/ Contributions New combinations in the terrestrial orchid genera Caladenia R. Br., Corybas Salisb. and Pterostylis R. Br. (Orchidaceae) for south-eastern Australia GaryN Backhouse DepartmentofSustainabilityandEnvironment,BiodiversityandEcosystemServicesDivision, 8NicholsonStreet,EastMelbourne3002.Email:[email protected] Abstract NewcombinationsaremadeintheterrestrialorchidgeneraCaladeniaR.Br.,CorybasSalisb.andPterostylisR. Br.toaccommodate32newspeciesfromsouth-easternAustraliaandonefromMacquarieIslandthatwerede- scribedingeneranotgenerallyrecognisedbyherbariainAustralia. (7heVictorianNaturalist127(2),2010,55-57) Keywords:Arachnorchis, Bunochilus, Hymenochilus, Oligochaetochilus, Speculantha Introduction Since 2001, over 60 new species of terrestrial References orchids from southern Australia have been de- Backhouse GN (2007) Newcombinations in theterrestrial scribed in genera presently not recognised by orchidgenera CaladeniaR. Br. andPterostylisR. Br. (Or- State and Territory herbaria in Australia (e.g. c1h2i4d,a1c2e3ae-)12f4o.rVictoria,Australia. TheVictorianNaturalist Jones 2006a; 2006b; 2006c; 2007). Some of BarkerRMandBatesRJ(2008)NewcombinationsinPtero- these new species have had new combinations stylis and Caladenia and other name changes in the Or- made in recognised genera (e.g. Backhouse ctahniidcacGeaaredeonfsS2o2u,th102Au-s1t0r3a.lia.JournaloftheAdelaide Bo- 2007; Barker and Bates 2008). However, ad- ClementsMAandJonesDL(2007)AnewspeciesofNema- idnitisoonamlenoefwtshpeesceiesuncroenctoignnuiesetod bgeendeersacri(eb.egd. 4ate0o)c5,e-rb4ao1st1ha.nfdrochmarsaucbtaenrtiasracttiiocnMoafcNq.uadriieenIesmlaunmd.(TOerlcohpiedaac1e1-, Clements and Jones 2007; Jones 2008a; 2008b; Jones DL (2006a) Miscellaneous new species ofAustralian 2008c;2009a;2009b),necessitatingfurthernew JonOerschDiLda(c2e0a0e6.bA)usTtorwaalridasnaOrrcehviisdiRoensoefaBrucnho5c,hi4l5u-s1D1.1L..Jones coNmebiwnactoimobnisnatotiboenmsadaere. made in Caladenia Jon&esM.DA.LCl(e2m00.6Acu)stSrpaelciuanlaOnrtchhaidveRrensaelairsch(5O,rc1h1i2d-a1c4e2a.e), a R. Br. (from Arachnorchis D.L. Jones & M.A. ScroiuttichalWlayleensd.aTnhgeerOerdchnaedwiasnpe1c5i,es27f7r-o2m81s.outh-easternNew Clem,andStegostylaD.L. Jones &M.A. Clem.), JonesDL (2007) Twonewspecies ofOrchidaceaefromthe Corybas Salisb. (from Corysanthes R. Br. and Australian Capital Territory and southern New South Nematoceras Hook, f.) and Pterostylis R. Br. JoWnaelsesD.LTh(e20O0r8cah)adTiwaon1n5e,w32s2p-e3c2i6e.s ofHymenochilus (Or- (from Bunochilus D.L. Jones & M.A. Clem., chidaceae) from south-eastern Australia. The Orchadian Hymenochilus D.L. Jones 8cM.A. Clem., Oligo- Jon1e5s,4D9L2-4(92060.8b) Twelve newspeciesofOrchidaceaefrom chaetochilusSzlach. andSpeculanthaD.L. Jones south-easternAustralia. TheOrchadian 15,546-558. 8c M.A. Clem.) to accommodate these species. JonesDL(2008c)TwothreatenednewspeciesofSpeculantha These combinations are made so recognised (Orchidaceae) from south-eastern Australia. The Orchad- ian16,30-35. validnamescanbeincludedinchecklistsofflo- Jones DL (2009a) Three new species of Oligochaetochilus ra maintained by herbaria in Australia. Many (Orchidaceae) from South Australia. The Orchadian 16, ofthese new species are highly localised and Jon1e1s9-D12L6.(2009b) Five threatened newspecies ofHymeno- or threatened with extinction, and a validly chilus(Orchidaceae)fromsouthernAustralia.TheOrchad- uacncdeeprteSdtastceieantnidficConmammoenwmeaakletshlibsitoidnigveerassiiteyr WaoilfasnVhi1cN6t,oGr1i7aa6n8-d1eS8dt7a.j(sRiocyVal(B2o0t0a7n)iAcGceanrsduesnosfMtehelbvoasucrunlea)rplants protection legislation (Backhouse 2007). Pres- entation ofauthor names follows the standard usedinWalshand Stajsic (2007). Acknowledgements I am grateful to Peter Weston and Wayne Cherry, National Herbarium of New South Wales, and Received9July2009;accepted10September2009 Neville Walsh, National Herbarium ofVictoria, for commentsandcheckingthenewcombinations. Vol 127 (2) 2010 55 Contributions Newcombinations Caladeniaarmata (D.L. Jones) G.N. Backh.,comb. nov. Basionym: Arachnorchisarmata D.L. Jones,Australian OrchidResearch 5: 52 (2006) Caladenia atrovespa (D.L. Jones) G.N. Backh.,comb. nov. Basionym:Arachnorchisatrovespa D.L. Jones, The Orchadian 15: 546 (2008) Caladenia branwhitei(D.L. Jones) G.N. Backh., comb. nov. Basionym:Arachnorchis branwhiteiD.L. Jones,Australian OrchidResearch 5: 46 (2006) Caladenia cadyi (D.L. Jones) G.N. Backh., comb. nov. Basionym:ArachnorchiscadyiD.L. Jones,Australian OrchidResearch 5: 52 (2006) Caladenia moschata (D.L. Jones) G.N. Backh., comb. nov. Basionym:Stegostyla moschata D.L. Jones, The Orchadian 15: 555 (2008) Caladenia orestes (D.L. Jones) G.N. Backh., comb. nov. Basionym: Arachnorchis orestes D.L. Jones,Australian OrchidResearch 5: 48 (2006) Caladenia ustulata (D.L. Jones) G.N. Backh., comb. nov. Basionym: Stegostyla ustulata D.L. Jones, The Orchadian 15: 323 (2007) Caladenia whiteheadii (D.L. Jones) G.N. Backh.,comb. nov. Basionym:Arachnorchis whiteheadii D.L. Jones,Australian OrchidResearch 5: 64 (2006) Corybasgrumulus (D.L. Jones) G.N. Backh., comb. nov. Basionym: Corysanthesgrumula D.L. Jones, The Orchadian 15: 550 (2008) Corybassulcatus (M.A. Clem. &D.L. Jones) G.N. Backh., comb. nov. Basionym: Nematocerassulcatum M.A. Clem. & D.L. Jones, Telopea 11: 406 (2007) Pterostylisagrestis (D.L. Jones) G.N. Backh.,comb. nov. Basionym: HymenochilusagrestisD.L. Jones, The Orchadian 16: 176 (2009) Pterostylis barringtonensis (D.L. Jones) G.N. Backh., comb. nov. Basionym: Bunochilus barringtonensis D.L. Jones,Australian OrchidResearch 5: 121 (2006) Pterostylischocolatina (D.L. Jones) G.N. Backh., comb. nov. Basionym: Bunochiluschocolatinus D.L. Jones,Australian OrchidResearch5: 122 (2006) Pterostylis clivicola (D.L. Jones) G.N. Backh., comb. nov. Basionym: Hymenochilusclivicola D.L. Jones, The Orchadian 15: 492 (2008) Pterostylisconferta (D.L. Jones) G.N. Backh.,comb. nov. Basionym: Hymenochilus confertus D.L. Jones, The Orchadian 16: 178 (2009) Pterostyliscrassicaulis (D.L. Jones) G.N. Backh., comb. nov. Basionym: Hymenochilus crassicaulis D.L. Jones, The Orchadian 15: 493 (2008) Pterostylisexalla (D.L. Jones) G.N. Backh., comb. nov. Basionym: Oligochaetochilusexallus D.L. Jones, The Orchadian 16: 119 (2009) The VictorianNaturalist 56 Contributions Pterostylisferruginea (D.L. Jones) G.N. Backh., comb. nov. Basionym: OligochaetochilusferrugineusD.L. Jones, The Orchadian 16: 120 (2009) Pterostylisglyphida (D.L. Jones) G.N. Backh., comb. nov. Basionym: Speculanthaglyphida D.L. Jones, The Orchadian 16: 31 (2008) Pterostylis incognita (D.L. Jones) G.N. Backh., comb. nov. Basionym: Hymenochilus incognitusD.L. Jones, The Orchadian 16: 179 (2009) Pterostylislepida (D.L. Jones) G.N. Backh., comb. nov. Basionym: Oligochaetochilus lepidusD.L. Jones, The Orchadian 16: 122 (2009) Pterostylislineata (D.L. Jones) G.N. Backh., comb. nov. Basionym:BunochiluslineatusD.L. Jones,Australian OrchidResearch 5: 121 (2006) Pterostylismacrosepala (D.L. Jones) G.N. Backh., comb. nov. Basionym:Bunochilus macrosepalusD.L. Jones,Australian OrchidResearch 5: 125 (2006) Pterostylismajor(D.L. Jones) G.N. Backh., comb. nov. Basionym: BunochilusmajorD.L. Jones,Australian OrchidResearch 5: 115 (2006) as‘majus Pterostylis multiflora (D.L. Jones) G.N. Backh., comb. nov. Basionym: Speculantha multiflora D.L. Jones, The Orchadian 15: 554 (2008) Pterostylisparca (D.L. Jones) G.N. Backh., comb. nov. Basionym: BunochilusparcusD.L. Jones,Australian OrchidResearch 5: 123 (2006) Pterostylis rubescens (D.L. Jones) G.N. Backh., comb. nov. Basionym: Speculantha rubescensD.L. Jones, The Orchadian 15: 554 (2008) Pterostylisspissa (D.L. Jones) G.N. Backh., comb. nov. Basionym: HymenochilusspissusD.L. Jones, TheOrchadian 16: 182 (2009) Pterostylisstenosepala (D.L. Jones) G.N. Backh., comb. nov. Basionym: BunochilusstenosepalusD.L. Jones,Australian OrchidResearch 5: 130 (2006) Pterostylis tenuis (D.L. Jones) G.N. Backh., comb. nov. Basionym: Bunochilus tenuisD.L. Jones,Australian OrchidResearch 5: 127 (2006) Pterostylis umbrina (D.L. Jones) G.N. Backh., comb. nov. Basionym: Bunochilus umbrinusD.L. Jones,Australian OrchidResearch 5: 126 (2006) Pterostylis ventricosa (D.L. Jones) G.N. Backh., comb. nov. Basionym: Speculantha ventricosa D.L. Jones, The Orchadian 16: 30 (2008) Pterostylis vernalis (D.L. Jones) G.N. Backh., comb. nov. Basionym: Speculantha vernalis D.L. Jones, The Orchadian 15: 277 (2006) Vol 127 (2) 2010 57