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LLLLoooosssstttt CCCCrrrroooovppppGolrusssasmin es ooooI ffff AAAAffffrrrriiiiccccaaaa e oric forhorit om thpecifie aut mposed from XML files created frng styles, and other typesetting-snt version of this publication as th Board on Science aNnda tTioencahln Roleosgeya rfcohr CInotuenrncailtional Development en recos, headie the pri eks bau as brese n of the original work hnal; line lengths, word dentally inserted. Plea epresentatioe to the origive been acci s new digital rbreaks are truerrors may ha ut this PDF file: Thisetting files. Page some typographic NATIONAL ACADEMY PRESS Abotypeand Washington, D.C. 1996 Copyright © National Academy of Sciences. All rights reserved. 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AmwbNfsNDouyPacuierpt ta mtaEhpIird on io-RTTbretn5terieemhheoat5grr pieln4wsanys o ,A5r ar oarrdoea-tetcfsiAfp n op faR tfodSo-hdnouer 0rceearOnt v0ti mlad ewc if-wpDenefo8yapwidcam0e csroes v6 e obmC rsf e8ppye ,oEol -rivr otft0etimhn ithappR0eeegtemmawe.e ei Nr nB siereebteeeanddutaeasett rliprr,bebaeo icoy Oynnanchn uc gaof,aase f,ln nAf ii .gaobcAs rganierlosced ed Atuoa n i fhdtpnfcofh ore gyIroeicmn c ttof tIhhaaoyefn,eed rr sm Born vtrI ifaeitnuet shtpuEmtrioaeeootnrnerbarnyng tueaot aiphrw nlftfs ae ieoA eMon oerarnfe freuefR ail atdnt clheohi ighrDsecfos,oe i r,tNenaas shvNenre aeacen.dt achliB toofct iohaopoonnreamnra ddrtlaIdh ei nlADn n esoRgitctenr,i ea v tt ussdouSepene tlcpeaedomi crreepocoiynrmafhc c Glo eMe eCcfdnr oaoaeuSt,nmndurc edtOinip cse NcfiTe ninaftolecipee.c.c en.pSe hscTr t,nooae htovff heafleeondgdy e oric forhorit Library of Congress Catalog Card Number: 93-86876 mposed from XML files created from thng styles, and other typesetting-specifint version of this publication as the aut ITtCPhSrhoeBiinp sNNty edra dio0tg i-cioh3nunt0 m ta19hle9- eAn09 Ut46c 9m anb9diayt0eey -mtd 3hb yeSe toNraefta peStsrico ooidnefuan Alcc eAemdsc.e sarodicleeamlyy f oofr Sedcuiecnactieosn. aAl lpl urirgphotsse rse wseirthvoedu.t the written permission of en recos, headie the pri eks bau as brese n of the original work hnal; line lengths, word dentally inserted. Plea epresentatioe to the origive been acci s new digital rbreaks are truerrors may ha ut this PDF file: Thisetting files. Page some typographic Abotypeand Copyright © National Academy of Sciences. All rights reserved. Lost Crops of Africa: Volume I: Grains http://www.nap.edu/catalog/2305.html iii ald, ne gin he orie retai A NOTE FROM THE SPONSORS ginal paper book, not from tmatting, however, cannot bative version for attribution. hcAMtsidhuuafeuarrpsiacrnc pehs aFson. 'euorstot vpr fo efp ntrnwtrouoh eortmer,bt ei dt eoipdtoefhnoo nce vprta ehauisdn.erli eiatt oueCsutuiar,onln settndia hoae rtenrieirslo ypU eni,som .xaS rptlotdh .slcr oee Aeoit ft egrmevporeduimrmono lncrcu baeeyBnlter riiOfentamoyagbSr, l,lTseI bnn aI ueoDttnetoef 'd rsi dAtn m saatfIhr naltisiicntolsaonao nnu ,ivna sgtiarel si itbatDe iipolo deeansns ver etaaPrailcin oonruhodpudlog ams srssr ateltcuy amnoos rttfntv.i am (fcaAriTeneteiIhreosDlnyidhns).. e oric forhorit This book offers many such ideas and is part of a commitment AID om thpecifie aut mThaedree ,a t mtheem Inbteerr nactoiounnatrl iCeso,n fenroenngcoev oenrn Nmuetnrittaiol n (oIrCgNan) iizna Dtioencse,m baenrd 1 9t9h2e. mposed from XML files created frng styles, and other typesetting-snt version of this publication as th istrb"wtonihelmtopohtaidesotsieriuc rotkvd, nhcBFna cecaeh oystrociito or no ihatu fowphcnoidltngsdle,erhue a "h vewtd b.lhvleo sieeiguedreb cc,hl en v mrootmesieaimmonapsrnaugnatrmedi se nybo t luaah tiyflao pn dei tmtitchaiy nhtbuayea rietrnno n jsorowgdf pa rh eoos rldteceerhrfe nolldapedrpAunigso o fttdtepotrroureiiiidrnvct sboniu seaoluvyts, rnitt i aesootd,ahwli . re tatrf ye- oteenr of raioolds i amtAtfo khe mrdtf ieenhrnp ii eeccsaIeCpdraestooel e'cNespnw u al uaoeoriostitnw hbtrreytc i jonede ihesi n cnnufunttot a niobttvmo thg sicedveaethrsa eoilra.rsnn . p b vhTuntsAiieohtoaa rt rewdlti tltibyat viiisohvle geloene ee rp-r aencsswrsrdo a,iaeti uttytltmaehlhhec tsciieenedos,t en recos, headie the pri ppraogjeecst 'ws imll asjtoimr usplaoten smorusc. hW ien theorepset tahned w meaalnthy osfu ibnsfoerqmuaetniot na icnt ivthiteie fso.l loIf wthinagt eks occurs, the now largely overlooked resources described herein should bau as brese contribute substantially toward achieving the goal of eliminating hunger and n of the original work hnal; line lengths, word dentally inserted. 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Lost Crops of Africa: Volume I: Grains http://www.nap.edu/catalog/2305.html iv ald, ne gin he orie retai ginal paper book, not from tmatting, however, cannot bative version for attribution. sTohcei eNtayt ioofn dails Aticnagdueimshye do fs Schcioelnacress e ins gaa pgreivda itne, sncoinenprtoiffiict, asneldf -epnegrpineteueartiinngg om the oripecific fore authorit ruthesseee Cfaorocrn htgh, rede egsdse inicnear t1ae8ld 6w t3oe, l tfthaheree fA.u Urctaphdoeenrma tnhyce eh aaousft hasco mireiatnync deoa fat tneh dteh tcaehtc ahrrentqoeurlo igrgeryas n aittne tddo t taood tivht iebsiyer een recomposed from XML files created frks, heading styles, and other typesetting-sse the print version of this publication as th tATcosrEnaATheehcuehhnlcslehabeteeegap sried cf dioeIttNrentseeannvtide,msaornsse e eeitntidomybrrisin fioita oi nneo ucplnltfignf ogrht tag eiye tu Eetalosse s orflniA Nnv dsooamfgego cegrfaiMr e inanteasennmitdesdpnoe meeo evobdngeerfmieneiaiidr scnnntrssliuhyis gnoe.tAn,ce .gre oIoe sasct rNfn ht t asw ehiia Ea.dssone art Dcenii gnasofnmig iu ereengan.idtnys an neWoewt tleedao nriAriebfa mfotirirll hcnce mSii.ga sg nstcAahdoho egipneeveeua. rdmd e wnrWsoN r c ciytgnainaeheun smr otcs,al 1 i effi,hoema9 t snannSins7tssstaai cd0 . ac lbai pa T deabrAilr nmilmepheys cccames heaieto rinhedNdsdaagdei. etle aasn ltmnNittteeini rtlrzmoyaa so1eon tt.oeisi9far oefDog l6n ttt nSahAhi4rann. cee,la c g iiB AuanezNs dnnrnaducauedacta pitimcteedoineiesrooenyr tnmnttiM hhhoooaayeeefrfll. bau as brese of Sciences to secure the services of eminent members of appropriate n of the original work hnal; line lengths, word dentally inserted. Plea ptAfcIhnearcdesorta eefipdert,uua serltbsem eiglsdoiyoecincv a.os ierT nfcri ehnnhS.e m ,ct haIieenennn sdtet cx iaeetnaudsmdt ube,i c yunaa pacitottitissono n cuni.otn soDnd foge rwpr.r eoHtnhsl iasiecnir yorivte neimsaayptal ioV vtctne.eh s,Fra itsbiro ntpi elieeidrbtr eyttenao rgit gnibif vieyines agni psn sttr oouea e sdttsihhvd eeoies fhNne metra a oettliodftoh ittnc hhoaaeefll epresentatioe to the origive been acci StaTedhccveihie Nsnnioancltgeoi osgtnh yiean wl f 1Rei9dtehe1s re6taha ltre ogc Ah oav csCeasrodonuecmnmiceayintl’e tsw. tpFahsuue rno pbcrgotriaosoneanisdzi neo cgdfo fibmnuy r amtthchuceeo nrNriidnatyatgin o okcnfnea o lsw wAciiclteheand dgcgeeemen aeyarn noaddfl s new digital rbreaks are truerrors may ha poAanpocdelair dctahieteeimns s gydc eiaoetgfne eErtnimnfcigyicni noaeenfd ed bbr oieyntn hgtg h tiiehnn ee Ap eNrcroiaanvdtgiiedo mcinnoaygml, Astmehcrueav ndCiicetoemieuss ynt. o coT iftlhh Sheec a Cgiseo obnvuceenecrcsnoi mmla nieesdn ta tthd,h emteh piNenr ipainstuitcboeirlpnieaacdll, ut this PDF file: Thisetting files. Page some typographic jANwoawlibtniweotrl.nnytasa lbat Riyno ednbs aoDelat-rhra.c cWAha cCdmaeo.dmu Aenime.c siWil.e.o usr lgaf nadre t hceh aIinr satnitdu tvei coef cMhaeidr,i criensep.e cDtirv. eBlyru, coef tMhe. Abotypeand Copyright © National Academy of Sciences. All rights reserved. Lost Crops of Africa: Volume I: Grains http://www.nap.edu/catalog/2305.html v ald, ne gin he orie retai PANEL nal paper book, not from tatting, however, cannot bve version for attribution. NJGOLOHERTIRNNMnre dNiAsAgie aoXWaNn,rT a Mc.E EhBe L.S xULBeiRc,rO voTDiR cCOeLepiN,tAa yU,rU, t G.mMSGe.ee, on DxrtCig ecoeipoafna ,tA rCrCtomgoh raeaIosnnintrttaom elorm afPn nyaAlac, giiPnoru incEraudxllu ptdeuee rrUe im,Mn Tieevnijfetotr orsSanittm,ay tG,ii eoWennot,ero gsA tid agLer aicMfauylateíuztrt aeyl, gimati JACK R. HARLAN, Department of Agronomy, University of Illinois (retired), e oric forhorit New Orleans, Louisiana ed from thng-specifias the aut KENNETH O. RACHIE, Winrock Inte*r*n*ational (retired), Pensacola, Florida mposed from XML files creatng styles, and other typesettint version of this publication NSFM.TORAAE.R TSFRLKceFU ice DRhSnnK.t. i oDfIilVNocA Ig,EFE yBdFT fOiOMotoSrRr ETINnIYDt,e ES rERtnada,f itfti SooAernsnasiolo cDri aePvteerloogprmamen tO, Afffirciecra, CBroopasrd S tuodny SDciireencctoe r aanndd coadipri ELIZABETH MOUZON, Senior Secretary een reks, hese the BDROENNATL DSI MOSPBSOORNN, M, MUUCICAI AIn Itnetrenrn bau as brese MICHAEL McD. Dow, Acting Director, BOSTID ginal work hngths, word serted. Plea n of the orinal; line ledentally in epresentatioe to the origive been acci s new digital rbreaks are truerrors may ha hie c ut this PDF file: Tsetting files. Pagsome typographi Abotypeand Copyright © National Academy of Sciences. All rights reserved. Lost Crops of Africa: Volume I: Grains http://www.nap.edu/catalog/2305.html CONTRIBUTORS vi ald, ne gin ginal paper book, not from the orimatting, however, cannot be retaiative version for attribution. ogbefr avitenhrBsea, gy le of1 rsp9utl9 iat3cnsr,,t o smp nwosu roteosr t,ft hh yAav neof grf1i ec,i0tnaa0c.b C0lluM epssoieo,o osnntpl .el thIegnra uh dmaia bdsep espaun,ar strtieotcio,ici lapisrlpaela tsetethdedesd ,s beiny sp pBeniocOoepmSsle,Ti nwIsaDweti'rsnee ego ct veosennpretaerrlcislbi ,se utsuta ondordysf e oric forhorit to this, the first product from the study. However, the following list includes only mposed from XML files created from thng styles, and other typesetting-specifint version of this publication as the aut tcwSOOhhA.eLoaC MsUpa.eNNr tP eUeAiirOw ggEtWsree MhLuorroOlii fIy aa A Rt AphgGHrJirosaAB, v tpDeOYiadfeIIureR,ptld i.IacE Crut,tR elmaNcIrehS anbnAttioi ocTooan-fklaW .Fl dToACeootSe AdaarIi ePFllTlsa, R e ltKshctI hheCaRa nncAteoo osl,bnoe Netagrcryiicagb,hm euU rteIiona nrissivnt, eicbtruoostritepthy,o lrBoiasfitt deeIaddb, a aidNnnatidnog , eu rIvn balSairstditaoaetundes,,, en recos, headie the pri FJAOCROBSuBOn NsAo K., AG. AYhaYUnEKaN-TSAUK, PElMan,t IGnestnietutitc dRee lsao uRreccehs eUrcnhite, CArgorposn oRmesieqaurec,h Y Inasotuitnudteé,, eks bau as brese Cameroon n of the original work hnal; line lengths, word dentally inserted. Plea RPMJSAATO.CAEUBQPRH.LE HeUa RBBgrEEaTiEENorS eNCn C,CB, KH ZUGAEMUiDmhYRRaAJObRnAONaa,, b,E,IZ wn EoDseAdmteieZtbpunIaat AF,r d tMoWmeu aeROnlndeaRtacw ,th iioeoGfrn cr,ah BZienii oAnsc dghDereroem, nvNioesimltgoreyipqr,m ueeU,n nMt ivBaerorosauirtady,, CoHfa moZheoirmoe,ob naVbowltea, epresentatioe to the origive been acci CAABREIRBLnes Esp WoDeucE.rt ciMoeCnsIA SCSSSeeTIrnEvLtri,Ec ePe/TEl,a OtUnhNt.i SoG, .p eDiIranme,t epApralndraatdsmtimisoe AnnEatbx loap fblS oAaer,a cgEttriiitoochnnuio ,la tpunAirdaen C,i mAolbaleli dcjtaainondn, ,I vPPollaarnyn ttC GoHeaensaetlttihc s new digital rbreaks are truerrors may ha STUEWSUUAOnnNLii vvDeeBErrssU iiBttRyyEN,, RAAEHddLddALiiNss AA EGDbb/aaEWbbGaaA,,Z REEIAttDhhBSiiooH, ppEiiTaaRh,e ThNea tNioantiaoln aHl eHrberabriaurmiu,m ,A Addddisi s AAbbaabbaa ut this PDF file: Thisetting files. Page some typographic SWAAHLDERTxe EpNveeR.rl oiFmpFOmeRMnOetBn LStA ItAaC,t sHiMsoo,na c,nS iCgaotrrrioogovphnesu, mFRSrwe esaaeenmtaodrpw c hMnR ,Ii icSnllesiee ttRir trueaSts eeLec,a etTriocoanhnme ,S atNlaety,i oaGnnhk,a pWnaaleas t AAgfrriiccua lRtuircael Abotypeand Copyright © National Academy of Sciences. All rights reserved. Lost Crops of Africa: Volume I: Grains http://www.nap.edu/catalog/2305.html CONTRIBUTORS vii ald, ne gin he orie retai KIFLMEi niGstOryZ oEfG AUgZriEc,u ltRuereg,i oAndadl isS Aoibla baan, dE thWioapteiar Conservation Department, ginal paper book, not from tmatting, however, cannot bative version for attribution. SLITS.EACRLN.A IZCTATnEiarNtGImofLeGaDprUb wnEAsa Pa GabRRFtT-iwAN.eBoA senMH ae,Ka laOel rE JwRCcUIRhDaerS oEgIEUEpniDAon,ss En tiNRRiva teuIeelK, rgst seASeii SoactfSynroohcS,arro h gABlot hhIla,Sn u euooZ smcSfrto hige Stnihum,ca uatiNleienm - niAdTfg oc aerereniar d Mtdimaha nTe iMdilr nlSo eSi epltSclsmie coietisis-nl,I A cmBICemrupio dElprna odrTswovuervearcovmeyapamotieitc,oine soZnnt,n ,itB m ,A PPuBbbrrlaaooaubwhggbwrrraadaakyemmeaorr,,,, e oric forhorit Ethiopia om thpecifie aut HILDNAjo roK, IKGeUnTyaHA, Department of Home Economics, Egerton University, en recomposed from XML files created frs, heading styles, and other typesetting-se the print version of this publication as th AJHKPJGO.l .UBEuM YLsEEA AAACPICMBYbNMlrEUbgaaEEoUAardt ID poe LbCKTaNMsaA aOnH K uID,R,cRA W AEOSNt KeUiMttsoEKREihRaeBgnLAt,iDUaEe eo ,D r- rD,YMpIIcF iKENneaihOSLaApfNY iAoIAUagRnSErrePS,mstI, Srm,t ,EiW i aMaMPtAKetuieenUrtoa,asek tsn tr, leo fl o ASArofDyu MregBfd, err tr'vpLoihisiclcia teleFcaaeur senSntotlom st yeotu,MPhm deraIron eninoiLc t-sdg Aart roiobaMtarbfuonim ditrdioecaB , Tl rtooooorSSgftorbaa yiopAihnco,ii aeyglclD lol,sr i iog,aRc CUuyNnuea, onlis ltanUCieauvmsa,erne eraCneircrlvstyvh areiR,ame t rytCN,esi e soieirtIeognynonafe, totr orercneJfrh, nGo I,asC beA,ta rieomddnJnadotarnaisnesl,, eks bau National de Recherches Agronomiques, Bambey, Senegal n of the original work has nal; line lengths, word bredentally inserted. Please FPMG.EECARB.TJHSANR.Et udAeUOadlMNl miSesTDAn bAMbEOoRobK,EE saAAcEGbAhna, I. ,g,SB M SoEOD.tl tEea,eh lMHpilSoeaEoAnprRtibiRmlao ECCesTcnEoAhtLn ,, sI CoeNNrafOv paae,tAt u iPorgIarnnrolo s vntDiRiotneuemcptseoayo, r utSmrdoaceneue ndsth It ,n DAPvMefearpssiinttaciuriagsrtametrçsyeã, noot U,f AnMAigvgrienorriinscsóiuttrmylyt u icrooaeff,, epresentatioe to the origive been acci IK.MOU. ASAMtgaMrtHiicoÉAun lM,Rt uCAIrhEePi,ZrA AeARddNzAdi,,i, sW Z ARiemebsstba eAbaabafr,wrc iEhce tah aRinoidpc iea SDpeevceilaolipsmt eSnet rAviscseosc,i aCtiohnir,e Bdzoiu aRkee,s eIvaorcrhy s new digital rbreaks are truerrors may ha GHEELTCAEAogNCar HiscMtEuloWtuss rB,e ,EI AnYtdeEdrNnisaE tAi MobnaIaSbla K,P EElaRtnh,t iC oGpoimeanmetuicn itRye Fsooruersctersy DInesptiatruttme,e nUt,n Mivienrissittryy ooff ut this PDF file: Thisetting files. Page some typographic Zimbabwe, Harare, Zimbabwe Abotypeand Copyright © National Academy of Sciences. All rights reserved. Lost Crops of Africa: Volume I: Grains http://www.nap.edu/catalog/2305.html CONTRIBUTORS viii ald, ne gin he orie retai S.C. ENcAonNoAm-iSc ICNoKmAmMis,s iJoonin fto rE ACfAri/cFaA, AOd Adigsr Aicbualtbuar,e EDthiivoipsiioan, United Nations ginal paper book, not from tmatting, however, cannot bative version for attribution. JNGAN.CO.LR .AERS BTPGNHMOreueODAcBrAthuoRUKINnnrEY iOod aFNli F,OoSU EP.gARGN ryNeX,.A, t SoOORT, rAhwENiaDeKTe,, erUTHSrDpeio,Ma, rt ureyDItAtm,lmh iaMNr oneeA d,ncoS f ttzrDti iaaoFcetomnaaepf r,b amd NireCqtisimnurg goeSee prnei Rrta vPeoisrcfeoe adsPr ulcGachnté iton anéPnr,dr ao uFdDxeu decTevteireoaclnloh ,pn UmiUqneunineivvtse ,e rPrsGsiritoitytyje e gcooatff,, e oric forhorit Ministry of Agriculture, Katumani, Machakos, Kenya om thpecifie aut P. SOInMstAituNte, Pfoear rtlh Me Silelemt iP-Arorgidra Tmr,o Spaichse,l iNania Cmeenyt,r eN, iIgnetrernational Crops Research en recomposed from XML files created frs, heading styles, and other typesetting-se the print version of this publication as th PJJJAGJ.EOA.HD.SKNHN..E .SIPNRANES WBnW rT egi seATaVsIErtRtEmeiLiOoCtYOc.aBurNLeuLHriNNtEycaIle.LOtA h,,R ,,u , TN SD PMrB,II eAonriInWiA,geSunvs tYAtteettiR,Aoi hesrrLtdriGPrnu ToiOAndateaEnCeaita,Rfsi ,tNrHro So i,iTlAo fcn, oI naDaSMaFbumaNlOatoe lhibC poayl,I la laAdanere,ron tsC ,fStE tpkmrGioPcitsptcihmuer haeaoRintalnomegatnecp rosuaeaioeAfan mfa aiT gnrt,Fycrr doih oS cTp oauFIiedhnlcoct esaurhSltelriin scaatAuotinlre tgl yeonrEC gicfcxeeyDoupn,,r e let NtUrptruheiaarnm,erte i it,IveomS nInenebtertame asnlrid tSintF,-ay atAona tMoit,rooi idNofdni nnR aiPTig,l ser ertesCCorretyiporraa ooirrcicppoashssf,, eks bau Research Institute for the Semi-Arid Tropics, Niamey, Niger as brese n of the original work hnal; line lengths, word dentally inserted. Plea DDJAAJICVBQLRIiDUnILc Eo JSA.l nWBA, NAN, ReRDbeRRrsaAiEsdkUWean,,S t UL,M aSDibAseospirOoaantrTot,m HiSreeiEl nvRdte ' ErRo tSfhE pnAGroignIbOrgoo,tN naMnoSimaqruyye,l a-UBndino,i gvUéeSorsAgirtyap hoife ,N Mebursaésukma, epresentatioe to the origive been acci DJ.PO.N ANBAagALtrioUDonnD oamOFl .dIi qN'HBu, ieEssPtE odhCieyrH etGo ,Nt eemacCthubonrlmoeieluml xeod,, nePGswae remiRasb,lé tlgFhoir uoaSxnn,scc iBeeCenlhtgiafiiuucdm easn, dF aIcnudlutés trdiaels RSecsieenacrcehs s new digital rbreaks are truerrors may ha GPAILULODLEréAgSv a eBnlBoiRzEpaApZteiMAomnNEe, LÇnBt-O rCeiNsnOb ,CXa noI, neoD,sp tQeéitpruuaaettr ietoFmnnsre aldannentç daMo,i fsAo Anudtgseptr eroalRlnliieoeacrmh, yeMr, coKhneat pneSslaclsiie eSrn,tt aiFftrieqa nuUcene ipvoeursri tyle, ut this PDF file: Thisetting files. Page some typographic Manhattan, Kansas, USA Abotypeand Copyright © National Academy of Sciences. All rights reserved. Lost Crops of Africa: Volume I: Grains http://www.nap.edu/catalog/2305.html CONTRIBUTORS ix ald, ne gin he orie retai FRRICAHNAKR BDR ELY. EBRR,U BGluGdEenRzS, ,V Ionrtaerrlnbaetrigo,n Aalu sPtrrioagrams Research Section, U.S. ginal paper book, not from tmatting, however, cannot bative version for attribution. LWGWYE.A ONYDDCLEFNNniEoeFvEgplREReo lara ErErpCanBtKYomddARoo eRlYP,Cn, UL.DEtL SCSnAOoOAHgYfNlN AaT,An ,P OdMgMDNraieAc,sp ukNaRlart,lotu mWyFreaoe,yln r eatDB gCeeoosnotf,avl lnIeeniSrgcc o e.W,c,G iCUoaiallrndolddilgeviwyfnee,esr s,l liRU,tK yeIn dseoieawvfahe ,ro Lrcs,Roh iUtn iycdSC hoAemonnf,ot enWrdL, y,i veDSe, eurKpnrroveeoneyrlt,,,, e oric forhorit England om thpecifie aut MAXN eDb.r aCsLkaE, GUGS,A Department of Agronomy, University of Nebraska, Lincoln, mposed from XML files created frng styles, and other typesetting-snt version of this publication as th EWRRJOLOOIHILNBZBUGSNALENAeeIeenNRmAYBrowikNvTr MEi ege-D ERTlAriP easSeHC.yri. l itEh,R dyMD ECxiE I,TAAUp OT.lIM ernmGCCNLord.ipeHSLsC mitriaEOEeLAcreirNsSNEtnNd,oF GtYa,A,, I m SDEC,En tD,LeCaadL AptlehSDeiipaofr,nm oa,rant tI rrrsm,PInnte tnmGetr ieetaafrnererodd,irn ntfraeU naf toms aiDtShfnito ,oAi,e, A o fTnvIG nnnaheAaedtlleh looi g ParNproCrglmoeapirntanooeh,otnlp e oUmtsrG g lSCyayeRA,nno, edeoUUsstpeinnceaii rrvvacReethrier osssiInoitnty uysS r toceifoteru vfsCt iecaIG enlifsesfo,oto rirrFt guntritiheaeaee,,, en recos, headie the pri CONURnAivDe rsLit.y , ESVtilAlwNaSte, r,O Offkilcaeh oomf a,I nUteSrAnational Programs, Oklahoma State eks bau CHARLES A. 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Page some typographic WNAAZYDLMNoenUpEdaL rot.nW mH, e.LA noQtH no,dA fIo nNAnt,Nge ErrAninca,ug tillotaGunneradeol, r CgTieiafn totrCne, o fGaosret oaUrl ngdiPael,r aU-iUnS tAilEizxepde rCimroepnst, KSitnagti'so nC, olUle.gSe. Abotypeand Copyright © National Academy of Sciences. All rights reserved.