Phytologxa (July 1991) 71(l):58-65. NOMENCLATURE. NOTES FOR THE NORTH AMERICAN FLORA. VI John T. Kartesz h Kancheepuram N. Gandhi The North Carolina Botanical Garden, Dept. of Biology, Coker Hall, University ofNorth Carolina, Chapel Hill, North Carolina 27599-3280, U.S.A. ABSTRACT The followingnew combinationsare proposed: Artemisia triden- tataNutt.ssp. spiciformis(Osterhout)Kartesz& Gandhi; Cistanthe parryi (A. Gray) Hershkovitz var. arizonica (J.T. Howell) Kartesz & Gandhi; Cistanthe parryivar. hesseae (Thomas)Kartesz & Gandhi; Cistanthe parryi var. nevadensis (J.T. Howell) Kartesz & Gandhi; and Cistanthe umbellata (Torr.) Hershkovitz var. caudicifera (A. Gray) Kartesz & Gandhi. Oonopsis foliosa E. Greene var. mono- cephala (A. Nels.) Kartesz & Gandhi; Pyrrocoma carthamoides Hook. var. cusickii (A. Gray) Kartesz & Gandhi; Pyrrocoma hirta (A. Gray) E. Greene var. sonchifolia (E. Greene) Kartesz & Gandhi; Pyrrocoma linearis (Keck) Kartesz & Gandhi; Pyrrocoma lucida (Keck) Kartesz & Gandhi; Pyrrocoma racemosa (Nutt.) Torr. & Gray var. pinetorum (Keck) Kartesz & Gandhi; Pyrrocoma race- mosa var. paniculata (Nutt.) Kartesz & Gandhi; Pyrrocoma uni- ftora (Hook.) E. Greene var.gossypina(E. Greene) Kartesz& Gandhi; Solidago simplex Kunth ssp. var. randii (Porter) Kartesz & Gandhi; Stenotus acaulis (Nutt.) Nutt. var. glabratus (D.C. Eat.) Kartesz & Gandhi; and Stenotus armerioides Nutt. var. gramineus (Welsh & F.G. Sm.) Kartesz & Gandhi. KEYWORDS: Floristics, nomenclature, Asteraceae, Portulacaceae, Artemisia, Cistanthe, Oonopsis, Pyrrocoma, Solidago, and Steno- tus INTRODUCTION Continuing with the "NOMENCLATURAL NOTES FOR THE NORTH AMERICAN FLORA" (Kartesz k Gandhi 1989, 1990a, 1990b, 1990c, 1991), a sixth note in the series is presented here toward advancing our understanding of North American plant names. The data pertaining to type specimens, mentioned in this article, were collected from literature. 58 Kartesz & Gandhi: Nomenciatural notes for North America VI 59 ASTERACEAE Artemisia tmdentata ssp. spiciformis In Goodrich, et al. (1985), the combination Artemisia tridentata Nutt. ssp. spiciformis (Osterhout) Goodrich &: McArthur, which was based on A. spici- formisOsterhout,was proposed intheabstract section ofthat work. Goodrich, et al. used this trinomial in the text and provided a reference (in the literature cited section of their work) to the place of publication of the basionym as: "Osterhout, G.E., 1900. New plants from Colorado. Bull. Torrey Bot. Club 27:506-508." Since the basionym actually appeared on p. 507, and since a page reference must be the page or pages on which the basionym was validly published, but not to the pagination ofthewhole publication (ICBN Art. 33.2, footnote 1), the new combination remains invalid. The invalidity of their new combination was indicated by Chapman (Taxon 35:767. 1986) and by the Spe- cial Committeeonvalid pubbcation (Taxon 35:768. 1986). Although basionym publication details were provided by Chapman and by the Special Commit- tee, neither indicated the acceptance ofthis trinomial; hence, the combination must not be attributed to any of them (Art. 34.1a). We accept the trinomial and provide the complete bibliography to validate the new combination. Artemisia tridentataNutt.ssp. spiciformis (Osterhout) Kartesz&: Gandhi, comb. & stat. nov. BASIONYM: Artemisia spiciformisOsterhout, Bull. Torrey Bot. Club 27:507. 1900. TYPE: U.S.A. Colorado: Larimer Co., North Park, 3 Sep 1899, Osterhout 2011 (?RM). Oonopsis, Pyrrocoma, and Stenotus The historical treatment and application of the Haplopappus Cass, com- plex has varied widely among North American authors. Lane, et al. (1987) considered Haplopappus sensu Hall to be an artificial genus. The complex has been viewed to represent a single genus (Torrey &: Gray 1842; Hall 1928), two genera, i.e., Haplopappus and Machaeranthera Nees (Cronquist &: Keck 1957; Munz & Keck 1973), or manygenera (Rydberg 1906; Correll k Johnston 1970; Turner & Hartman 1976; Clark 1979; Brown & Clark 1982; Turner & Sund- berg 1986; Nesom 1989a, 1990; Nesom & Morgan 1990; Nesom, et al. 1990). The authors of the third category restrict the application of Haplopappus to South America. Nesom &: Morgan provided a synopsis of the complex ac- counting for all sections of North American Haplopappus sensu Hall and their current placement. For the North American flora (north of Mexico), we do not recognize Haplopappus. The taxa, previously known as Haplopappusin the U.S.A., are assigned to the following genera: CroptilonRaf., EricameriaNutt., Hazardia E. Greene, HesperodonarZ. Greene, IsocomaNutt., Machaeranthera Nees, Oonopsis Nutt., Prionopsis Nutt., Pyrrocoma Hook., Stenotus Nutt., Tonestus A. Nels., and XylothamiaNesom, et al. The recognition ofthe above PHYTOLOGIA 60 volume 71(l):58-65 July 1991 genera necessitates several new combinations. We propose new combinations in Oonopsis, Pyrrocoma, and Stenotus. Note: Dr. R.A. Mayes (1976), in his Ph.D. dissertation (TEX), proposed several new combinations in Pyrrocoma at infraspecific ranks. To date, his new combinations remain unpublished and therefore invalid. Our attempts to locate Dr. Mayes have been unsuccessful. Since we plan to use five of his new combinations (*), we considered publishing them on his behalf. But such a publication requires his approval; hence, we reluctantly propose them here along with our own. Oonopsis foliosa E. Greenevar. monocephala(A. Nels.) Kartesz & Gandhi, comb. & stat. nov. BASIONYM: Oonopsis monocephala A. Nels., Bot. Gaz. (Crawfordsville) 31:399. 1901. TYPE: U.S.A. Colorado: 1900, Archibald 257. Pyrrocomacarthamoides Hook. var. cusickii (A. Gray) Kartesz & Gandhi, comb. nov. (*). BASIONYM: Haplopappus carthamoides (Hook.) A. Gray var. cusickii A. Gray, Syn. Fl. 1(2):126. 1884. TYPE: U.S.A. Oregon: Union Co., Jun 1877, Cusick 44$ (GH). Pyrrocoma hirta(A. Gray) E. Greenevar. sonchifolia (E. Greene) Kartesz & Gandhi, comb. & stat. nov. (*). BASIONYM: Pyrrocoma sonchifo- lia E. Greene, Lean. Bot. Observ. Crit. 2:18. 1909. TYPE: U.S.A. Washington: Yakima Co., 1882, Brandegee 111 (US). Pyrrocoma linearis (Keck) Kartesz & Gandhi; comb. & stat. nov. BA- SIONYM: Haplopappus uniflorus Hook. ssp. linearis Keck, Aliso 4:103. 1958. TYPE: U.S.A. Idaho: Owyhee Co., 17 Jun 1943, Christ 14081 (NY). Note: The grasslike leaves (linear and entire) of Pyrrocoma linearis warrant specific recognition from P. uniflora (Hook.) E. Greene, which has lanceolate or oblanceolate leaves with dentate margins. Pyrrocoma lucida (Keck) Kartesz & Gandhi, comb. nov. (*). BASIONYM: Haplopappus racemosus Nutt. ssp. lucidus Keck, Madrono 5:167. 1940. Haplopappus lucidus (Keck) Keck, Aliso 4:103. 1958. TYPE: U.S.A. California: Plumas Co., 9 Aug 1938, Keck 4897(CAS); Isotype: CAS. Note: Although Keckdid notjustify elevating Haplopappus racemosus ssp. lucidus to H. lucidus, we assert that the latter taxon with its glandular nature and acuminate-caudate phyl- lariesis distinct from theformer(usually eglandular; phyllaries obtuse-acute). Kartesz k Gandhi: Nomenclatural notes (or North America. VI 61 Pyrrocomaracemosa(Nutt.)Torr.k Grayvar. paniculata (Nutt.) Kartesz k Gandhi, comb. nov. BASIONYM: Homopappus paniculatus Nutt., Trans. Amer. Philos. Soc. II. 7:331. 1840. Haplopappus paniculatus (Nutt.) A. Gray var. paniculatus (automatically created by Haplopappus paniculatus var. virgatus A. Gray). TYPE: U.S.A. Oregon: Union Co., 1836, Nuttall s.n. (BM). Haplopappus paniculatus (Nutt.) A. Gray var. virgatus A. Gray, Proc. Amer. Acad. Arts 7:354. 1868. TYPE: U.S.A. California: Mono Co., Bolander 6168 (GH). Pyrrocomaracemosa(Nutt.)Torr.k Gray var. pinetorum (Keck)Kartesz k Gandhi, comb. & stat. nov. (*). BASIONYM: Haplopappus racemo- sus (Nutt.) Torr. k Gray ssp. pinetorum Keck, Madrono 5:166. 1940. TYPE: U.S.A. California: Siskiyou Co., 8 Aug 1938, Keck 4862(CAS). Pyrrocomauniflora (Hook.) E. Greenevar.gossypina (E. Greene) Kartesz k Gandhi, comb. & stat. nov. (*). BASIONYM: Pyrrocoma gossyp- ina E. Greene, Pittonia 3:23. 1898. TYPE: U.S.A. California: San Bernardino Co., Aug 1882, Parish 1558 (US). Stenotus acaulis (Nutt.) Nutt. var. glabratus (D.C. Eat.) Kartesz k Gandhi, comb. nov. BASIONYM: Haplopappus acaulis (Nutt.) A. Gray glabratus D.C. Eat. in S. Watson, Botany [fortieth parallel] 161. 1871. Chrysopsis caespitosus Nutt., J. Acad. Nat. Sci. Philadelphia 7:33. 1834. Stenotus caespitosus (Nutt.) Nutt., Trans. Amer. Philos. Soc. II. 7:334. 1841. TYPE: U.S.A. Missouri River (PI1; fiche !). Stenotus armerioides Nutt. var. gramineus (Welsh k F.G. Sm.) Kartesz k Gandhi, comb. nov. BASIONYM: Haplopappus armerioides (Nutt.) A. Gray var. gramineus Welsh k F.G. Sm., Great Basin Naturalist 43:371. 1982. TYPE: U.S.A. Utah: Uintah Co., 29 May 1982, Thome & Neely 1836 (BRY); Isotypes: NY, CAS. Sohdago simplex var. randii Nesom (1989b) reported that Sohdago glutmosa Nutt. (Trans. Amer. Phi- los. Soc. II. 7:328. 1841) and S. simplex Kunth (in H.B.K., Nov. Gen. et Sp. 4:163. 1820) are conspecific. In S. simplex, Ringius (in Ringius k Semple 1991) made seven new combinations including S. simplex ssp. randii (Porter) Ringius. Since we recognize the preceding taxon at varietal rank, a new com- bination is proposed below. Solidago simplex Kunth var. randii (Porter) Kartesz k Gandhi, ssp. et var. comb. nov. BASIONYM: Sohdago virgaurea L. var. randii Porter, PHYTOLOGIA 62 volume 71(l):58-65 July 1991 Bull. Torrey Bot. Club 20:208. 1893. Solidago glutinosaNutt. ssp. randu (Porter) Cronq., Rhodora 49:77. 1947. Solidago glutinosa Nutt. var. randii (Porter) Cronq., Rhodora 49:77. 1947. Solidago glutinosa Nutt. ssp. randii (Porter) Ringius, Phytologia 70:397. 1991. LECTOTYPE (vide Ringius, Taxon 36:156. 1987): U.S.A. Maine: Mt. Desert Island, 30 Aug 1892, Rand s.n. (NY). PORTULACACEAE We accept Hershkovitz' (1990) segregation of the genus Cistanthe Spach from the genus CalandriniaKunth and his inclusion ofthe genus Calyptridium Nutt. in Cistanthe. Four additional new combinations are required and made below. Cistanthe parryi (A. Gray) Hershkovitzvar. arizonica (J.T. Howell) Kartesz & Gandhi, comb. nov. BASIONYM: Calyptridium parryi A. Gray var. arizonicum J.T. Howell, Leafl. West. Bot. 4:215. 1945. TYPE: U.S.A. Arizona: Pima Co., Hills above Rosemont, 13 Mar-23 Apr 1903, Griffiths (US). Cistanthe parryi (A. Gray) Hershkovitz var. hesseae (Thomas) Kartesz &; Gandhi, comb. nov. BASIONYM: Calyptridium parryi A. Gray var. hesseae Thomas, Leafl. West. Bot. 8:10. 1956. TYPE: U.S.A. Califor- nia: Santa Cruz Co., Ben Lomond Mountains, 2 Jul 1954, Hesse 1317 (CAS). Cistanthe parryi (A. Gray) Hershkovitz var. nevadensis (J.T. Howell) Kartesz & Gandhi, comb. nov. BASIONYM: Calyptridium parryi A. Gray var. nevadense J.T. Howell, Lean. West. Bot. 4:216. 1945. TYPE: U.S.A. Nevada: Gold Mt., 23 Jun 1930, Keck 559 (CAS). Cistanthe umbellata(Torr.) var.caudicifera (A. Gray) Kartesz& Gandhi, comb. nov. BASIONYM: Spraguea umbellata Torr. var. caudicifera A. Gray (in Patterson, Numb. Checkl. N. Amer. PL 14. 1892, nom. nud.). Syn. Fl. 1:278. 1897. Calyptridium umbellatum (Torr.) E. Greene var. caudiciferum (A. Gray) Jepson, Fl. Calif. 1:465. 1914. Note: Typification of Gray's trinomial is under our inves- tigation. Kartesz <fc Gandhi: Nomenclatural notes for North America VI 63 ACKNOWLEDGMENT We are grateful to: Dr. Larry E. Brown (SBSC) and to Dr. Jimmy Massey (NCU) for their valuable comments on the manuscript; and Dr. Guy Nesom (TEX) and Mr. Bryan Dutton (GH) for providing literature used in this study. REFERENCES Brown, G.K. &: W.D. Clark. 1982. Taxonomy of Haplopappus sect. Gymno- coma (Compositae). Syst. Bot. 7:199-213. Clark, W.D. 1979. The taxonomy of Hazardia (Compositae: Astereae). Madrono 26:105-127. Correll, D.S. & M.C. Johnston. 1970. Manual of the Vascular Plants of Texas. Texas Research Foundation, Renner, Texas. Cronquist, A. &; D.I? Keck. 1957. A reconstitution of the genus Machaer- anthera. Brittonia 9:231-239. Goodrich, S., E.D. McArthur, & A.H. Winward. 1985. A new combination and a new variety in Artemisia tridentata. Great Basin Naturalist 45:99- 104. Greuter, W., H.M. Burdet, W.G. Chaloner, V. Demoulin, R. Grolle, D.L. Hawksworth, D.H. Nicolson, P.C. Silva, F.A. Stafleu, E.G.Voss,&J. Mc- Neill. 1988. International Code ofBotanical Nomenclature. 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