Vestnik zoologii, 45(3): 195—207, 2011 Ôàóíà è ñèñòåìàòèêà UDC 395.766.4(47) NEW SPECIES OF THE GENUS LASIODERMA (COLEOPTERA, ANOBIIDAE) FROM CRIMEA AND THE CAUCASUS I. N. Toskina 107241, post box 25, Moscow, Russia E-mail: [email protected] Received 12 October 2010 Accepted 30 March 2011 New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus. Toski- naI.N.– Fivespecies of the genus LasiodermaStephens, 1835 from Crimea(L. dolinisp. n., L. tau- ricumsp. n.), North Caucasus (L. prolixumsp. n., L. sparsumsp. n.), and Armenia (L. morulumsp. n.) are described. A key to 17 species of the genus Lasioderma occurring in these regions is compiled. Key words: Coleoptera, Anobiidae, Xyletininae, Lasioderma, Crimea, Caucasus, new species, key. Íîâûå âèäû ðîäà Lasioderma (Coleoptera, Anobiidae) èç Êðûìà è Êàâêàçà. ÒîñêèíàÈ. Í. – Îïèñàíû 5 âèäîâ ðîäà Lasioderma Stephens, 1835 èç Êðûìà (L. dolini sp. n., L. tauricum sp. n.), Ñåâåðíîãî Êàâêàçà (L. prolixum sp. n., L. sparsum sp. n.) è Àðìåíèè (L. morulum sp. n.). Ñîñòàâëåíà îïðåäåëèòåëüíàÿ òàáëèöà 17 âèäîâ ðîäà Lasioderma äëÿ íàçâàííîãî ðåãèîíà. Êëþ÷åâûå ñëîâà: Coleoptera, Anobiidae, Xyletininae, Lasioderma, Êðûì, Êàâêàç, íîâûå âèäû, òàáëèöà äëÿ îïðåäåëåíèÿ âèäîâ. Introduction Anobiid beetles of the genus Lasiodermawere studied in Crimeaand the Caucasus by A. Reitter (1884, 1913), who described L. punctulatum Reitter, 1884 and L. babadjanidesi Reitter, 1913. Approximately at the same time,Schilsky (1899) described L. kiesenwetteriSchilsky, 1899, and Pic (1904) gave very short descrip- tion of L. redtenbacheri var. caucasicum Pic, 1904. Later, F. Español (1970, 1992) figured and commented L. haemorrhoidale (Illiger, 1807) and S. M. Iablokoff-Khnzorian (1976) illustrated and gave new records of L. aterrimumRoubal, 1916. V. D. Logvinovskij (1977) described L. kryzhanovskiiLogvinovskij, 1977 and pro- vided a key to Lasiodermaspecies of the USSR in his “Fauna of USSR” Anobiidae monograph (Logvinovskij, 1985). Recently, I. N. Toskina described L. mazokhini Toskina, 1999, L. mikheechevi Toskina, 1999 and L.oculeum Toskina, 1999 and provided the most comprehensive key to the species of this genus occurringin the USSR and adjacent countries. In this work, 5 additional new species from Crimea and the Caucasus of the genus Lasioderma are described, and a key to species occurring in Crimea and Caucasus regions is pro- vided. Material and methods Length of thepronotum was measured in profile in a position with theapical and basal margins aligned at the same level. Measurements of thepronotum from above give higher error. The elytral length was mea- sured from the basal margin of scutellum along elytral suture. The elytral width was measured somewhat pos- teriorly of the shoulders. Aedeagus is figured from its dorsal side, and the pseudopositors from their ventral side. In cases when hind tarsi are lost the ratio of the segments in the middle or fore tarsi is used, and then the ratio of tibia and tarsus lengths is not used. Measurements with asterisk (*) are those of the holotype. Material is deposited in the collections of Hungarian Museum of Natural History (Budapest) (HMNH), the Zoological Museum of Moscow University (ZMUM), and Zoological Institute of the Russian Academy of Sciences, St.-Petersburg (ZISP). 196 Descriptions of new species Lasioderma dolini sp. n. (fig. 1, 1—10) Material.Holotype }: USSR, [Crimea], Kizil-Dzhar, 8.05.1975 (Dolin) (HMNH). 2 paratypes with the same label data (HMNH, ZMUM). Description. Habitus as on fig. 1,1. Dark yellow beetles; antenna light yellow. Pubescence light yellow, homogenous, fine, appressed on elytral disc, raised on sides. Fig. 1. Lasioderma dolini sp. n., holotype }: 1 – general outline of body; 2 – antenna; 3 – maxillary pal- pus; 4 – anterior part of body, lateral view; 5 – pronotum, dorsal view (with pubescence pattern); 6 – punctation on pronotal disc; 7– structure of elytral surface; 8– the middle of metasternum and 1st visible abdominal sternite; 9– fore tarsus (part); 10– pseudopositor: coxite top with style. Scale bar: 1.0 mm(1); 0.5mm (4, 5, 8); 0.2 mm (2, 9); 0.1 mm (3, 6, 7); 0.05 mm (10). Ðèñ. 1. Lasioderma dolini sp. n., ãîëîòèï }: 1 – îáùèé êîíòóð òåëà; 2 – óñèê; 3 – ïîñëåäíèé ÷ëåíèê ÷åëþñòíûõ ùóïèêîâ; 4 – ïåðåäíÿÿ ÷àñòü òåëà, âèä ñáîêó; 5 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ ðèñóí- êîì îïóøåíèÿ); 6– ïóíêòèðîâêà íà äèñêå ïåðåäíåñïèíêè; 7– ñòðóêòóðà ïîâåðõíîñòè íàäêðûëüåâ; 8 – ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî áðþøíîãî ñòåðíèòà; 9 – ïåðåäíÿÿ ëàïêà (÷àñòü); 10 – ëîæíûé ÿéöåêëàä: âåðøèíà êîêñèòà ñî ñòèëåì. Ìàñøòàáíûå ëèíåéêè: 1,0 ìì (1); 0,5 ìì (4, 5, 8); 0,2ìì (2, 9); 0,1 ìì (3, 6, 7); 0,05 ìì (10). 197 Head. Frons slightly convex. Eyes round, convex, separated by 1.8 eye diameters. Antenna: 3rd segment almost as long as the 2nd one; 4th segment with oblique and concave upper margin; 5—10th segments with concave upper margin and convex late- ral one. 11th segment 3.3 times as long as wide and 1.6 times as long as the 10th seg- ment. All segments oblong (fig. 1, 2). Last segment of maxillary palpus is spindle- shaped, 3.3 times as long as thick, with oblique truncate top and with carina on it (fig.1, 3). Pronotum 1.5—1.6* times as wide as long, with base arcuate, anterior angles rectangular, posterior ones not developed (fig. 1, 4). Pronotum slightly convex, lateral margins narrowly flattened, sides very slightly bulging. Pubescence not parted to two sides (fig. 1, 5). Surface with dual punctation, small punctures very fine (fig. 1, 6). Scutellum triangular. Elytra 1.8*—2.0 times as long as wide and 2.8*—3.0 times as long as pronotum. Surface very gently wrinkled (fig. 1,7). Metasternum nearly as long as 1st visible abdominal sternite. Anterior rims a little approximated and evenly rounded medially. Medial portion significantly stronger convex in distal third of metasternum (fig. 1, 8). Surface finely wrinkled with puncture traces. Legs. Fore tibiae not widening to their apices. 1st segment of fore tarsus 1,7 times as long as 2nd; 2nd segment 1.7 times as long as 3rd (fig. 1, 9). Other tarsi lost or strongly damaged in type specimens. Pseudopositor. Style looking as elongated cone with long chaeta on its top; coxite top with rather long chaetae (fig. 1, 10). Length 4.5*—5.0 mm, width 1.8—1.9* mm. Differential diagnosis. L. dolini differs from other species of the genus Lasioderma with having yellow-rufous or reddish-brown colour and with pubescens not parted to two sides on pronotum occurring in this region (L. haemorrhoidale, L.kryzhanovskii, L. mazokhini, L. oculeum, L. prolixum sp. n., L. punctulatum and L.serricorne (Fabricius, 1792) as follows. L. dolini differs from L. serricorne and L. pro- lixum by the non-dilated fore tibiae (fore tibiae dilated to their apices in the compared species); it differs from L. haemorrhoidale, L. mazokhini, and L. oculeum by evenly rounded anterior rims of metasternum (rims approximated in the middle and straigh- tened or curved forwards in the compared species); the new species differs from L.kryzhanovskii by homogenous pubescence on elytra (in the compared species, pubescence dual on elytra) (see Logvinovskij, 1985: 134). At last, L. dolini differs from L.punctulatum in fine punctation (in L. punctulatum, punctation is already visible with magnification x10). Etymology. The new species is named in memory of Ukrainian entomologist Vladimir Gdalich Dolin (1932—2004), who collected the type series. Lasioderma morulum sp. n. (fig. 2, 1—10) Material. Holotype {: “USSR, Armenia, Gokh-Guekhard, 1500 m, 30 km E from Yerevan, 29.05.1980, N 129” (J. Papp) (HMNH). Description. Habitus as on fig. 2, 1. Black beetle except dark yellow elytral apices; continued as narrow stripe along lateral margin of elytron to metasternum. Pubescence very fine, dark, homogenous, appressed on elytra. Head. Frons almost flat. Eyes round, moderately convex, separated by 2.7 eye diameters. Antenna: 3rd segment 1.5 times as long as the 2nd one; 4—5th segments with oblique and concave upper margin and convex lateral one; 6—9th segments with straight upper and lateral margins (10—11th segments are lost). All segments except 5th oblong (fig. 2, 2). 198 Fig. 2. Lasioderma morulumsp. n., holotype {: 1– general outline of body; 2– antenna; 3– anterior part of body, lateral view; 4 – pronotum, dorsal view (with pubescence pattern); 5 – punctation on pronotal disc; 6– punctation on elytral disc; 7– the middle of metasternum and 1st visible abdominal sternite; 8– hind tibia and tarsus; 9 – aedeagus; 10 – distal part of genital ring. Scale bar: 1.0 mm (1); 0.5 mm (3, 4); 0.2 mm (2, 7—10); 0.1 mm (5, 6). Ðèñ. 2. Lasioderma morulum sp. n., ãîëîòèï {: 1 – îáùèé êîíòóð òåëà; 2 – óñèê; 3 – ïåðåäíÿÿ ÷àñòü òåëà, âèä ñáîêó; 4 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ ðèñóíêîì îïóøåíèÿ); 5 – ïóíêòèðîâêà íà äèñêå ïåðåäíåñïèíêè; 6 – ïóíêòèðîâêà íà äèñêå íàäêðûëüåâ; 7 – ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî áðþøíîãî ñòåðíèòà; 8– ãîëåíü è ëàïêà çàäíåé íîãè; 9– ýäåàãóñ; 10– äèñòàëüíûé êîíåö ãåíèòàëü- íîãî êîëüöà. Ìàñøòàáíûå ëèíåéêè: 1,0 ìì (1); 0,5 ìì (3, 4); 0,2 ìì (2, 7—10); 0,1 ìì (5, 6). Pronotum 1.5 times as wide as long; anterior angles rectangular, posterior angles slightly developed (fig. 2, 3). Sides not bulging, lateral margins not flattened. Pubescence not parted to two sides (fig. 2, 4). Surface with dual punctation, looking as if pitted because of large punctures (fig. 2, 5). Scutellum triangular. Elytra 1.5 times as long as wide and 2.1 times as long as pronotum. Surface with dual punctation and wrinkles (fig. 2, 6). Metasternum 1.5 times as long as 1st visible abdominal sternite. Anterior rims approximated and straightened medially. (fig. 2, 7). Legs. Fore tibiae not widened to their apices. Hind tarsus more than 0.8 times as long as hind tibia; 1st segment 1.3 times as long as 2nd segment; 2nd segment 1.9 times 199 as long as 3rd one; 3rd segment 1.3 times as long as 4th segment; 5th segment almost as long as the 2nd one; claws large (fig. 2, 8). Aedeagus. Penis narrows to its apex, 9.3 times as long as wide at apex; sudden- ly widened in basal third. Internal penial sac with longitudinal row of 6—7 small spines and with 3 spines of medium sizes; one of them probably representing end of short bifurcated “brush”. Parameres shortly bifurcated at ends, outgrowths nearly reaching paramere apices (fig. 2, 9). Cross-piece of distal end of genital ring looking like thin band slightly widened at every side (fig. 2, 10). Length 2.25 mm, width 1.10 mm. Differential diagnosis. The new species differs from other of black-coloured Lasioderma species occurring in examined region (L. aterrimum; L. obscurum (Solsky, 1867)) in the following characters. L. morulum differs from L. aterrimum in oblong 6—9th antennal segments (4—10th antennal segments strongly transverse in L. aterri- mum (Iablokoff-Khnzorian, 1976, fig. 10)). L. morulum differs from L. obscurum in round eyes and pubescence not parted to two sides on pronotum (eyes shortly oval and pubescence parted to two sides on pronotum in L. obscurum). Etymology. Species name comes from the Latin morulus (black, dark, gloomy), reflecting colour of its body. Lasioderma prolixum sp. n. (fig. 3, 1—12) Material. Holotype {: [Russia,] Krasnodar Terr., Severskij Distr., Ubinskoe Forestry, 21.06.1971 (Nikitsky) (as “Lasioderma obscurum Sols.” M. Dubrovina det.) (ZMUM, coll. Nikitsky). Description. Habitus as on fig. 3, 1. Beetle blackish-brown, tinged with dark red; margins and elytral apices not light-coloured. Pubescence dark grey, fine, homoge- nous, appressed. Head. Frons slightly convex. Eyes round, slightly convex, separated by about 2.5eye diameters. Antenna: 1st segment unusually flattened; 3rd segment 1.4 times as long as the 2nd one; 4—5th segments with oblique and concave upper margin and con- vex lateral one; 6—9th segments strongly transverse, their upper and lower (i. e. later- al) margins straight. (10th and 11th segment lost) (fig. 3, 2). Pronotum 1.47 times as wide as long; anterior angles slightly acute, posterior ones very clear, not rounded (fig. 3, 3). Lateral margins not flattened. Pubescence not parted to two sides (fig. 3, 4). Surface with dual punctation, and more large punctures are nearly as large as small ones (fig. 3, 5). Difference between large and small punc- tures is more distinct near basal margin (fig. 3, 6). Scutellum triangular with blunt top. Elytra 1.4 times as long as wide and 2.3times as long as pronotum. Surface with fine, uniform punctation (fig. 3,7). Metasternum 1.5 times as long as the 1st visible abdominal sternite; anterior rims approximated in the middle, 1st rim is straight, 2nd one a little curved forwards (fig. 3, 8). Legs. Fore tibiae dilated to their apices (fig. 3, 9); all tibiae flattened. Tarsi short: hind tarsus 0.54 times as long as its tibia; 1st segment of middle tarsus 3 times as long as the 2nd one; 2nd segment 1.2 times as long as the 3rd one; the latter 1.2 times as long as the 4th one; 5th segment 1.4 times as long as the 2nd one (fig. 3, 10). Aedeagus. Penis narrows to rounded apex, 7.2 times as long as wide before rounded apex. Internal penial sac has apical and basal “brushes”; basal ends are com- pleted with 3 small needle-shaped sclerites in apical “brush”. The apical end of basal “brush” is completed with rather small digitate sclerite. Basal parts of parameres (from dorsal surface) with separated lobes. Side outgrowth consists of two halves and reach- es the paramere apex. Phallobase short (fig. 3, 11). Cross-piece of genital ring is rib- bon-like, strongly curved (fig. 3, 12). 200 Fig. 3. Lasioderma prolixumsp. n., holotype {: 1– general outline of body; 2– antenna; 3– anterior part of body, lateral view; 4 – pronotum, dorsal view (with pubescence pattern); 5 – punctation on pronotal disc; 6– punctation near pronotal basal margin; 7– punctation on elytral disc; 8– the middle of metaster- num and 1st visible abdominal sternite; 9– fore tibia; 10– middle tarsus; 11– aedeagus; 12– distal part of genital ring. Scale bar: 1.0 mm (1); 0.5 mm (3, 4, 8, 9); 0.2 mm (2, 10—12); 0.1 mm (5—7). Ðèñ. 3. Lasioderma prolixum sp. n., ãîëîòèï {: 1 – îáùèé êîíòóð òåëà; 2 – óñèê; 3 – ïåðåäíÿÿ ÷àñòü òåëà, âèä ñáîêó; 4 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ ðèñóíêîì îïóøåíèÿ); 5 – ïóíêòèðîâêà íà äèñêå ïåðåäíåñïèíêè; 6– ïóíêòèðîâêà ïåðåäíåñïèíêè ó áàçàëüíîãî êðàÿ; 7– ïóíêòèðîâêà íà äèñêå íàä- êðûëüåâ; 8 – ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî áðþøíîãî ñòåðíèòà; 9 – ïåðåäíÿÿ ãîëåíü; 10 – ñðåäíÿÿ ëàïêà; 11 – ýäåàãóñ; 12 – äèñòàëüíûé êîíåö ãåíèòàëüíîãî êîëüöà. Ìàñøòàáíûå ëèíåéêè: 1,0 ìì (1); 0,5ìì (3, 4, 8, 9); 0,2 ìì (2, 10—12); 0,1 ìì (5—7). Length 3.20 mm, width 1.75 mm. Differential diagnosis. The new species belongs to the group of yellow- or red-brown beetles of the genus Lasioderma with pubescence not parted to two sides on pronotum, fore tibiae dilated to their apices. L. prolixum differs from the cosmopolitan 201 L. serricorne by clearly developed posterior angles of the pronotum, in strongly trans- verse middle antennal segments with straight lower margins, approximated anterior rims of the metasternum, by very dark (nearly black) body colour (in L. serricorne, prono- tum posterior angles not developed, middle antennal segments with convex lateral mar- gin, anterior rims evenly rounded in metasternum, body greyish-red, antenna yellow). These species differ also in penial armaments (see article by Español, 1972, fig. 33; and fig. 3, 11 in this article). Etymology. Species name comes from the Latin prolixus (wide, broad, exten- sive), reflecting its body shape. Lasioderma sparsum sp. n. (fig. 4, 1—10) Material. Holotype {: [Russia, Krasnodar Terr.] Anapa, 27.07.1962 (V. Meshcheryakov) (ZMUM). Description. Habitus as on fig. 4, 1. Beetle brown: head, pronotum and abdomen black-brown, elytra reddish-brown, elytral apices with yellow-red spot. Pubescence light grey, heterogeneous, raised, rather long, arranged with uneven and unclear transverse stripes and spots on elytra. Elytral surface with 2—3 costae near suture and lateral margins (fig. 4, 2). Head. Frons almost flat. Eyes round, convex, separated by 1.6 eye diameters. Antenna: 3rd segment 1.5 times as long as 2nd; 4th segment with oblique and straight upper and straight lateral margins; 5—10th segments with slightly convex lateral and straight upper margin. 11th segment 3 times as long as wide and 1.5 times as long as 10th segment. All segments oblong (fig. 4, 3). Last segment of maxillary palpus spin- dle-shaped. Pronotum 1.55 times as wide as long; anterior angles rectangular, posterior ones slightly developed (fig. 4, 4). Pronotum flattened at posterior angles. Pubescence part- ed to two sides and arranged unevenly on pronotum surface (fig. 4, 5). The latter with uniform punctation, its punctures separated by 0.25—0.5 puncture diameters (fig. 4, 6). Scutellum semioval. Elytra 1.6 times as long as wide and 2.6 times as long as pronotum. Surface with dual punctation, small punctures separated by 1—2 puncture diameters (fig. 4, 7). Metasternum twice as long as the 1st visible abdominal sternite; anterior rims approximated a little in the middle and widely straightened (fig. 4, 8). Legs. Fore tibiae not dilated to the apices. 1st segment of middle tarsus 1.5 times as long as the 2nd one; 2nd segment 1.8 times as long as the 3rd one; 3rd segment 1.1times as long as the 4th one; 5th segment almost as long as the 2nd one (fig. 4, 9). Aedeagus. Internal penial sac turned out, its armaments consists of longitudinal row of 8 small spines, 2 detached small spines, and 3 large spines (fig. 4, 10). Length 3.5 mm, width 1.65 mm. Differential diagnosis. The new species differs from other species of the genus Lasioderma without black body colouration, with pubescence parted to two sides on pronotum, and distributed in the named region (L. babadjanidesi, L. redtenbacheri (Bach, 1852), L. mikheechevi, L. kiesenwetteri, L. tauricum sp. n.) in the following char- acters. L. sparsum has rims of metasternum which are approximated and straightened in the middle (these rims evenly rounded in L. babadjanidesi, L. redtenbacheri; and rims slightly approximated and 1st rim hardly straightened in L. mikheechevi). L. sparsum differs from L. kiesenwetteri by brown colour of body and pubescence parted to two sides on pronotal disc (body yellow-red and pubescence parted near apical margin of pronotum in L. kiesenwetteri). L. sparsum differs from L. tauricum in tarsi structures (2nd segment about as long as the 1st one, and 5th segment 1.7 times as short as the 2nd one in L. tauricum, whereas 1st segment 1.5 times as long as the 2nd one and 5th segment as long as the 2nd one in L. sparsum). Also L. sparsum differs from the named 202 Fig. 4. Lasioderma sparsumsp. n., holotype {: 1– general outline of body; 2– right elytron with pubescence pattern; 3 – antenna; 4 – anterior part of body, lateral view; 5 – pronotum, dorsal view (with pubescence pattern); 6 – punctation on pronotal disc; 7 – punctation on elytral disc; 8 – the middle of metasternum and 1st visible abdominal sternite; 9 – middle tarsus; 10 – internal penial sac (turned out). Scale bar: 1.0mm (1); 0.5 mm (2, 4, 5, 8); 0.2 mm (3, 9); 0.1 mm (6, 7, 10). Ðèñ. 4. Lasiodermasparsumsp. n., ãîëîòèï {: 1– îáùèé êîíòóð òåëà; 2– ïðàâîå íàäêðûëüå ñ ðèñóí- êîì îïóøåíèÿ; 3 – óñèê; 4 – ïåðåäíÿÿ ÷àñòü òåëà, âèä ñáîêó; 5 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ ðèñóíêîì îïóøåíèÿ); 6 – ïóíêòèðîâêà íà äèñêå ïåðåäíåñïèíêè; 7 – ïóíêòèðîâêà íà äèñêå íàä- êðûëüåâ; 8 – ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî áðþøíîãî ñòåðíèòà; 9 – ñðåäíÿÿ ëàïêà; 10 – ýíäîôàëëóñ (âûâåðíóò). Ìàñøòàáíûå ëèíåéêè: 1,0 ìì (1); 0,5 ìì (2, 4, 5, 8); 0,2 ìì (3, 9); 0,1 ìì (6, 7, 10). species in spotted pubescence on elytra and armaments of internal penial sac (see Toskina, 1999: figs 44, 92, 140, Logvinovskij, 1985: fig. 228, and this paper: fig. 4,10 and 5,10). 203 Etymology. Species name comes from the Latin sparsus (sparse, diffuse, dis- persed), reflecting diffused or spotted distribution of pubescence on the body. Lasioderma tauricum sp. n. (fig. 5,1—11) Material. Holotype {: Crimea, Karadag, 5.08.1991, meadow, net sweeping (I. T. A.). (“Lasioderma kiesenwetteri Schils.” I. T. A. det., 1992) (ZMUM). Description. Habitus as on fig. 5,1. Elytra brownish-yellow with yellow apices; pronotum and head brownish; ventral surface black-brown; antennae (except brown 1stsegment) and legs light yellow. Pubescence yellowish, homogenous, slightly raised. Head. Frons slightly convex. Eyes round, convex, separated by 1.7 eye diameters. Antenna: 3rd segment almost as long as the 2nd one; 4—5th segments with concave upper margin and convex lateral one; these two segments are of smaller size than next 6—10th segments; the latters have straight upper and lateral margins. 11th segment about 3 times as long as wide and 1.9 times as long as the 10th segment (fig. 5,2). Last segment of maxillary palpus is fusiform, about 3.5 times as long as its thickness, with oblique truncate apex (fig. 5, 3). Pronotum 1.7 times as wide as long; anterior angles rectangular, posterior angles obtuse, rounded. Pronotum convex more strongly near basal margin; sides not bulging (fig. 5,4). Pubescence parted to two sides on disc (fig. 5, 5). Surface with fine, uniform punctation, where punctures separated by 0.5—1.0 puncture diameter (fig. 5, 6). Scutellum semioval. Elytra 1.45 times as long as wide and 2.4 times as long as pronotum. Surface with uniform punctation, punctures separated by 1—2 puncture diameters (fig. 5, 7). Metasternum about 1.5 times as long as the 1st visible abdominal sternite; ante- rior rims approximated in the middle, 1st rim is straightened, the 2nd one slightly curved forward (fig. 5, 8). Legs. Fore tibia not dilated to the apices. Tarsi as long as or longer than tibiae. 1st segment of hind tarsus 1.2 times as long as 2nd segment; 2nd segment 3 times as long as 3rd segment; 3rd segment 1.2 times as long as 4th segment; 5th segment 1.7times as short as 2nd segment (fig. 5, 9). Aedeagus. Penis straight, almost parallel-sized, not narrowed to its apex; about 8 times as long as wide at apex; the latter has quadrate notch. Internal penial sac with two longitudinal rows (4 + 9) of small spines, 5 small spines without arranging in rows, and 1 large spine in apex. Penis is filled with thick, double “brush” in its two fifth of basal part. Ends of parameres not deeply bifurcated, paramere outgrowth not reaching paramere apex (fig. 5, 10). Cross-piece of genital ring segment very thin medially but strongly widened to its sides (fig. 5,11). Length 2.4 mm, width 1.2 mm. Differential diagnosis. The new species belongs to the group of yellow- brown species with pubescence parted to two sides on pronotum and tibiae not widened to their apices, which includes L. babadjanidesi, L kiesenwetteri, L. mikheechevi, L.redtenbacheri, and L. sparsum sp. n. L. tauricum differs from L. babadjanidesi and L.redtenbacheri by approximated anterior rims of metasternum (anterior rims evenly rounded in those species), and from L. kiesenwetteri, L. mikheechevi, and L. sparsum sp. n. first of all in shape of antennal segments: 4—5th segments are smaller and with convex lateral margin whereas the next segments are larger and with straight lateral margin; secondly, 1st tarsal segment almost as long as the 2nd segment and 5th seg- ment is very short (1.7 times as short as the 2nd segment) in L. tauricum. In compared species, only 4th segment has oblique upper margin, 5—10th segments more or less uni- form, the 1st tarsal segment 1.5 times and more as long as the 2nd segment, and 5th tarsal segment as long as the 2nd one in L. mikheechevi and L. sparsum and a little 204 Fig. 5. Lasioderma tauricumsp. n., holotype {: 1– general outline of body; 2– antenna; 3– last segment of maxillary palpus; 4 – anterior part of body, lateral view; 5 – pronotum, dorsal view (with pubescence pattern); 6 – punctation on pronotal disc; 7 – punctation on elytral disc; 8 – the middle of metasternum and 1st visible abdominal sternite; 9– hind tibia and tarsus; 10– aedeagus; 11– distal part of genital ring. Scale bar: 1.0 mm (1); 0.5 mm (4, 5, 8); 0.2 mm (2, 9—11); 0.1 mm (3, 6, 7). Ðèñ. 5. Lasioderma tauricum sp. n., ãîëîòèï {: 1 – îáùèé êîíòóð òåëà; 2 – óñèê; 3 – ïîñëåäíèé ÷ëå- íèê ÷åëþñòíûõ ùóïèêîâ; 4 – ïåðåäíÿÿ ÷àñòü òåëà, âèä ñáîêó; 5 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ ðèñóíêîì îïóøåíèÿ); 6 – ïóíêòèðîâêà íà äèñêå ïåðåäíåñïèíêè; 7 – ïóíêòèðîâêà íà äèñêå íàä- êðûëüåâ; 8– ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî áðþøíîãî ñòåðíèòà; 9– çàäíèå ãîëåíü è ëàïêà; 10– ýäåàãóñ; 11– äèñòàëüíûé êîíåö ãåíèòàëüíîãî êîëüöà. Ìàñøòàáíûåëèíåéêè: 1,0 ìì (1); 0,5ìì (4, 5, 8); 0,2 ìì (2, 9—11); 0,1 ìì (3, 6, 7). shorter than the 2nd segment in L. kiesenwetteri. Difference in penial armaments is shown in fig. 5, 10 and 4, 10 (also compare to: Toskina, 1999: fig. 44, 92, 140, and Logvinovskij, 1985: fig. 228). Etymology. Species name is derived from the Latin geographic name Tauria (Crimea), the type locality of the new species. Key to species of the genus Lasioderma from Crimea and the Caucasus Òàáëèöà äëÿ îïðåäåëåíèÿ âèäîâ ðîäà Lasioderma èç Êðûìà è Êàâêàçà In this key, some data from the keys by Iablokoff-Khnzorian (1976), Logvinovskij (1985), Toskina (1999), and articles and books by Arnoldi (1965), Español (1970, 1972, 1992), Zahradník (1998, 2000 et al.), and some others, were used. 1. Beetles black (excepting light margins of pronotum and elytra) or having something of black colour. .............................................................................................................................................................. 2 - Beetles without black colour, but can be blackish-brown. .................................................................. 4 2. Antenna: 5—10th segments transverse, their lateral (i.e.,lower) margins convex (Iablokoff-Khnzorian, 1976, fig. 1,10). Pronotum slightly campaniform, pubescence forms parting on pronotal disc; beetles wholly black, pubescence grey (Roubal, 1916, S. 48). Length 2.5—3.4 mm. Czechoslovakia, Caucasus; Crimea. ...................................................................................................... L. aterrimum Roubal, 1916.