SPIXIANA 25-42 München, 01. März 1998 ISSN 0341-8391 New data on Chrysomelidae of Nepal (Insecta, Coleoptera) L. N. Medvedev and E. Sprecher-Uebersax Medvedev, L. N. & E. Sprecher-Uebersax (1998): New data on Chrysomelidae of Nepal (Insecta, Coleoptera). 4 Spixiana 21/1: 25-42 Recently studied material from Nepal has resulted a new genus Sphenorella and 7 new species (Oreomela annapurnae, Apophylia brancuccii, Japonitata diformis, Mono- lepta impressicollis, Doryscus niger, Paraplotes nepalensis, Dercetina nepalica) as well as 56 species firstly recorded for Nepal. 7 new synonymies, 1 new homonymy, 2 re- descriptions and 7 rectifications are proposed. Keys for Nepalese Apophylia, Orien- tal Euphitrea micans-group and one to divide Paridea octomaculata from P. tetraspilota are given. Dr. Lev. N. Medvedev, Inst. Evol. Morphol. & Ecol. Anim., Leninsky prospect 33, Moscow 117 071, Russia Eva Sprecher, Naturhistorisches Museum, Augustinergasse 2, CH-4001 Basel, Switzerland. Introduction As a part of our project <Catalogue of Chrysomelidae from Nepal= we propose the next part of taxonomical investigation on leaf beetles from Nepal. Our studies based mostly on the material of the Natural History Museum in Basel, but also on the collection of Staatliches Museum für Naturkunde Stuttgart. In the Natural History Museum of Basel the material of Nepalese leaf beetles is very rich and therefore gives us the chance of large investigations. In this article descriptions of new taxa, synonym- ical notes, nomenclature changes as well as a large list of species firstly recorded for Nepal and keys for a few genera or species group are given. The material is deposited in the collections mentioned below. The following abbreviations are used: NHMB Naturhistorisches Museum, Basel SMNS Staatliches Museum für Naturkunde, Stuttgart LM Collection of L. Medvedev, Moscow List of species Subfamily Eumolpinae Nodina robusta Jacoby N. crassipes Jacoby, 1908 and N. major Kimoto & Gressitt, 1982 are new synonyms of this species, which has rather variable colour of legs, from entirely fulvous to strongly darkened with fulvous apices of tibiae and tarsi. 25 Subfamily Chrysomelinae Agrosteomela fallaciosa Stal Material: 1 ex., Mt. Everest. New for Nepal, was known from Sikkim and Assam. Phrathora abdominalis Baly Material: 1?, Langtang National Park, Kyanjin Gompa, 4000 m, 12. V. 1988, leg. S. Bily. New for Nepal, was known from North India. Gonioctena inconditus Weise Material: 13, East Nepal, Arun Valley, Num-Chichila, 1500-1900 m, 17. VI. 1980, leg. C. Holzschuh. New for Nepal, was known from Malaysia. Oreomela (s. str.) annapurnae, spec. nov. Fig. 1 Types. Holotype: d, Nepal, Annapurna Mts. (Everest) (LM). Diagnosis. Near O. indica Jacoby, 1896, differs in colour, evenly rounded (not cordiform) sides of prothorax, and structure of aedeagus with well developed and acute apical process. Description Upperside and legs dark bronze, antennae and underside practically black.Body elongate, widened posteriorly, broadly rounded at apex. Head much more narrow than prothorax, sparsely punctate and extremely finely shagreened, with deep transverse groove dividing frons and clypeus. Antennae with segment 2 subglobose, segment 3 elongate, about 1.5 times as long as 2= and slightly longer as 4"; next segment elongate, subequal to 4th. Prothorax 1.6 times as wide as long, with maximal width in middle, sides evenly rounded, anterior angles rounded, posterior angles obtuse, surface moderately convex, shagreened and not densely uniformly punctate; all punctures of the same size. Scutellum very short, triangular, densely shagreened. Elytra ovate, 1.25 times as long as wide, with maximal width just behind middle, surface moderately convex, finely and sparsely punctate, punctures distinctly smaller than on prothorax, interspaces shining. Fore tarsi of d not widened, pubescent below. Claws simple. Aedeagus (Fig. 1) longitudinally concave on underside, especially on apical process. Length: 5.5 mm. Oreomela (s. str.) himalayensis nepalica, subspec. nov. Fig. 2 Types. Holotype: d, Nepal, Prov. Karnali, Distr. Dolpa, Kagmara Lekh Garpung Khola, Hochplateau, 4100 m, 12.V.1995, leg. A. Weigel (LM). Diagnosis. This subspecies is identical with nominative form in structure of aedeagus, but differs in having distinct metallic colour, more finely punctured prothorax and very shining upperside. Description Upperside aeneous green, antennae, legs and underside black. Body elongate, widened posteriorly. Head finely and sparsely punctate, shining. Antennae with segment 3 distinctly longer than 2" or 4". Prothorax 1.7 times as wide as long, feebly cordiform, with obtuse hind angles; surface shining, finely 26 un | Figs 1, 2. Oreomela, aedeagus ventral. 1. O. annapurnae, spec. nov. 2. O. himalayensis nepalica, subspec. nov. Figs 3-5. Apophylia, antennae of d. 3. A. brancuccii, spec. nov. 4. A. aeruginosa Hope. 5. A. samoderzhenkovi Medvedev. punctate and shagreened. Scutellum short, triangular, shining. Elytra 1.3 times as long as wide, with maximal width in middle, surface shining, finely punctate, more or less wrinkled among punctures. Fore tarsi of d not widened. Aedeagus (fig. 2) longitudinally concave on underside, especially in apical part, with membranous apical third and simple rounded triangular apex. Length: 5.2 mm. Subfamily Galerucinae Apophylia aeruginosa (Hope) Figs 4, 15 We have at our disposal 384 from Nepal with a specific structure of abdomen and rather moderate size, but in all other characters fully identical with A. aeruginosa Hope. Maulik (1936) in his description of A. aeruginosa mentions nothing about spines on abdomen, but very possibly he overlooked this character (as in the case of A. nilakrishna Maulik). Because of this we propose a description of the d which we accept for A. aeruginosa Hope. Redecription Black, basal half of antennae and legs fulvous, elytra metallic green or bluish green, tarsi more or less darkened. Anterior part of head, including frontal tubercles practically smooth and shining, hind part very densely punctate, dull. Antennae (Fig. 4) about % of body lenght, in d with thickened preapical and shortened apical segments. Prothorax 1.7-1.8 times as wide as long, with maximal width in anterior third, anterior margin practically not elevated and not incised in middle, hind margin feebly concave, surface with shallow impression on middle line and groove on each side, finely and densely punctate. Elytra very densely punctate. Segment 1 of anterior and mid tarsi of d very narrow, elongate and curved. Abdominal sternites 1 and 2 of d with two sharp teeth each on hind margin (Fig. 15), last sternite with small groove before apex. Aedeagus symmetrical, parallel-sided, with narrowed apical third (Fig. 6) Length: 6.3-6.7 mm. Material used for description: East Nepal, Dhawalagiri, Myagdi Distr., Ghara Khola, Shikka-Tatopani, 1100- 2000 m (J. Probst); Nepal, Bagmati, Sindhupalchok; Gangiwal-Parahang, 1700-2500 m, 8.V1.1989 (M. Brancucci); 13 without head and prothorax, same region, Sarmatang-Gangiwal, 2500 m, 5.V1.1969 (M. Brancucci). 27 Apophylia brancuccii Medvedev, spec. nov. Fig. 3 Types. Holotype: 3, East Nepal, Mechi, Dobhan-Phulvari, 800-1200 m, 8.V1.1985, leg. M. Brancucci (NHMB). - Paratypes: 1 ex., E. Nepal, leg. M. Brancucci: same locality; 10 ex., Arun Valley, Pharicot, 550 m, 13.V1.1983; 2 ex., Koshi, Dharan, 18.V1.1985; 1 ex.,Koshi, Simraghat-Lumbughat, 450 m, 14.V1.1985; 1 ex., Arun Valley, Tumlingtar, 450 m, 26.V.1983; 4 ex., Hile, Arun R., 200-300 m, 26.V.1983; 3 ex., East Nepal, Arun Valley, Mongmaya-Surtibari, 400 m, 1.V1.1992, leg. J. & J. Probst (NHMB, 2 in LM). Diagnosis. Near A. metallica Jacoby, 1904, poorly known species from South India, differs by bicolor- ous head and unicolorous elytra. From Nepalese species it differs by characters indicated in the key given below. Description Fulvous; head behind frontal tubercles and middle of mesosternum black, apical half of antennae darkened, elytra metallic green. Head roughly and densely punctate. Antennae thin, with elongate seg- ments (Fig. 3). Prothorax twice as broad as long, with maximal width in anterior third, anterior margin elevated, incised in middle, hind margin concave, surface with impressed middle line and large shallow groove on each side, densely punctate. Elytra very densely punctate. Segment 1 of anterior and mid tarsi not enlarged in d. Aedeagus (Fig. 7) lanceolate, with triangular acute apex, slightly asymmetrical. Length 4.8-5.7 mm. Key to Apophylia of Nepal Itothoraxtulvous,sometimesswithäblackespotss re... rs 2: =" Brothoraxeblaek.......n. en. een nee er ee DE REINE 3. 2 Erothoraxswithlblackspols. .......n.....02..22 une erseanensnesan sten naneeber ern ecte ehren EEE © - Prothorax without black spots. Head bicolorous, legs and underside except metasternum fulvous. kensthr 1.8.9 7ammee. eeneen ee ee ee brancuccii, spec. noV. 3. Erothoraxswith.1 black spot. ...........eunuanssee sBnrsetesnesteenteeaerse ssseenne esessEeIeER 4. - Prothorax with 3 black spots. Head fulvous with black vertex. Legs fulvous. Elytra metallic green. imedeasusiseertie29. Pensth, 9.5.0. 910m reR e EEE crotchi Jacoby 4. Antennae and legs entirely or mostly fulvous. Head bicolorous. Elytra metallic green. Length ATI ee ee EURE sericea Fabricius -4 Antennae and legs entirely or mostly black. Head black. Elytra usually bronze or dark green. kensth8 6.3.7.5. mm en ee anieeenensensasren ge eetor ernsten maculata Kimoto 5. Head bicolorous. Margins of prothorax often fulvous. Legs fulvous. ...........eeeneeseneeeee 6. = Headrentirelyoblacke....:eterenreanseeneenssraentesnn nneen enen eress aengnesn sensaSsSe neEneEnEs 78 6. Elytra metallic green. Antennae of d see fig. 5. Aedeagus lanceolate, asymmetrical, with very acute äpexulengthr4.8-5.5.mmi......er. ne na es samoderzhenkovi Medvedev - Elytra practically black. Length 6-7.3 mm. Possibly a dark form of preceding species ........... sp. A 7.>legs. black, /knees sometimes fulveussr..nn.ta. en see ee EEE 8. 4eFegszfulvous, tarsifandapices/offfemorarsometimes darkened...... RER 10. 8. cd antennal segments 7-8 not thickened, about 3 times as long as wide. Body smaller. ................ 9). - d antennal segments 7-8 thickened, about 1.5 times as long as wide; abdominal segments 1 and 2 withytworteethreach<Bodyzlarser, 7.7.8: mer ne nilakrishna Maulik 9. Frontal tubercles smooth. Elytra green. d abdominal segment 1 with 2 hairy brushes. Aedeagus not narrowed in apical part, extreme apex short triangular, curved downwards (Fig. 8). Length SPS LO EEREREN schawalleri Medvedev 28 Figs 6-9. Apophylia, aedeagus lateral and dorsal. 6. A. aeruginosa Hope. 7. A. brancucci, spec. nov. 8. A. scha- walleri Medvedev. 9. A. crotchi Jacoby. - Frontal tubercles usually shagreened. Elytra dark violaceous or green with cupreous suture. d abdomen simple, aedeagus cuneiform with apex not curved downwards. Length 4.1-5.9 mm. SUR RRREIRIRER RER REEL. On Saninneneeecunetn-anineensniaesschnensnenrens pen antenne sen ee er himalayana Medvedev 10. Elytra green. d antennal segments 7-9 thickened (Fig. 2).Abdominal segments 1 and 2 with two teethleach. Aedeagus see fie. 6. Length 6.36.72 MM ........eueeceseenenenesennensesnensneransteenenes aeruginosa Hope - Elytra green with golden suture. Body small, 5 mm. Unclear species, very possibly identical with prececdimegoner te eeB rennen rennt assamensis Jacoby Sastroides dohertii Maulik Material: 1 ex., Koshi, Chauki, 2700 m, 2.V1.1985 (M. Brancucci); 1 ex., Ganesh Himal, Somathang, 3270 m, 15.V1.1993 (M. Hroblay). New for Nepal, was known from Burma and Laos. Mimastracella pubicollis Samoderzhenkov Material: 2 ex., Arun Valley, Lamobagar Ghola, 1400 m, 28.-31.V.1980 (C. Holzschuh). New for Nepal, was described from Vietnam. Mimagitocera flava (Jacoby) Material: 1 ex., Taplejung Distr., Gunsa Khola, 2400-2600 m, 12.1X.1983 (J. Martens & W. Schawaller). New for Nepal, was known from Assam. Trichocerophysa latifascia Gressitt & Kimoto Material: 1 ex., Arun Valley, Lamobagar Gola, 1400 m, 8.-14.V1.1983 (M. Brancucci). New for Nepal, was known from China. 29 Agetocera lobicornis Baly Material: 2 ex., <Himalaya, Mt. Everest=. New for Nepal, was known from North India. Aulacophora carinicauda Chen & Kung Material: 15, Gorkha Distr., Ghorka-Khanckok, 1700-2300 m, 22.V.1990, leg. J. Probst. New for Nepal, was known from South China and Vietnam. Cerophysella viridipennis Allard Material: 14, 12, Arun Valley, Lamobagar Ghola, 1400 m, 8-14.V1.1983 (M. Brancucci); 17, Arun Valley, Mure, 2000 m, 2.-8.V1.1983 (M. Brancucci); 15, Thamur Valley, Dhankuta-Bile, 1150-2000 m, 24.-25.V.1983 (M. Brancucci). New for Nepal, was known from Indochina. Genus Taumacera Thunberg The genus Oedicerus Kollar & Redtenbacher, 1848 is anew synonym of Taumacera. A typical Taumacera has no enlarged antennal segments in d, or segments 3-7 are modified; in the subgenus Cerophysa Chevrolat, 1837 segments 5-6 or more often segment 8 are enlarged. In Oedicerus 85 have modified 5th and 6th segments, Therefore we can reduce Oedicerus to a full synonym of Taumacera, because all differences are only in secondary sexual characters. Taumacera monstrosa Jacoby Material: 1 ex., Arun Valley, Tumlingtar-Khandbari, 450-1100 m, 27.V.1983 (M. Brancucci; 1 ex., Arun Valley, Khandbari-Bhotebas, 950-1850 m, 5.V1.1992, leg. J. Probst. New for Nepal, was known from Burma. Taumacera mandarensis Jacoby The species is rather variable: prothorax entirely red or with black bilobed spot in middle, rarely entirely black; antennae black or with red basal segment; head more or less darkened, legs from fulvous to red. Form of aedeagus and structure of d antennae are identical in all these colour variations. T. andrewesi Jacoby, 1904 is very possibly identical with this species. Phyllobrotica komiyai Takizawa Material: 1 ex., Dolakha, Tama Kola, 850-1100 m, 24.-29.V.1989 (M. Brancucci). New for Nepal, was known from West Bengal. Liroetis apicicornis Jacoby Material: 1 ex., Arun Valley, Num, 1550 m, 5.-6.V1.1983 (M. Brancucci); 1 ex., Bagmati, Sindhupalchok, 2500 m, 6.-7.V1.1989 (M. Brancucci). New for Nepal, was known from South India. 30 E R I l | FL Figs 10-11. Paridea, aedeagus lateral and dorsal. 10. P. tetraspilota Hope. 11. P. octomaculata Baly. Figs 12-13. Doryscus, aedeagus ventral. 12. D. niger, spec. nov. 13. D. testaceus Jacoby. Fig. 14. Paraplotes nepalensis, spec. nov, antenna. Fig. 15. Apophylia aeruginosa Hope, spines on abdominal sternites 1 and 2. Figs 16-18. Sphenorella maculata (Kimoto & Takizawa), elytral pattern. 16. From Manegero. 17. From Phulcho- ki. 18. From Chautara. Mimastra nitida Maulik Material: 2 ex., Bagmati, Nuwakot, 2300 m, 15.V1.1989 (M. Brancucci). New for Nepal, was known from North India. Japonitata diformis, spec. nov. Types. Holotype: 2, East Nepal, Arun Valley, Mure, 2000 m, 2.-8.V1.1983 (M. Brancucci), spotted specimen (NHMB). - Paratypes: 69%, same locality (NHMB, 2 in LM). Diagnosis. We place this species in Japonitata, although it has closed anterior coxal cavities, because all other characters fully correspond to this genus. It resembles more or less J. eberti Kimoto, 1970, but differs immediately in having only one ridge on elytra. Description Fulvous, antennae dark brown except one or two basal segments, elytra often with two large black spots: one behind basal convexity, another behind middle. Head shining, impunctate; interantennal space narrow and carinate, frontal tubercles delimited behind with transverse groove. Antennae reach behind middle of elytra, segment 3 distinctly longer than 2, segment 4 a little longer than 3, next segments subequal to 4. Prothorax 1.8 times as wide as long, with maximal width before middle, sides feebly rounded, anterior margin feebly concave, basal margin straight, with oblique emargination at hind angles, surface shining, impunctate, with large shallow impression on each side. Elytra with high humeral tubercle and sharp lateral ridge, dividing vertical part of elytra and distinct to apical slope, basal convexity distinct, surface smooth, shining, impunctate, but with transparent, more or less darkened dots. Anterior coxal cavities narrowly closed. Length: 4.8-5.8 mm. 31 Paridea (Semacia) lateralis Medvedev & Samoderzhenkov Material: 254, 1?, West Nepal, Chitre, Ghar Khola, 2400 m, 26.-31.V.1984 (C. Rai) (NHMB, LM). New for Nepal, was described from Vietnam. Paridea octomaculata (Baly) Fig. 11 This very usual species is rather variable, the elytral spots are sometimes enlarged and more or less united together. In such case the species resembles very much another species, P. tetraspilota (Hope, 1831). However, they differ well in structure of aedeagus as follows: Key to Paridea octomaculata and tetraspilota 1. Aedeagus rather straight and thin in lateral view, apex rounded, with long apical process, curved downwards under right angle. Underside without grooves (Fig.10) ........... tetraspilota Hope -4 Aedeagus thickened in middle and curved in lateral view, apex truncate with short apical process, rather feebly curved downwards; underside with groove before apex and another along the middle are octomaculata Baly Cneorane birmanica Jacoby Material: 1 ex., Arun Valley, Num, 1500 m, 5.-6.V1.1983 (M. Brancucci). New for Nepal, was known from Burma. Cneorane dohertii Maulik Material: 1 ex., Bagmati, Nuwakot, Pati Bhanjyang, 1900 m, 16.-18.V1.1989 (M. Brancucci); 1 ex., Godavari, 1500- 2500 m, 1.V11.1987 (C. Rai). New for Nepal, was known from Assam. Morphosphaera prava Maulik Material: 1 ex., Bagmati, Sindhupalchok, 1700-2500 m, 8.V1.1989 (M. Brancucci). New for Nepal, was known from Assam and Burma. Genus Calomicrus Stephens The genus Charaea Baly, 1878 is a new synonym of Calomicrus. All characters given in the original description correspond fully with the latter genus. Calomicrus balyi, nom. nov. = Charaea flaviventris Baly, 1878 (homonymy with Calomicrus flaviventris Motschulsky, 1860). Material: 12 ex., Kali-G. Khola, Kalopani, 1600-2500 m, 26.V1.1986 (C. Holzschuh); 6 ex., Mure-Num, 1500- 2000 m, 4.-7.V1.1983 (M. Brancucci). New for Nepal, was known from Punjab. 32 Monolepta labiata Jacoby Material: 2 ex., Dhaulagiri, Kali-G. Khola, 1600 m, 18.V1.1986 (C. Holzschuh). New for Nepal, was known from Bengal. Monolepta rufa Takizawa Material: 1 ex., Modi Khola, Randrung, 1100-1800 m, 3.-6.V1.1984 (C. Holzschuh). Monolepta impressicollis, spec. nov. (det. by L. Medvedev) Types. Holotype: d, Assam, Kasiranga (LM). Diagnosis. Resembles M. bimaculipennis Kimoto, 1970, but prothorax with very distinct depressions and more closely punctate, head bicolorous, elytra with clearly transverse fulvous band (not rounded spot). Description Black; anterior part of head, including frontal tubercles, 3 basal segments of antennae, prothorax, broad transverse band in the middle of elytra and legs fulvous; tibiae and tarsi darkened. Body elongate, flattened. Head with deep transverse groove behind frontal tubercles, vertex punctate and finely wrinkled. Antennal segments 2 and 3 subequal, segment 4 almost as long as two preceding together, next segments subequal or slightly shorter than 4th. Prothorax 1.4 times as wide as long, lateral margins almost straight, hind angles distinct, almost rectangular, anterior and posterior margins arcuate, surface with distinct depression on each side just behind middle, densely punctate. Elytra 1.7 times as long as wide, very densely punctate, with sparse hairs, mostly on apical slope. Segment 1 of posterior tarsus about twice as long as next segments combined. Length: 4.2 mm. Monolepta bimaculicollis Jacoby Material: 1 ex., Lamobagar Khola, 1400 m, 8.-14.V1.1983 (M. Brancucci). New for Nepal, was known from South India. Paleosepharia tomokunii Kimoto & Takizawa Material: 1 ex., Godavari, 1500 m, 21.-27.V.1989 (M. Brancucci). This rare species was described from the Nepal by Kimoto & Takizawa, 1983. Paleosepharia pokharensis Kimoto (comb. nov.) This species, described as Monolepta, is transferred to the genus Paleosepharia; P. costata Takizawa & Basu, 1987 is a new synonym of this species. Sinoluperoides marginalis Kimoto Material: 2 ex., Kathmandu Valley, Godawari, 1500 m, 17.V.1983 and 31.V.1987 (M. Brancucci). New for Nepal, was known from Vietnam. 33 Doryscus niger, spec. nov. Fig. 12 Types. Holotype: d, West Nepal, Pothara-Landrung, 1600-1900 m, 7.V.1984, C. Holzschuh (NHMB). Paratypes: 1 ex., same locality; 2 ex., Central Nepal, Bagmati, Sindhupalchok, Gangjwal, 2500 m, 6.-7.V1.1989, C. Holzschuh (NHMB, LM); 12, Bagmati, Sindhupalchok, Dubhachaur-Sarmatang, 1600-2500 m, 3.V1.1989, M. Brancucci (NHMB); 1 ex., Myagdi Distr., Myagdi Khola N Boghara, 1800-2000 m, 21.V.1995, Martens & Schawaller (SMNS). Diagnosis. Very near to the single known species of the genus, D. testaceus Jacoby, 1887, and might be only a dark variation of the latter, but differs in obtuse apex of aedeagus (not pointed as in D. testaceus, fig. 13), black body, smaller size and a little more robust antennae. D. testaceus has sometimes head and elytra more or less darkened to entirely black, but prothorax seems to be always fulvous. Description Body black with fulvous knees, antennae fulvous or darkened, anterior part of head sometimes fulvous. Head impunctate. Antennae with segments 3-10 subequal, each of them about 2.5-3 times as long as wide. Prothorax strongly cordiform, impunctate, with numerous white erect hairs along anterior and lateral margins. Elytra are practically the same as in D. testaceus. Last abdominal sternite of d with deep rounded groove. Aedeagus (Fig. 12) narrow and elongate, with obtuse apex. Length: 3.3-4.4 mm. Palpoxena konbirensis Weise Material: 13, Koshi, Simraghat-Lumbughat, 450 m, 14.V1.1985 (M. Brancucci); 135, Godavari, 1500 m, 23.V.1985 (M. Brancucci). The specimens in question have entirely fulvous upperside, but structure of head is quite identical with P. konbirensis Weise. New for Nepal, was known from Bengal. Paraplotes nepalensis, spec. nov. (det by L. Medvedev) Fig. 14 Types. Holotype: ?, Central Nepal, Kathmandu Valley, Godavari, 1500 m, 23.V.1985, leg. M. Brancucei (NHMB). 4 Paratypes: 2 ex., same locality, 21.-27.V.1989 (M. Brancucci) (NHMB); 1 ex., same locality, 10.- 12.V1.1984 (C. Rai) (LM). Diagnosis. This genus is known from East China, North Vietnam and Taiwan and includes 5 species. Although we have at our disposal only a single 9, it differs well from all known species. P. antennalis Chen, 1942 and P. clavicornis Gressitt & Kimoto,1963 have antennal segments 6-7 robust; P. rugosa Laboissiere, 1933 has other proportions of antennal segments and fulvous body with black elytra; P. frontalis Laboissiere, 1933 has vertex and abdomen black, elytra not fully metallic. Description Head black with fulvous labrum and palpi, antennae fulvous with darkened apical half, prothorax fulvous, scutellum black, elytra blackish blue, underside black with fulvous abdomen, legs fulvous with darkened tarsi. Body robust, widened posteriorly, flattened above. Head impunctate, frontal tubercles elongate triangular with acute anterior angles invading in an interantennal space. Vertex with lon- gitudinal impressed line. Antennae (Fig. 14) about % of body length, thin, with segments cylindrical, except segments 8 and 9, that are more or less triangularly widened at apex; proportions of segments are as 15-6-10-12-11-10-10-8-8-10-15. Prothorax 2.6 times as broad as long, transversely concave, with almost straight and parallel side margins, fore angles protruding, hind angles with quadrate emargin- ation; surface shining and impunctate. Scutellum triangular, impunctate. Elytra 3 times as long as wide in shoulders, broadened behind, with sharp lateral ridge, divided elytra on horizontal and vertical parts; surface without distinct basal convexity, moderately shining, very densely subrugosely punctate. Length: 6 mm. 34