MemoirsofMuseumVictoria57(2): 225-236 (1999) NEW SPECIES OF THE WATER MITE GENUS ARRENURUS FROM EASTERN AUSTRALIA (ACARI: HYDRACFTNIDIA: ARRENURIDAE) Harry Smit DM Emmastraat43-a, 1814 Alkmaar, The Netherlands ([email protected]) Abstract Smit,H., 1999. NewspeciesofthewatermitegenusArrenurusfromeastern Australia(Acari: Hydrachnidia: Arrenuridae). Memoirs ofMuseum Victoria 57: 225-236. Five new species of the water mite genus Arrenurus from Queensland, Victoria and Tasmania are described, viz. A. acutipetiolatus sp. nov., A. maria sp. nov., A. hyhrichts sp. nov., A. queenslandicus sp. nov andA. perplexus sp. nov. The nameArrenurus mantonensis Smit ispreoccupied, andtherefore thenewnameArrenurusbifurcatusnom. nov. isproposed. ArrenurusmadarasziDadayisrecordedforthefirsttimefrom Australia. Somemeasurements, additional characters andnew recordsare given foralready known species. Introduction Natural History. For the description of the Ofthe cosmopolitan water mite genus Arrenurus glandularia, Jin and Wiles (1996) and Wiles 35 species and subspecies are known from (1997)are followed. All measurementsare infim, measurements of leg and palp segments are of Australia (Smit, 1997), a relatively low number the dorsal margins. Measurementsofparatypes in compared to the 150 species from Europe (Smit, v1i9d96e)d.bAy kSemyitof(1al9l97A)u.strInalitahinssppeacpieers fwiaves npreow- tbhreackdeetssc.riSpctailoenlionefsnareew20s0pe|ciimesforarmeosgtivfiegnureisn and 50 |im for figures ofthe palp. species from Victoria, Tasmania and Queensland are described, and one new name is given to a Arrenurus (Arrenurus) acutipetiolatus sp. nov. previously described species. Additionally, Figures 1-6 one species new to the fauna of Australia is reported and some additional characters and Material examined. Holotype. Male, Victoria, swamp measurements are given for already known atjunction ofVictoria Valley Road and Bundol Road, species. SWofGrampiansNational Park,30 Sep 1997 (NMV). All material has been collected by the author. Paratypes: Victoria: 8 males, 22 females, same data Victorian and Tasmanian holotypes and para- asholotype (NMV, TM, ZMAN). types have been deposited in Museum Victoria, Tasmania: 14 females. Reservoir of Darlington, Maria Island National Park, Tasmania, 18 Oct 1997 Melbourne (NMV), and in the Tasmanian (ZMAN). Museum and Art Gallery, Hobart (TM) respect- Othermaterial.New South Wales: 1 female,MtVic- ively. Holotypes and paratypes from Queensland toria, 24 Oct 1936, leg. F. Under (NHRS, slide 3414, have been deposited in the Queensland Museum, "A. fissipetiolatus"). Brisbane (QM). Otherparatypes and all non-type Diagnosis. Petiole without ligulate process, nar- material have been deposited in the Zoological rowed posteriorly, with pointed extension. Museum of the University of Amsterdam Female with L4 and V2 on small humps. (ZMAN). The following abbreviations have been used Description. Male: Body 1499 (1464-1584) in (see figs 1, 2 and 6): Al and A2, pre- and post length and 1222 (1128-1248) in width. Body antennal glandularia; C2^I, coxoglandularia2-4; brownish. Anterior body margin concave. Body Dl-4, dorsoglandularia 1-4; Ll-4, lateroglandu- with well developed pygal lobes. Dorsal shield laria 1—4; V2, ventroglandularia 2; PI-PV, palp 601 in width, dorsal furrow incomplete. Dl on segments 1-5; IV-leg-4-6, fourth-sixth segments small humps, D3 on large humps. Setae associ- of fourth leg; NHRS, Swedish Museum of ated with D4 on long tubercles. Genital plates 225 226 H. SMIT Figures 1-4.Arrenurusacutipetiolatus. 1, holotypemale,dorsal view. 2,ventralview. 3, lateralview. 4,palp. NEW SPECIES OF THE WATER MITE GENUS ARRENURUSFROM EASTERN AUSTRALIA 227 Figures 5, 6. Arremtrus uculipetiolatus. 5, paratype female, ventral view. 6, dorsal view. straight, medially widened, extending to lateral Gonopore 1 16 long. Gonopore without chitinized body margin. Petiole posteriorly narrowed, with a patches. Genital plates wide, slightly bowed. V2 pointed extension; ligulate process absent. Hya- on small humps. Lengths ofPI-PV: 48, 132, 118, line membrane well developed, with more or less 144, 89. Palp as in male, antagonistic bristle of pointed lateral angles, posteriorly concave. Setae PIV 103 long. Lengths of l-leg-4-6: 281, 233, associated with petiole extending beyond pos- 242. Lengths of IV-leg-4-6: 349, 301, 267. terior margin of petiole. Lengths of PI-PV: 60, Second, third and fourth legs with numerous 130, 108, 149, 88; PII with 4 setaeonmedial side, swimming setae. ofwhich 2 in anteroventral corner. Antagonistic Etymolog}'. The name refers to the pointed bristle of PIV 106 long. Lengths of I-leg-4-6: petiole. 301, 262, 252. Lengths of IV-leg-4-6: 407, 204, 262; IV-leg-4 witha short spur. Second, third and Remarks. The new species is close toA. fissipeti- fourth legs with numerous swimming setae. olalus Lundblad. Both have an almost similar Female: Body 1656 (1608-1992) in length and palp (longantagonistic bristleofPIV, 2-3 setae in 1416 (1344-1764) in width. Anterior margin of anteroventral corner ofPII). Males are easily dis- body concave. Body truncated posteriorly, pos- tinguished by the shape ofthe petiole, but distin- terolateral corners present. Dorsal shield 1056 guishing the females is more difficult. Both have (1032-1344) in length and 563 (446-582) in a rather slender dorsal shield, which character width. Dorsal shield slender, usually widest in they share with A. balladoniensis Halik and A. anterior part, slightly tapering posteriorly, poste- ensiferSmit. The last species isthe smallest, 1416 rior margin straight. Body shape and shape of in length. A. balladoniensis is the largest, measur- dorsal shield variable, less truncated specimens ing 1848-2232 in length (Smit, 1997). A. bal- can be found, as well as specimens in which the ladoniensis can be told apart fromfissipetiolatus dorsal shield is contracted in middle or with par- and acutipetiolatus by the shape of the genital allel margins. L4 on small humps. Distance of plate, which is much narrowerthanthatofthe last fourth coxal plates larger than width of 1 geni- two species.A. acutipetiolatus differs fromA.fis- tal valve. Medial margin of fourth coxal plates sipetiolatus by having L4 on tubercles and a pos- larger than medial length of third coxal plates. teriorly more truncated body. Another useful 228 H. SMIT character by which to distinguish the two species humps, body truncated, ratio dorsal shield 1.89), is the shape of the dorsal shield, which is more and should therefore be assigned to that species. slender in acutipetiolatus. The ratio length/width ofthe dorsal shield is 1.73-2.04 inacutipetiolatus (usually >1.80) and 1.50-1.67 mfissipetiolatus. Arrenurus (Arrenurus) hybridus sp. nov. Lundblad (1947) described the female of A. Figures 7-10 fissipetiolatus, but he had some doubts if the — assignment to this species was correct, because Arrenurus sp. Harvey, 1998: 106, fig. 32C. the male and the female came from different loca- Material examined. Holotype. Male, Victoria, pond, tions. The female described by Lundblad matches Hospice Plain, Mt Buffalo National Park, 10 Oct 1997 the description of A. acutipetiolatus (L4 on (NMV). Figures 7-10. Arrenurus hybridus, holotype male, 12, dorsal view, 13, ventral view. 14, palp. 15, paratype female, ventral view. Figure 1 1. Arrenurusfissipetiolatus Lundblad, holotype male, detail ofpetiole. 1 1 NEW SPECIES OF THE WATER MITE GENUSARRENURUS FROM EASTERN AUSTRALIA 229 saPmaeradtaytpaeass.hVoilcototryipae:;11mmaallee((nZoMtAsNcl)e,rot1imzeadl,eZ(MNMAVN),). fhoorem,onthyemspoefciAe.smdaenstcorniebnesdisbyGemoergeis aandjutnhiuosr 1 female(NMV), smallponds, Kowan Plain, ± 1400 m requires a new name for which I proposeArrenu- abovesealevel,MtBuffaloNational Park, 10Oct 1997. rus bifurcatus nom. nov. The new name refers to Diagnosis. Body with distinct cauda. Petiole pre- the bifurcated setae ofthe petiole. sent, spatulate. Hyaline membrane and ligulate process absent. Arrenurus (Arrenurus) fissipetiolatus Lundblad Description. Male: Body 951 (912-951) longand Figure 1 543 (504-553) wide. Anterior body margin Arrenurus (Arremmis) fissipetiolatus Lundblad, rounded. Body posteriorly gradually tapering into 1947: 73, tigs 46A—D. Cook. 1986, figs a distinct cauda. Cauda with a shallow concavity. 1632-1637. Dorsal furrow not closed posteriorly. Posterior Material examined. Holotype. Male, Victoria, Mary- margin almost straight, with small indentations. borough. 24 Sept 1926. leg. E.J. Semmens (NHRS, First coxal plates extending beyond anteriorbody slide 3413). margin. Second and third coxal plates lying very Othermaterial. Victoria: 1 female, swampatjunction close. Gonopore 72 long. Genital field 197 long, of Victoria Valley Road and Bundol Road, SW of not extending to lateral body margin, slightly Grampians National Park, 30 Sep 1997. undulating. Petiole spatulate, hyaline membrane Tasmania: 1 male, 14 females, Blackmans Lagoon, and ligulateprocessabsent. LengthsofPl-PV: 38, Waterhouse Protected Area, 21 Oct 1997: 4 females. 82, 65, 91, 31; PI1 with 7 (8?) setae on medial LittleWaterhouseLake,WaterhouseProtectedArea,2 Oct 1997. side. Lengthsofl-leg-4-6: 136, 136, 146. Lengths of lV-leg-4-6: 194, 184. 155. lV-leg-4 without Description. Male: Body 1488 (1337-1483) spur, but lV-leg-5 with dorsodistal extension. in length and 1272 (1064-1138) in width (in Second, third and fourth legs with numerous brackets the measurements of Lundblad, 1947 swimming setae. and Cook, 1986). Female: Body 1106 long and 825 wide, egg- Female: Body 1680-1944 (Cook, 1986: 1581) shaped, without posterolateral corners. Anterior in length and 1392-1656 (Cook, 1986: 1292) in body margin rounded. Dorsal shield 970 long and width. Dorsal shield 1104-1320 in length and 679 wide, dorsal furrow closed. Medial lengthsof 281-364 in width; usually widest in middle. LI third and fourth coxal plates of equal length, shifted dorsally towards dorsal shield and there- fourth coxal plates almost without posteromedial fore visible in dorsal view. corner. Gonopore large, 165 in length; gonopore Remarks. Contrary to Lundblad's description, the with indistinct sclerotized patches. Genital plates male does not have a long spur. The holotype bowed, laterally somewhat rectangular. Lengths (Figure 16) andmy own specimen have a notch in ofPI-PV: 46, 98, 82, 108, 48; PII with 8 setae on the posterior margin of the petiole. However, medial side. Second, third and fourth legs with Cook (1986) could not observe this in all his numerous swimming setae. specimens. The genital plate of most of my Etymology. The name refers to the somewhat female specimens fit well with the description of intermediate position ofthe new species between Cook (1986). However, occasionally some speci- the subgenera Arrenurus and Megaluracarus. mens have a narrower genital plate, while others Remarks. No other Indo-Australian Arrenurus have a genital plate with a slightly undulating posterior margin. The species has been reported species has a distinct cauda with a spatulate peti- from Victoria and Tasmania. However, the num- ole. The female is characterized by the combina- ber ofspecimens collected so far is very limited. tion ofthe absence of posterolateral corners, the Therefore, some additional characters and absence of posteromedial corners of the fourth measurementsare given. The female described by coxal plates and the medial margins of the third Lundblad (1947) belonged to anotherspecies (see and fourth coxal plates being ofequal length. underA. acittipetiolatus). Arrenurus (Arrenurus) bifurcatus nov. nom. Arrenurus (Megaluracarus) niaria sp. nov. Remarks. When describingA. (Arrenurus)manto- Figures 12-16 nensis Smit, 1997, I was not aware of the exis- tence of A. (Megaluracarus) mantonensis Material examined. Holotype. Male, Tasmania, reser- George, 1903. This species was synonymized by voirofDarlington. Maria Island National Park, 18 Oct Viets (1956) with A. buccinator (Miiller). There- 1997 (TM). 230 H. SMIT Figures 12-16. Arrenurus maria. 7, holotype male, dorsal view, ventral view. 9, lateral 10, palp. paratype female, ventral view. Paratypes. Tasmania: 1 female, same data as holo- Description. Male: Body 1465 longand 708wide. type (TM); 1 female, Apsley River, at crossing with Anterior and posterior body part brownish, mid- Tasmanian Highway, 19 Oct 1997 (ZMAN). dlepartbluish, caudapurple; legsbluish. Anterior Diagnosis. Male with a long cauda, posterior bodymarginalmost straight, bodytruncatedante- margin of cauda with a small hyaline area and riorly. In the middle between the eyes an area 4 rounded hyaline extensions, 1 peg-like without body pores. Postocularia setae and setae rudimentary petiole. associated with dorsoglandularia 2 and 4 very NEW SPECIES OF THE WATER MITE GENUS .4RRENURUS FROM EASTERN AUSTRALIA 23 1 long. Cauda much longer than wide, width of Material examined. Victoria: 10 males, 21 females, cauda 359. D4 on small humps. Posterolateral swamp atjunction ofVictoria Valley Road and Bundol corners ofcauda rounded. Cauda posteriorly with Road, southwest ofGrampians National Park, 30 Sep aextdeinsstiinocnts.hPyeatliionlee raurdeiameanntdar4y,rpoeugn-dliekde,. hGeynailtianle P1a9r9k7,; 130 Ofcetmal1e9s9,7;LIakmealeC,atuannin,amMeld cBruefefkal4o.5Nkatmioenaaslt of Shipwreck Creek, Croajingolong National Park, 23 field 252 in width. Genital plates short and Oct 1997. rounded. Lengths ofPI-PV: 24, 94, 62, 1 16. 53. Tasmania: 1 male,2females,oldriverbranchofCoal P1I with a large patch ofsetae lying on a bulge. River, north of Richmond. 17 Oct 1997; 1 female, Lengths of I-leg-4-6: 206, 243, 180. Lengths of swamp 12 km south ofGladstone, along road B82, 20 IV-leg-4-6: 272, 184, 243; IV-leg-4 with a short Oct 1997; 2 males, 10 females. Big Waterhouse Lake, spur. Second, third and fourth legs with numerous Waterhouse Protected Area, 21 Oct 1997; 2 females. swimming setae. LittleWaterhouseLake, WaterhouseProtectedArea,21 Female: Body 1416 (1320) long and 1077 Oct 1997. (1009) wide. Dorsal shield 897 long and 902 Description. Male: Body 689-786 in length and wide; dorsal furrow complete. Body brownish, 543-640 in width. Petiole occasionally reaching posterior and anterior body parts purple; legs posterior body margin. Caudal lobes in some bluish. Between eyes an area without body pores. specimens rounded. Body egg-shaped, without posterolateral or with Female: Body 776-936 in length and 650-786 indistinct posterolateral corners. Medial margin in width. Dorsal shield complete. ofthird coxal plates larger than medial margin of Remarks. The species was previously reported fourth coxal plates. Medial distance of fourth from Victoria, South Australia and Queensland. coxalplates slightly smallerthan width of1 gono- Thespecimens fromQueensland reportedby Smit pore valve. Gonopore 194 long. Genital plates (1992) do not belong to this species but to a new short and wide, sloping posteriorly, extending species described below. The record from China beyond posterior margin ofgonopore. Lengths of (Uchida and Imamura, 1951) needs confirmation PI-V: 36,94,67, 122, 50; palpasinmale. Lengths ofI-leg-4-6: 165, 204, 146. Lengths of IV-leg- as only females have been collected. Females of 4-6: 233, 272, 276. Second, third and fourth legs the subgenus Micruracarus with two pairs of with numerous swimming setae. rounded chitinized patches on the gonopore are difficultto identify. Moreover, the genital field of Etymology: Named after the island where thetype the specimens illustrated by Uchida and Imamura specimen was collected. Noun in apposition. ( 1951) is laterally narrowed,acharacternotfound Remarks. The male ofthe new species is close to in the Australian specimens. anumberofAustralian species with a longcauda. Arrenurus (Micruracarus) queenslandicus i.e.A. otodus Cook.A. gilvus Smit andA. vander- palae Smit. The new species differs from sp. nov. A. gilvus and.-I. vanderpalae by its large size and Figures 17-20 — the presence of only one peg-like rudimentary Arrenurus (Micruracarus) forpicatoides. Smit, petiole (two ingilvus and vanderpalae), and from 1992: 109. A. otodus by the lack of the pointed postero- Materialexamined. Flolotype. Male, Queensland, pond lateral corners ofthe cauda. Moreover, A. maria north ofNormanton, 14 Aug 1989 (QM). is larger than A. otodus, has a larger hump on Paratypes. Queensland: 3 males, 5 females which D4 are located, and its rounded hyaline (QM), 3 males, 5 females (NMV), 3 males, 5 extensions are not found inA. otodus. The female females (ZMAN), all same data as holotype; 3 can be distinguished from all other species by males, 1 female (ZMAN), pond near Crocodile having the medial margin of the third coxal Road, Bowling Green Bay National Park, 31 Jul plates larger than the medial margin ofthe fourth 1989; 3 males, 1 female (ZMAN), pond Townsville Common National Park, Aug 1989; coxal plates. 1 male, Freshwater Lagoon, Horseshoe Bay, 1 Magnetic Island, 3 Aug 1989. Arrenurus (Micruracarus)forpicatoides Othermaterial. Queensland: 2 females,pondnorth of Longreach, 15 Aug 1989. Lundblad Arrenuru—s (Micruracarus)forpicatoides Lundbla—d, Diagnosis. Caudaofmale indistinctly setofffrom 1941: 160. Lundblad, 1947: 75, figs 47A—-D. anterior part of body. Petiole tong-shaped, Uchida and Imamura, 1951: 353, figs 18a-d. Smit, hyaline, fused with body by small hyaline area. 1992: 109. Dorsal shield offemale incomplete. ' i.' SMI Figures 17 20. Arrenurus queenslandicus. 17, holotype male, dorsa] view. IS, ventral view. 19, palp. 20, paratype female, ventral mow. Description. Male. Body 70S (660 795) in 4 6: 110, 113, 132. Lengths of IV-leg-4 6: 144, length and 61 1 (553 679) in width. Body yellow- 134, 142; IV-leg-4 without a spur. Second, third ish brown to greenish. Anterior margin of body and fourth legs with numerous swimming setae. straighl to slightly concave. Dorsal shield com- female: Body egg-shaped, 951 (936 1048) in plete, 423 (359 456) in length and 417 (373-441) length and 786(771 844) in width. Anteriorbody in width. Genital plates long anil narrow, extend- margin straight or slightly concave. Posterolateral ing onto dorsum. Cauda indistinctly set off from corners of body almost absent. However, occa- anterior pari ofbody. Cauda with median cleft. sionally posterolateral corners more pronounced, Petiole hyaline, long-shaped, fused wilh body by and in these specimens body truncated pos- a small hyaline area. Lengths of PI I'V: 31, 61, teriorly. Dorsal shield incomplete, 524 (495 572) 40, 84, 50; I'll with 4 setae on medial side, of in width. Medial distance of fourth coxal plates which 2 elose to ventral margin. Lengths ofI-lea- longer than width of I genital valve. Medial NKW SPECIES OF THE WATER MITE GENUS ARRENURUS FROM EASTERN AUSTRALIA 233 margin of fourth coxal plates longer than medial coxal plates slightly larger than I genital valve. margin ofthird coxa] plates. Gonopore 98 long. Medial margin of fourth coxal plates larger than Genital valves with large ehitinons patches, ante- medial margin ofthird coxal plates. Gonopore 97 rior and posterior patches connected by ehitinons long, each valve with 2 rounded ehitinons strip. Genital plates straight to slightly bowed, patches. Genital plates straight, slightly sloping tapering laterally. Lengths of PI-PV; 36, 70, 48, posteriorly and laterallv somewhat enlarged. 89,43; palpas in male. Lengths ofI-leg-4 6: 130, Lengths ofPI-PV: 31, 55, 41, 86, 43. Palp as in 134, 125. Lengths oflV-leg-4 6: 155, 155, 136. male, but medial side ofPII with 3 setae. Lengths Second, third and fourth legs with numerous of l-leg-4 6: 121, 125, 120. Lengths of IV-lcg- swimming setae. 4 6: 170, 156, 130. Second, third and fourth legs with numerous swimming setae. Etymology. The name refers to the state of Queensland in which the species has been found. Etymology, The name refers to (he complicated structure ofthe petiole. Remarks. erroneously identified the species as I A. forpicatoides (Smit, 1992). However, in the Remarks, flic new species is close lo A.jorpica- last species the eauda is distinctly set offfrom the toides, but differs in size, body colour and shape body, the body is reddish brown and the tong- of the cauda. The hyaline petiole of the two shaped part ofthe petiole is narrower. A. anbang- species arc nearly identical, although the tong- hang Smit has a different shaped petiole which is shaped part ofthe petiole is narrower in the new not fused with the Cauda. The female ofthe new species. In A. qiieenslaudicus the cauda is not species closely resembles A. forpicatoides but is set off from the body, but the petioles of the larger, has a different colour and an incomplete two species are very similar. The female ol dorsal shield. .1. pcrplexus can be distinguished from A. forpi- catoides by the body colour, shape of the body and the broader genital plates, and from Arrenurus (Micruracuius) pcrplexus sp. nov. A. queenslandicus by the complete dorsal shield. Figures 21 24 The species of Micruracarus with a hyaline nMeaatrerBiuaclkleaxnadmiRnievde,r,laltoclrootyspsei.ngMwailteh.BuVcikctloarmila,Vaplolnedy pAertrieoilneinifso.rmInamcyommaptleerxialgr(ofurpomwiltlhasilnietshSewgaemnpu,s Road (west ofBright), 11 Oct 1997(NMV). Alherton Tablelands, Queensland) another Paratype. Victoria: 1 female, same data as holotype species might be present, of which the only male (NMV). is somewhat intermediate between .1. queens- bDpeiotadigyon,loeslaitshe.yraallCianmuea,dragiloonfngom-fasclhaeaupddeiads.tcionBncotvledyxy,sebtoofodfyffebfmlrauoleme, lmsmpaaenlscdeihieacsapu.nesdUaonfnftdohterhtAe.umnaepalacteuresdplalyeo,axfaunlsdtl.htethThrheteewebrooefdeamyafraoelbreeedstimwfefefnreetornimeonttnchheeiidsss tapering posteriorly. location arc different, and I refrain from describ- Description. Male: Body 635 long and 519 wide. ing it as a new species until more material is Dorsal shield complete, 320 long and 320 wide. available. Body blue. Cauda distinctly set offfrom anterior part of body, lateral margins of cauda convex, caudal lobes rounded. Cauda posteriorly with a Arrenurus (Micruracarus) iiiuiluraszi Daday median cleft. Dl on small humps. Petiole hyaline, Arrenurus Madardszi Daday, 1898: 99, figs 49a i, tong-shaped, fused with body by a small hyaline Arrenurus (Micruracarus) forpicatoides. Smit, area. Genital plates narrow and long, extending 1992: 109 (part). onto dorsum. Lengths of PI-PV: 28, 50, 38. 72, 4l1c;g-4PI-I6:wi9t8h, 944,set1a1e3.onLemnegdtihaslosfidIeV.-lLcegn-g4th6s: o1f32I,- TMIaoWtwenMrsZivaiMllAlcNexC)ao.mmimnoedn NQauteieonnasllaPnadr:k,QIuemeanlsal,andp,onIdAuigll 125, 108; IV-leg-4 without spur. Second, third and fourth legs with numerous swimming setae. Remarks. Initially, the specimen has been identi- Female: Body 757 long and 630 wide. Dorsal fied erroneously as A. forpicatoides to which it is shield complete, 582 long and 432 wide; dorsal closely related. A. madaraszi is a very widespread shield somewhattapering posteriorly. Bodytaper- species, known from Japan, China, Burma, India, ing posteriorly, slightly truncated. Anterior mar- Sri Lanka throughout Indonesia. This is the first gin of body straight. Medial distance of fourth record for Australia. 234 II. SIV1IT tFeimgaulree,sv2e1n-t2r4a.lAvirerwe.numsperplexus. 21, holotypemale, dorsal view. 22, ventral view. 23 palp 24' pFarat}vyoe Figure 25. Arrenumshaswelli Cook, ventral view female. Figure 26. Arrenums novaehollandiae Lundblad, ventral view male.