in the 15.–17.4.[19]99”; 1 #, “Iran, NE Mazandaran, Nahar Khoran, female genitalia of the two taxa. The only differences be- (36.45 N/54.28 E), 6 km south Gorgan, J. Rejsek 15.– tween H. recidivus and H. goldschmidti are the colour of the 17.4.[19]99”; 1 $, “Iran, W Mazandaran, 10 km NW Hasan head and the pronotum: head in large parts reddish brown in Keif, (36.36N/51.10E), J. Rejsek 20.–21.4.[19]99” (all in H. recidivus, only at anterior and posterior margins (else- CJS); 1 #, “7.8.1995 Iran, W. Azerbayjan, 75 km N Saghez, where dark brown) in H. goldschmidti; sides of pronotum wetland (t15), 1400 m, Elmi leg. (#1650)”; 1 #, “2.9.1998 lighter brown to rather large extent in H. recidivus, only rim Iran, Ardebil, 53 km W Assalem, stream, 1980 m, Elmi leg. of pronotum lighter brown in most H. goldschmidti. All these (#2131)”; 1 #, “18.4.2001 Iran, Fars, 17 km SE Sepidan (= observations induced us to treat H.recidivus as a junior Ardakan), Shesh Pir”, “ditch with running water, Elmi & Fery subjective synonym of H. goldschmidti. leg.” (all in CHF). Tunisia: 1 #, “Tunesien, 28.3.[19]91, DISTRIBUTION. China (Xinjian), Kyrgyzstan, Uzbekistan. Toujane, Quelle [= spring], leg. Wewalka (T3)” [hw Wewalka] (NMW). 9 exs., “7.4.[19]92 Tunesien, NE Tunis, Carthago, Hydroporus pubescens (Gyllenhal, 1808) Rinnsal [= streamlet], Fery leg.”; 2 exs., “8.4.[19]92 Tunesien, S Tunis, El Zahra, Tümpel [= pond], Fery leg.” (all in CHF). Gyllenhal, 1808: 536 ( Hyphydrus pubescens) Algeria: 1 #, “Edough, juin [18]85”, coll. Bedel (MNHN). Type locality — Sweden Russia (first record from Asian Russia — the Urals): 1 #, = Hydroporus distinguendus var. estrellensis Schaufuss 1882: 3 $$, “Chelyab. obl. [= Chelyabinskaya oblast = Chelyabinsk 559 (Type locality — Portugal, Serra da Estr êla)* area], Ilmensk. zap. [= Ilmenskiy zapovednik = Ilmenskiy Stephens, 1828: 61; Zimmermann, 1920: 100 (including syno- Nature Reserve], 22.VI.1941” [Cyrillic] (DEUM). nymy of H. distinguendus var. estrellensis); Wewalka, 1992: 59; DESCRIPTIVE NOTES. Specimens from the southern Nilsson & Holmen, 1995: 50; Nilsson, 2001: 158, 2003: 64. part of the distribution area often have the elytra — in MATERIAL. Syntypes of Hyphydrus pubescens not stud- particular the basal part — somewhat lighter yellowish brown. ied. Type material: Lectotype of Hydroporus distinguendus In specimens from Iran these markings are particularly light var. estrellensis (by present designation): #, “Estrella, 18/ and rather sharply delimited. DISTRIBUTION. Europe; Russia (at least southern Eu- *Usually Hydroporus distinguendus Desbrochers des Loges, ropean regions and the South Urals); North Africa except 1871 is treated as junior subjective synonym of Hydroporus jonicus L. Miller, 1862 Egypt; Turkey; Cyprus; the Near East; Iran. Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus (Dytiscidae) 257 13 14 15 16 17 18 Figs. 13–18. Male genitalia of Hydroporus spp.: 13–15 — H. limbatus; 16–18 — H. brucki.; 13–15, 16–17 — penis; 15, 18 — paramere; 13, 16 — ventral view; 14–15, 17–18 — lateral view. Ðèñ. 13–18. Ãåíèòàëèè ñàìöîâ Hydroporus spp.: 13–15 — H. limbatus; 16–18 — H. brucki.; 13–15, 16–17 — ïåíèñ; 15, 18 — ïàðàìåðà; 13, 16 — ñíèçó; 14–15, 17–18 — ñáîêó. Hydroporus limbatus Aubé, 1838 specimen placed in the drawer behind a label “ limbatus Aubé” Aubé, 1838: 292 [hw Wehncke] (MNHN). This specimen originally was moun- Type locality — Italy, Sardinia ted on a pin, and it has been glued onto a rectangular card by the senior author. Zimmermann 1931: 134; Guignot, 1959: 390; Franciscolo, DESCRIPTIVE NOTES. This species is similar to H. 1979: 336; Wewalka, 1992: 48; Nilsson, 2001: 157, 2003: 62. brucki, but the punctation of the elytra is coarser and less MATERIAL. Type material : Lectotype of Hydroporus dense; the puncture lines are only exceptionally and very limbatus (by present designation): $, “Limbatus” [hw Aubé], weakly perceptible. The yellowish margin on sides of prono- “Museum Paris, Coll. Aubé”, “Lectotype, Hydroporus limbatus tum is somewhat narrower. In lateral view, the margin of the Aubé, 1838, Fery des. 2004” [red, printed]; specimen originally elytra is only weakly ascending towards humeral angle. Ac- pinned, placed in the drawer behind a label “1975 Hydroporus cording to Franciscolo [1979: 336] the total length is 4.0–5.0 Clairville, Limbatus Aubé” [hw Aubé] (MNHN). Notes: The lectotype has been glued onto a rectangular card by the senior mm; the examined specimens have 4.0–4.7 mm (lectotype 4.0 author. It lacks the last six articles of the right antenna and the mm). The male genitalia are given in Figs 13–15; compare last tarsomere of the left hind leg. Behind the lectotype stands those of the externally similar H. planus in Figs 10–12. a female prepared in the same manner; it does not bear any DISTRIBUTION. Italy, France, Spain, Portugal, Moroc- label, except “Museum Paris, Coll. Aubé” (MNHN). This co, Algeria, Tunisia, and Libya. In Franciscolo [1979: 336], specimen is a Hydroporus lucasi Reiche, 1866, as can be easily Corfu, Attica, and Crete are given additionally; these records, seen by the reticulated last abdominal sternum and the dense however, no doubt must be attributed to H. brucki. and rugose punctation of its apical third. This specimen is not treated as paralectotype because it does not fit some details of Hydroporus brucki Wehncke, 1875 the original description of H. limbatus, e.g., the rather coarse Wehncke, 1875: 234 punctation of elytra and the absence of puncture lines. Aubé’s Type locality — Greece, Thessaloniki [“Saloniki”]* description of H. limbatus suggests that the author had only a single specimen at his disposal. Since this is, however, not stated Zimmermann, 1931: 135; Zaitzev, 1953: 165, 1972: 174; explicitly, the existence of syntypes instead of a holotype must Wewalka, 1992: 48; Nilsson, 2001: 156, 2003: 60. be assumed (Recommendation 73F of the ICZN [1999]). MATERIAL. Type material : Lectotype of Hydroporus Additional material examined : Portugal: 1 $, “Portugal brucki (by present designation): $, “Brucki” [hw Wehncke], mer. reg. Faro, Trafal /1/, 02.03.2004 Manti lgt., louka-sladká “Turcia, Salonichi” [hw Wehncke], “ Hydroporus brucki Wenck., voda” (CJH); 1 $, “7.11.[19]89 Portugal, Algarve, Vila Real, aus Coll. Oberthur” [hw Wewalka], “Muséum Paris, ex coll. R. Tümpel, Fery leg.”; France: 11 exs., “4.2.[19]96 France, Ca- Oberthur, ex Wehncke”, “Lectotype, Hydroporus brucki Wehn- margue, Étang du Charnier, Fery leg.”; 1 #, 1 $, “Il [sic!] de Ré”; cke, 1875, Fery des. 2004” [red, printed]; specimen originally 1 $, “27.9.[19]88 France, Camargue, Aigues-Mortes, Lache, Fery glued onto a small triangular tip, placed in the drawer behind leg.”; 1 #, “31.3.[19]86 France, Herault, Agde, Wassergraben, a label “Brucki mihi” [hw Wehncke] (MNHN). Paralectotype: Fery leg.”; Italy: 1 $, “Sardinien 24.5./20.6.[19]65, Laloletta, 1 $, a very small quadratic label without any text, “Muséum Budberg”; 8 exs., “14.7.[19]81 Italia, Sardegna, Fertilia, Alghero, Paris, ex coll. R. Oberthur, ex Wehncke”, and the respective red Tümpel, Fery leg.”; 2 exs., “Italie: Sicile – Catania, embouchure paralectotype label; specimen placed behind the lectotype, du Simeto, 11.04.[19]99, leg. G. Carron” (all in CHF). Unknown prepared in a similar manner as the lectotype (MNHN). Notes: locality: 1 #, “Hydroporus exotique [?], velubinus mihi., h. Both specimens have been glued onto larger rectangular cards [Dejean often used an “h.” as abbreviation of “habitat”, but here by the senior author. The lectotype lacks the left protarsus, it was not followed by any locality.] D. Latreille”; a greyish and three right protarsomeres, left mesotibia and mesotarsus, four shiny label, text almost illegible, only perceptible if appropri- right mesotarsomeres, left metatarsus, and the last right ately illuminated; the word “exotique” in red ink and of metatarsomere. The paralectotype lacks three left mesotarsom- unknown hw, the rest in black ink, hw most probably that of eres and the right metatarsus. Dejean; on the reverse “1221, 1222, 1223” [hw?], “Ex Mus æeo Dejean” [printed], “limbatus” [hw Régimbart?], “D. Sharp Monogr.” *In 1875, Thessaloniki belonged still to the Turkish (Otto- [printed], “Muséum Paris, ex coll. R. Oberthur, ex Wehncke”; man or Osman) Empire 258 H. Fery, P.N. Petrov Additional material examined : Greece: 1 #, “Griech- 21.IV.1992, leg. Schawaller”; 23 exs., “Appennino Romagnolo, enland 1991, Ebrosebene (2), leg. Schödel 16.V.” (NMW); 1 #, Passo d. Muraglione, m 850 21.2.1993, leg. F. Pederzani” (all “Corfu, Paganetti”; 1 #, “Kephallenia 1905, Argostolion, O. in CHF). Leonhard”; 4 exs., “Kephallenia, Paganetti”; 1 #, “GR (38), DESCRIPTIVE NOTES. Hydroporus analis and H. de- Makedonia (CHA), Chalkidiki, Chlomón Oros, Tal am Südhang cipiens are the only members of the planus-group that have a QU 32, 16. 650 m, 29.05.[19]91., leg. CIT & GER”; 1 ex., strongly rugose abdomen. Their upper surface is similar to “Olympos, Graecia, 2100 m, 4.6.[19]37, Coll. Barto ò”; 1 ex., “Gr. that of the lighter coloured H. pubescens specimens. The Ins. Lesbos, 4 km W. Agiassos, leg. Malicky 26.5.[19]75”; 1 ex., male genitalia of H. analis are shown in Figs 19–21. The total “GR 95–21: Insel Lesbos, SE, 12 km W Mitilíni, Flußmündung length of the specimens studied is 3.4–3.8 mm (lectotype 3.7 [= river mouth], Tümpel [= pond]; leg. D. Grimm. 17.– mm). Franciscolo [1979: 331] gives 3.3–3.5 mm, Zimmer- 21.V.1995”; 10 exs., “GR Korinth 12.6.[19]86, Kato Alephori, leg. H. Hebauer” (all in CHF). mann [1931: 134] 3.5–3.75 mm. DESCRIPTIVE NOTES. The punctation of the elytra is DISTRIBUTION. Italy, France (Corsica); records from rather dense and fine, approaching almost that of Hydroporus Austria, Greece, Algeria, and Tunisia [Franciscolo, 1979: marginatus (Duftschmid, 1805). Other characteristic features 341] should be confirmed; those from North Africa possibly are the broadly brownish sides of the pronotum and smooth must be attributed to H. decipiens. abdominal sterna (in contrast to H. planus), the distinct punc- Hydroporus decipiens Sharp, 1878 ture lines on the elytra, and the margin of the elytra distinctly ascending towards humeral angle (in contrast to H. limbatus). Sharp, 1878: 113 According to Zimmermann [1931: 135], its total length is 4.0– Type locality — Spain, Sierra de Guadarrama 4.3 mm, yet most of the examined specimens are distinctly Zimmermann, 1931: 134 ( analis decipiens ); Franciscolo, 1979: longer: 4.5–4.7 mm, only one of these has 4.3 mm, another one 340 (analis [= decipiens]); Fery & Fresneda, 1988: 342 ( analis 4.4 mm; the total length of the two types is 4.5 mm. The male decipiens); Ribera et al., 1999: 57; Nilsson, 2001: 156, 2003: 60. genitalia are given in Figs 16–18. MATERIAL. Type material : Lectotype of Hydroporus DISTRIBUTION. Greece, Turkey, Cyprus, and Lebanon. decipiens (by present designation): $, “Hydroporus decipiens Sharp, Type D.S.” [on glue card, hw Sharp], “Type” [printed, Hydroporus analis Aubé, 1838 round label with red margin, most probably mounted by J. Balfour-Browne], “Spain” [printed], “Sharp Coll, 1905 – 313” Aubé, 1838: 294 [printed], “Guadarrama” [hw Sharp], “Type 410, Hydroporus Type locality — Italy, Sardinia decipiens, Europe” [hw Sharp], “Lectotype, Hydroporus deci- Zimmermann, 1931: 133; Guignot, 1959: 394; Wewalka, 1992: piens Sharp, 1878, Fery des. 2005” [red, printed] (BMNH). 47; Nilsson, 2001: 156, 2003: 60. Paralectotype: 1 $, “Hydroporus decipiens Sharp, Ind. typ. D.S., MATERIAL. Type material : Lectotype of Hydroporus Guadarrama” [on glue card, hw Sharp; the word “ decipiens” on analis (by present designation): #, “Sardaigne” [a small round a small piece of paper glued onto the card], “Spain” [printed], label, hw Aubé], “Museum Paris, Coll. Aubé”, “Lectotype, “Spain” [printed], “Sharp Coll, 1905 – 313.” [printed], “Syntype” Hydroporus analis Aubé, 1838, Fery des. 2004” [red, printed]; [printed, round label with light blue margin, most probably specimen placed in the drawer above a label “1976 Hydroporus mounted by J. Balfour-Browne], and the respective red paralec- Clairville, Analis Aubé” [hw Aubé?] (MNHN). Notes: The totype label (BMNH). Notes: The lectotype lacks the last three lectotype was originally pinned; it has been glued onto a articles of the right antenna. R. Booth (BMNH) kindly rectangular card by the senior author. It lacks two left communicated: “There is a label in the collection, probably in protarsomeres and the right hind leg. In the original descrip- Garth Foster’s handwriting, indicating that H. decipiens is a tion Aubé wrote “Il a été trouvé en Sardaigne et fait partie de valid species and not a synonym of H. analis.” According to the la collection de M. Chevrolat”. We have not been able to locate original description, Sharp has studied at least one further syntypes of H. analis in the collections of the IRSN or BMNH specimen from Monchique (southern Portugal) collected by where parts of the collection of Chevrolat are said to have been Camille van Volxem. In the BMNH, however, no further preserved (see Horn et al. [1990: 71)]. However, we have no material could be localised. doubt that the specimen from the collection Aubé has been Additional material examined : Portugal: 1 #, 2 ##, used by the author for his description. We assume that Aubé “9.8.[19]84 Portugal, Serra Estrella, b. Manteigas, Bach, 600 m, has kept the specimen for his collection after the study of Fery leg.”; 8 exs., “4.8.[19]89 Portugal, Serra Estrela, Umg. [= Chevrolat’s material. environs] Manteigas, kl. Bach, Fery leg.”; 1 #, “22.7.1991 (P) Additional material examined : France, Corsica: 1 #, Guarda-distr., Serra da Estr êla, N Manteigas, ca. 1100 m, “8.8.[19]74 Serriera, b. Porto (Corse), fast ausgetrock. Flu ß, Rinnsal, Fery leg.”; 7 exs., “9.8.[19]84 Portugal, Serra Estrella, Fery leg.”; 1 ex., “2.4.[19]80 Korsika, Serriera, Flu ß am Strand, südl. Guarda, Rinnsal, Fery leg.”; 9 exs., “14.8.[19]85 Portugal, Fery leg.”; 1 #, “3.4.[19]81 Korsika, Serriera, Lache am Strand, Serra Estrela, Penhas Douradas, Bach, 1150 m, Fery leg.”; 8 exs., Fery leg.”; 3 exs., “10.4.[19]81 Corse, Serriera (Porto), Tümpel, “14.8.[19]85 Portugal, Serra Estrela, Penhas Douradas, Bach, Fery leg.”; 12 exs., “10.4.[19]81 Corse, Serriera, Tümpel, Fery 1000 m, Fery leg.”; 9 exs., “21.7.[19]87 Portugal, Serra da Estrêla, leg.”; 1 ex., “18.7.[19]82 Corse, Serriera, Bach, Fery leg.”; 2 exs., Umg. [= environs] Torre, ca. 1900 m, Moortümpel, Fery leg.”; 1 “7.4.[19]80 Korsika, Francardo, Waldbach, Fery leg.”; 1 ex., ex., “30.1.[19]94 Portugal, Minho, Serra de Arga, Rinnsal, 800 m, “7.4.[19]80 Corse, Casamozza, ehem. Kiesgrube, Fery leg.”; 3 Fery leg.”; 1 $, “20.7.[19]87 Portugal, Odemira, Cercal, Rinnsal, exs., “13.4.[19]81 Corse, Casamozza, Tümpel, Fery leg.”; 1 $, Fery leg.” (all in CHF). Spain: 1 #, 1 $, “Esp. Salamanca prov., “13.4.[19]81 Corse, b. Barchetta, Flu ß Golo, Fery leg.”; 1 #, La Alberca 1000 m, 5.6.2002 R. Sejkora lgt.” (CJH); 15 exs., “24.7.[19]81 Corse, Sc. di St. Regina, Flu ß Golo, Fery leg.”; 1 #, “23.7.[19]87 España, Prov. Leon, Astorga (südl.), Bach/Tümpel, “28.7.[19]81 Corse, Nähe Fango, Bach, Fery leg.”; 25 exs., Fery leg.”; 1 #, “6.7.[19]92 Espa ña, Prov. Le ón, Noceda, NE “13.7.[19]83 Corse, Ponte Leccia, Stranciacone, Fery leg.”; 2 Ponferrada, Bach, Fery leg.”; 10 exs., “22.7.[19]87 Espa ña, Prov. exs., “19.4.[19]79 Korsika, Carg èse, Wiesenbach, Fery leg.”; 1 Zamora, Puente de Sanabria, Bach, Fery leg.”; 5 exs., “21.3.[19]89 ex., “26.7.[19]82 Corse, Südlich Carg èse, Lache, Fery leg.”; 1 ex., España, Prov. Zamora, Justel, östl. Sanabria, Bach, Fery leg.”; 12 “1.8.[19]74 Soccia (Corse), Tümpel, Fery leg.”; 8 exs., “8.4.[19]80 exs., “3.11.[19]89 España, Prov. Zamora, O. [= E of] P. d. Sanabria, Korsika, Nördl. Sagone, Flu ß, Fery leg.” (all in CHF). Italy: 2 Bach, Justel, Fery leg.”; 1 #, “4.4.1999 Spain, Burgos, ca. 3 km exs., “I. Sardinien (201), östl. Badde Suelzu, leg. Malicky W Soncillo, ca. 30 km E Reinosa, ponds on meadow, Fery leg.”; 19.10.[19]81”; 14 exs., “18.7.[19]81 Sardinien, Villagrande, 1 #, “6.7.[19]88 Espagna, S. d. Gredos, Avila, Tormellas, Bach, Bach, Fery leg.”; 8 exs., “18.7.[19]81 Sardinien, Villagrande, Fery leg.”; 2 exs., “9.7.[19]88 Espa ña, Prov. Avila, S. d. Gredos, Tümpel, Fery leg.”; 2 exs., “Sardinien: NU, Stat. Arzana 800 m, Gredos, 1800 m, Quelltümpel, Fery leg.”; 30 exs., “3.8.[19]89 Nomenclatural, taxonomic, and faunistic notes on selected species of Hydroporus (Dytiscidae) 259 19 20 21 22 23 24 25 26 27 Figs 19–27. Male genitalia of Hydroporus spp.: 19–21 — Hydroporus analis ; 22–24 — H. decipiens; 25–27 — H. zimmermanni; 19–20, 22–23, 25–26 — penis, 21, 24, 27 — paramere; 19, 22, 25 — ventral view; 20–21, 23–24, 26–27 — lateral view. Ðèñ. 19–27. Ãåíèòàëèè ñàìöîâ Hydroporus spp.: 19–21 — Hydroporus analis ; 22–24 — H. decipiens; 25–27 — H. zimmermanni; 19–20, 22–23, 25–26 — ïåíèñ, 21, 24, 27 — ïàðàìåðà; 19, 22, 25 — ñíèçó; 20–21, 23–24, 26–27 — ñáîêó. España, Prov. Avila, Sierra de Gredos, Gredos, Tümpel, 1800 m, 4.0 mm, paralectotype 3.9 mm). The male genitalia are fig- Fery leg.”; 3 exs., “9.7.[19]95 Espa ña, Prov. Avila, Sierra de ured for the first time (Figs 22–24). Gredos, Gredos, Rinnsal, 1800 m, Fery leg.”; 1 #, “22.5.[19]90 This taxon has been treated by most authors of the last España, Prov. Guadalajara, S. d. Alto Rey, Bustares, Bach, Wiese, century either as a synonym or a variety ofH . analis. Ribera et Fery leg.”; 2 exs., “16.6.[19]90 Espa ña, Prov. Guadalajara, S. d. al. [1999: 57] were the firsts who re-instatedH . decipiens as a Alto Rey, Bustares, Rinnsal, Fery leg.”; 1 #, “19.2.[19]90 España, valid species, but did not give any reason for their procedure. Prov. Guadalajara, Anquela del Ducado, salzh. Lache [= salty DISTRIBUTION. Portugal and Spain; see also the notes puddle], Fery leg.”; 8 exs., “24.7.[19]87 Espa ña, Prov. Madrid, Sierra de Guadarrama, Lozoya, Bach, 1300 m, Fery leg.”; 2 exs., under the distribution of H. analis. “1.7.[19]92 (E) Toledo, SW Los Yébenes, Bachtümpel, Fery leg.”; Hydroporus crinitisternus Shaverdo & Fery, 2001 2 exs., “27.5.[19]90 Espa ña, Prov. Cáceres, S Coria, Ca ñaveral, Bachtümpel, Fery leg.”; 1 #, “2/iv/1996 Spain Cácares [sic!], Shaverdo & Fery, 2001: 34 Arroyo de las Canteras, 5 km NE of Trujillo, granite stream, D.T. Type locality — Kazakhstan, Talassky Alatau range, 40 km Bilton leg.”; 1 $, “8.7.[19]95 (E) Palencia, Ledigos, E Sahagun, Rio SW Dzhambul [= Auliye-Ata] Cuenza, Bach, Fery leg.”; 1 ex., “27.5.[19]90 España, Prov. Ciudad Nilsson, 2001: 156. Real, W Retuerta del Bullaque, Rinnsal, Fery leg.”; 8 exs., “27.5.[19]90 Espa ña, Prov. Ciudad Real, W Retuerta del MATERIAL. Kazakhstan: 1 #, 1 $, “E Kazakhstan, S Bullaque, Bachtümpel, Fery leg.”; 1 ex., “4.8.[19]85 Espagna, Tarbagatay Mts., 4 km NE Petrovskoe [ca. 47.03N 82.18E], Sierra Nevada, Laujar, Paterna del Rio, Rio de Alcolea, ca. 100 1100 m, 22.VI.2001, leg. W. Schawaller” (SMNS). Distribution: This new record completes well the known m, Fery leg.”; 8 exs., “19.7.[19]87 Espa ña, Prov. Cadiz, nördl. Jimena d. l. Frontera, Zufluß d. Rio Hozgarganta, Fery leg.” (all distribution of this rather poorly known species, which so in CHF). far has only been recorded from south-eastern Kazakhstan DESCRIPTIVE NOTES. Externally similar to H. analis, and western Mongolia. The species seems to prefer the in particular, with respect to the rugose surface of abdomen. transition between the plain and mountainous regions. It Yellowish spots on elytra more prominent. On average lon- should be noted that the Mongolian locality and the Kazakh ger, total length of specimens studied 3.6–4.2 mm (lectotype ones are well connected by the Irtysh valley. 260 H. Fery, P.N. Petrov Hydroporus tessellatus-group DISCUSSION. The synonymy of H. habelmanni has been treated in different ways (cf. list of references above). Hydroporus tessellatus (Drapiez, 1819) Gozis [1914: 161] treated the taxon as a variety of H. tessel- latus, and some pages further [Gozis 1915: 178] as a variety Drapiez, 1819: 43 ( Dytiscus tessellatus ) Type locality — Belgium, Moosel of H. pubescens, however, remarking in a footnote that the majority of authors of that time preferred the former view. = Hydroporus habelmanni Wehncke, 1876: 76 syn.n. (Type Burmeister [1939: 226] called the taxon a “form” of locality — Dalmatia*) H.pubescens, but not expressly giving it infrasubspecific Heyden, 1883: 30 ( habelmanni), 1891: 62 ( pubescens var. rank (cf. ICZN [1999], Article 45.6, which might be applied habelmanni), 1906: 116 ( tesselatus var. habelmanni); Seidl- here analogously, although such cases are not treated by the itz, 1887: 72 ( pubescens var. habelmanni); Jacobson, 1908: 425 (pubescens habelmanni ); Zimmermann, 1920: 100 ( pu- Code). To our knowledge, Franciscolo [1979: 334] was the bescens habelmanni ); Burmeister, 1939: 226 ( pubescens f. first who gave it expressly infrasubspecific rank and called it habelmanni); Franciscolo, 1979: 334 ( pubescens ab. habel- “ab. habelmanni”. However, Wewalka [1986: 278] and Ange- manni); Wewalka, 1986: 278 ( pubescens habelmanni ); An- lini [1988: 47] still listed it as a subspecies of H. pubescens, gelini, 1988: 47 ( pubescens habelmanni ); Balke & Fery 1993: whereas B. Guéorgiev [2004: 398] was the first who pro- 93 (neotype designation of H. habelmanni ); Nilsson, 2001: vided this taxon simply as a synonym of H. pubescens. 166, 2003: 65. As shown above, populations of H. pubescens from the MATERIAL. Type material: Neotype of Hydroporus tessel- Mediterranean, which usually are longer, broader, more vault- latus designated by Balke & Fery [1993: 93], stored in the IRSN. ed, and have the base and the margins of elytra more yellow- Lectotype of Hydroporus habelmanni (by present designa- ish than more northern ones, in the last decades have been tion): $, a small quadratic label without text, “Dalmatia” [hw treated either as a subspecies or as a colour variety of this Wehncke], “ex Wehncke, Muséum Paris, ex coll. R. Oberthur”, species and called “ habelmanni”. The designation of the “Lectotype, Hydroporus habelmanni Wehncke, 1876, Fery des. 2004” [red, printed]; glued onto a small triangular tip, placed lectotype of H. habelmanni is at the same time a synonymi- in the drawer behind a label “Habelmanni mihi” [hw Wehncke] sation of this name with H. tessellatus. Thus, these southern (MNHN). Paralectotype: 1 #, “Steinheil” [? nearly illegible, hw populations of H. pubescens lose their name, but, at present, Wehncke], “Elba” [hw Wehncke], “ex Wehncke, Muséum Paris, we do not see the necessity to give them another one. ex coll. R. Oberthur”, and a respective red paralectotype label; placed behind the lectotype and prepared in the same manner Hydroporus zimmermanni G.[J.] Müller, 1926 (MNHN). Note. The lectotype is an immature Hydroporus Müller, 1926: 288 tessellatus (Drapiez, 1819) with a total length of 3.5 mm; it Type locality — Slovenia, Pivka [= St. Peter = San Pietro del lacks the left hind tarsus. Both types have been re-glued by the Carso], ca. 30 km E Trieste [Italy] senior author onto larger rectangular cards. NOTES. It was not possible to identify the paralecto- = Hydroporus bicolor J.[G.]M üller 1933: 202 (Type locality Slovenia, Pivka) type of H. habelmanni undoubtedly. It is an extremely imma- ture specimen of ca. 3.3 mm total length and resembles Müller,1928:260,1933:201;Zimmermann,1931:127; Gschwendt- H.pubescens insofar as it has the elytra as well as the last ner, 1939: 34 ( bicolor); Zaitzev, 1953: 167 (including bicol- or), 1972: 176 (including bicolor); V.B. Gu éorguiev, 1960: 27, abdominal sternum smooth between the punctures. On the 28 (bicolor), 1965: 492 (including bicolor), 1971: 13 (includ- other hand, the punctation on the apical third of this segment ing synonymy of H. bicolor); Nilsson, 2001: 167, 2003: 65. is rather sparse and not densely and almost rugosely punc- MATERIAL. Type material : Holotype of Hydroporus tate as it is typical for H.pubescens; the punctures are zimmermanni: #, “Carniolia, St. Peter, 9.4.[19]11”, “ Hydroporus distinctly separated one from another and the smooth spaces Guernei Rég.” [hw Zimmermann], “Museo Civico di Trieste”, between them clearly perceptible. The sculpture of this seg- “Typus” [red], “Zimmermanni Müll., Tipo” [green, hw J.[G.] ment resembles much more that of H. limbatus, a species Müller] (MCTR). Notes: Müller’s [1926: 288] original descrip- which has, however, usually a distinctly higher total length tion of H. zimmermanni contains no hint on the number of (see above). Additionally, due to the high immaturity it was syntypes studied. In a later publication [Müller 1928: 260] the not possible to identify the specimen by the shape of the author provided “… the single exemplaire of H. Zimmermanni aedeagus. A further reason to designate not the Elba speci- …”; Zimmermann’s [1931: 127] remark can be interpreted in the men but the “Dalmatia” one as lectotype is that the latter fits same way: “Type from St. Peter in Krain …”. Thus Müller has a single exemplare of a new species. So, this specimen is a better Wehncke’s description with respect to some minute holotype by monotipy (Recommendations 72.4.1.1 and 73.1.2. features which, however, shall not be explained here in detail. of the ICZN [1999]). In the Wehncke collection, there stands a third specimen Lectotype of Hydroporus bicolor (by present designation): behind the lecto- and paralectotype which, however, is not #, “Istr. [= Istria] St. Peter, 16.4.33, Springer” [printed, hw treated as belonging to the syntype series because it is not Springer (?) in part], “Petel. jezero” [hw Springer?], “Cotypus” mentioned in the original description: 1 ex., “Smyrna [= [red], “Museo Civico di Trieste”, “Lectotype, Hydroporus bicolor Izmir, Turkey]” [hw Wehncke], “ex Wehncke, Muséum Pa- J.[G.] Müller, 1933, des. H. Fery 2005” [red, printed] (MCTR); ris, ex coll. R. Oberthur” (MNHN). This specimen is — like Paralectotypes: 1 $, first four labels as in the lectotype; 1 $, the lectotype — an immature H. tessellatus. “S. Pietro, Petelinsko jezero” [hw Springer (?)], “Springer, 16.4.33” [hw Springer?], “Museo Civico di Trieste”, “Typus” [red], The designation of a specimen of H. tessellatus as lecto- “bicolor m., det. J. Müller” [hw J.[G.] Müller?] (all in MCTR). Each type is — at first sight — not in accordance with Recommen- paralectotype is provided with the respective red label. dation 74A of the ICZN [1999], since in the last decades Additional material examined: Slovenia: 1 ex., “S. Pietro, H.habelmanni was treated as a synonym of H. pubescens. It Petelinsko jezero” [hw Springer?], “Springer, 16.4.33” [hw seems, however, to be absolutely necessary to base the Springer?], “Museo Civico di Trieste”; abdomen externally glued designation of the lectotype on the specimen of clear identity onto the card, genitalia missing; 2 exs., “Istr. St. Peter, 16.4.33, and not on the one which might give rise to future doubts. Springer” [date hw Springer?], “Petel. jezero” [hw Springer?], “Museo Civico di Trieste” (all in MCTR). 2 exs., “Galško jezero [not found on any map], 7.V.1956”, “Slovenia, leg. Pretner”; *This coincides more or less with today’s Croatia genitalia absent (MRTO). 4 exs., “Petelinsko Jezero”, “7.V.1956.”,