Journal of the Bombay Natural History Society, 107(3), Sep-Dec 2010 213-219 FLORA OF SANDY COAST OF GANJAM DISTRICT, ORISSA, INDIA D. Sahu1'2 and M.K. Misra1,3 'Ecology & Floristic Laboratory, Post Graduate Department of Botany, Berhampur University, Bhanja Bihar 760 007, Orissa, India. "Email: [email protected] 3Email: [email protected] This paper deals with the systematic account of plants from the sandy coast of Ganjam district of Orissa, and reports 175 species of angiosperms under 134 genera belonging to 61 families. The specimens were deposited in the Herbarium of the P.G. Department of Botany, Berhampur University, Berhampur (BOTB). Poaceae was the dominant family followed by Euphorbiaceae, Cyperaceae, Fabaceae and Asteraceae. Families were arranged according to the modified Bentham and Hooker’s system of classification. The native species are represented by 155 species (89%), whereas the exotic species are represented by 20 species (11%), of which 16 species (9% of the total) are naturalized in the area. Ceropegia candelabrum - a species hitherto reported from inland is now reported from the coastal area. Bulbostylis subspinescens and Micrococca mercurialis are reported after 85 years of their first collection from the coast. Key words: strand flora, angiosperms, native and exotic species, Ganjam district, Orissa, sandy coast INTRODUCTION nesting. The district experiences a climate when near the sea with an active south-west monsoon and the average annual Orissa has a coastline of 481 km and is rich in strand rainfall at Gopalpur is 1,296 mm. Relative humidity is high flora and forests. Coastal Orissa harbours littoral and tidal (64-86%) throughout the year. forests (Champion and Seth 1968). However, these forests Ganjam coast is famous for the Kewda Pandanus have been degraded to a large extent due to various biotic fciscicularis used in perfume industry. There are about interferences. Ganjam, one of the six coastal districts of Orissa, 120 distillation units (bhatties) in the coastal area that extract bordering Andhra Pradesh on the south, is represented by kewda essence from the male flowers (Sahu and Misra 2007). sandy coasts and devoid of tidal forests. The strand flora of Moreover, Indian Rare Earth (IRE) Limited and Gopalpur Ganjam coast has not been studied in detail even though some port are situated in the Ganjam coast. sporadic reports on it are available (Rao 1971; Brahmam and An extensive floristic survey of the sandy coast of Saxena 1980; Subudhi et al. 2002). The flora of the Ganjam district was conducted during 2007-2009. The plants presidency of madras (Gamble 1915-1936) refers to the plants were collected from different localities along the coast during of the district, but Haines (1921-1925) and Mooney (1950) different seasons. Specimens collected were dried and did not include the plants of the district in their floras. preserved with saturated mercuric chloride solution in ethyl Brahmam and Saxena (1980) mentioned some sand dune alcohol (Jain and Rao 1977). The specimens were identified plants of Ganjam coast in their Ganjam flora. The present with the help of the local floras (Haines 1921-1925; Gamble paper covers the sandy coastal flora of Ganjam district in 1915-1936; Saxena and Brahmam 1994-1996) and the Orissa. voucher specimens were deposited in the Herbarium of the Department of Botany, Berhampur University (BOTB), MATERIAL AND METHODS Orissa, India. The field number cited after the scientific names of the plants are of M.K. Misra, D. Sahu and R.C. Sahoo. Ganjam district lies between 18° 58' to 20° 17' N and The specimens collected are arranged in a tabular form 84° 06’ to 85° IT E. The coast of Ganjam is bounded by indicating the family, flowering period and growth form. The Srikakulam district of Andhra Pradesh on the south and Puri families are arranged according to modified Bentham and district of Orissa on the north; the coast line runs over 64 km Hooker’s system (1862-1883) of classification and undereach long along the Bay of Bengal (Fig. 1). The Chilika lake, the family the species are arranged alphabetically. largest brackish water lagoon of Asia is located on the extreme north-east of the district. Rushikulya, the biggest river of RESULTS Ganjam, discharges its waters into the Bay of Bengal and the estuarine zone is famous for the annual visit of thousands of The present study reports a total number of 175 species Olive Ridley Lepidochelys olivacea turtles to the shore for of angiosperms from the sandy coast of Ganjam district, Orissa FLORA OF SANDY COAST OF GAN JAM Table 1: Plant species found in the strand flora of Ganjam district, Orissa, India Family Name of the taxa Flowering period Growth form Annonaceae Annona squamosa L.; 573 Mar-May T Polyalthia suberosa (Roxb.) Thw.; 475 Apr.-May T Menispermaceae Tinospora cordifolia (Willd.) Hook.f. & Thoms.; 587 Aug.-Dec. L Capparaceae Cleome viscosa L.; 578 May-Oct. H Violaceae Hybanthus enneaspermus (L.) F.v.Muell.; 533 Throughout year H Caryophyllaceae Polycarpaea corymbosa (L.) Lam.; 546 Nov.-Feb. H Portulacaceae Portulaca quadrifida L.; 1508 Most part of year H Malvaceae Abutilon indicum (L.) Sweet subsp. indicum\ 597 Jul.-Apr. S Sida acuta Burm.f.; 515 Sep.-May H Sida cordata (Burm.f.) Borssum; 1534 Jul.-Dee. H Sida cordifolia L.; 70 Aug.-Dec. H Sterculiaceae Waltheria indica L. var. indica', 509 Aug.-Jan. H Tiliaceae Grewia tilifolia Vahl; 1516 Apr.-Jun. T Triumfetta rhomboidea Jacq.; 1075 Jul.-Jan. H Zygophyllaceae Tribulus terrestris L.; 1011 Throughout year H Oxalidaceae Oxalis corniculata L.; 173 Throughout year H Rutaceae Toddalia asiatica (L.) Lam.; 600 Aug.-Apr. S Meliaceae Azadirachta indica A.Juss.; 572 Feb.-May T Celastraceae Cassine glauca (Rottb.) Kuntze; 271 Sep.-Dec. T Rhamnaceae Ziziphus mauritiana Lam.; 594 Aug.-Oct. T Ziziphus oenoplia (L.) Mill.; 1515 Jun.-Sep. S Vitaceae Cissus quadrangula L.; 53 Apr.-Jan. S Cissus vitiginea L.; 376 Jul.-Oct. L Sapindaceae Cardiospermum halicacabum L.; 113 Apr.-Nov. H Anacardiaceae Anacardium occidental L.\ 1025 Feb.-Apr. T Lannea coromandelica (Houtt.) Merr.; 177 Mar.-Apr. T Caesalpiniaceae Caesalpinia bonduc(L.) Roxb.; 125 Aug.-Oct. L Cassia occidentalis L.; 542 Aug.-Nov. S Cassia tora L.; 62 Sep.-Nov. H Tamarindus indica L.; 98 Apr.-Jun. T Fabaceae Abrus precatorius L.; 218 Aug.-Sept. S Alysicarpus vaginalis (L.) DC.; 13 Sep.-Jan. H Atylosia scarabaeoides (L.) Benth.; 279 Aug.-Jan. H Crotalaria umbel lata Wight ex Wight & Arn.; 498 Oct.-Dec. H Crotalaria pallida Ait.; 1062 Jul.-Apr. H Desmodium biarticulatum (L.) F.v. Muell.; 513 Aug.-Nov. H Desmodium triflorum (L.) DC.; 251 Most part of year H Tephrosia purpurea (L.) Pers. var. maritima Haines; 34 Most part of year H Tephrosia villosa (L.) Pers.; 310 Jul.-Apr. H Vigna trilobata (L.) Verde.; 585 Sep.-Feb. H Zornia diphylla (L.) Pers.; 1034 Aug.-Oct. H Mimosaceae Acacia auriculiformis A.Cunn. ex Benth.; 122 Throughout year T Myrtaceae Syzygium cumini (L.) Skeels; 570 Mar.-Apr. T Lythraceae Ammannia baccifera L.; 270 Sep.-Nov. H Onagraceae Ludwigia perennis L.; 577 Oct.-Nov. H Passifloraceae Passiflora foetida L.; 1536 Aug.-Nov. H Cucurbitaceae Coccinia grandis (L.) Voigt; 06 Most part of year H Cactaceae Opuntia stricta (Haw.) Haw.; 200 Jun.-Sep. S Molluginaceae Glinus lotoides L.; 1537 Feb.-May H Glinus oppositifolius (L.) A.DC.; 326 Mar.-Oct. H Mollugo pentaphylla L.; 12 Throughout year H Apiaceae Centella asiatica (L.) Urban; 599 Most part of year H Rubiaceae Hedyotis graminifolia L. f. subsp. arenaria (Haines) Deb & Dutta; 1047 Aug.-Nov. H Hedyotis herbacea L.; 508 Aug.-Jan. H 214 J. Bombay Nat. Hist. Soc., 107 (3), Sep-Dec 2010 FLORA OF SANDY COAST OF GANJAM Table 1: Plant species found in the strand flora of Ganjam district, Orissa, India (contd.) Family Name of the taxa Flowering period Growth form Hedyotis puberula (G.Don) Am.; 74 Sep.-Mar. H Hydrophylax maritima L. f.; 1066 Feb.-Nov. H Morinda pubescens Sm.; 1501 Apr.-Jul. T Pavetta crassicaulis Bremek.; 539 Jun.-Aug. S Spermacoce articularis L. f.; 521 Jul.-Oct. H Spermacoce hispida L.; 1018 Jul.-Dee. H Asteraceae Chromolaena odorata (L.) R. King & H.Robins.; 528 Oct.-Dec. S Eclipta prostrata (L.) L.; 112 Aug.-Apr. H Emilia sonchifolia (L.) DC.; 131 Aug.-Apr. H Grangea maderaspatana (L.) Poir.; 107 Jan.-Apr. H Launaea sarmentosa (Willd.) Schult-Bip. ex Kuntze; 04 Mar.-Nov. H Mikania micrantha Kunth; 590 Jan.-Mar. L Parthenium hysterophorus L.; 102 Oct.-Apr. H Tridax procumbens L.; 1058 Throughout year H Vernonia cinerea (L.) Less.; 1053 Most part of year H Primulaceae Anagatiis arvensls L.; 304 Jun.-Mar. H Sapotaceae Manilkara zapota (L.) P.Royen; 541 Apr.-Jul. T Salvadoraceae Azima tetracantha Lam.; 305 Mar.-May S Apocynaceae Carissa carandas L.; 591 Mar.-Apr. S Catharanthus roseus (L.) G Don; 536 Throughout year H Holarrhena pubescens (Buch.-Ham.) Wall.ex G.Don; 118 May-Jul. T Ichnocarpus frutescens (L.) R.Br.; 105 Sep.-Dec. L Asclepiadaceae Calotropis gigantea R.Br.; 239 Dec.-Jul. S Ceropegia candelabrum L; 1519 Aug.-Oct. L Hemidesmus indicus (L.) R.Br.; 564 Aug.-Oct. L Pergularia daemia (Forssk.) Chiov.; 59 Aug.-Dec. L Tylophora indica (Burm.f.) Merr.; 1026 Aug.-Oct. L Boraginaceae Heliotropium indicum L.; 154 Most part of year H Convolvulaceae Evolvulus alsinoides (L.) L.; 209 Jul.-Feb. H Evolvulus nummularius (L.) L.; 1056 Jul.-Nov. H Ipomoea campanuiata L.; 158 Nov.-Feb. L Ipomoea pes-caprae (L.) R.Br.; 09 Most part of year L Merremia tridentata (L.) Hall. f. subsp. tridentata; 151 Feb.-Nov. H Solanaceae Physalis minima L.; 1502 Aug.-Jan. H Solanum trilobatum L.; 88 Throughout year L Solanum viarum Dunal; 1006 Most part of year H Solanum virginianum L.; 1005 Throughout year H Scrophulariaceae Lindernia anagallis (Burm. f.) Pennell; 324 May-Dec. H Lindernia caespltosa (Bl.) Panig.; 93 Aug.-Dec. H Lindernia ciliata (Colsm.) Pennell; 99 Sep.-Nov. H Pedaliaceae Pedalium mu rex L.; 03 Aug.-Nov. H Acanthaceae Asystasia gangetica (L.) T. Anders.; 374 Oct.-Dec. H Verbenaceae Clerodendrum inerme (L.) Gaertn.; 484 Most part of year S Lantana camara L. var. aculeata (L.) Mold.; 545 Throughout year S Phyla nodiflora (L.) Greene; 596 Most part of year H Lamiaceae Geniosporum tenuiflorum (L.) Merr.; 391 Jun.-Oct. H Hyptis suaveolens (L.) Poit.; 56 Sep.-Jan. H Leucas lanata Benth.; 463 Oct.-Dec. H Ocimum basilicum L.; 54 Sep.-Jan. H Nyctaginaceae Boerhavia diffusa L.; 10 Most part of year H Amaranthaceae Achyranthes aspera L.; 275 Oct.-Feb. H Aerva lanata (L.) Juss. ex Sch.; 103 Aug.-Jan. H Allmania nodiflora (L.) R.Br. ex Wight; 34 Jul-Dec. H Alternanthera sessilis (L.) R.Br. ex DC.; 84 Jul.-Jan. H Amaranthus viridis L.; 1003 Throughout year H J. Bombay Nat. Hist. Soc., 107 (3), Sep-Dec 2010 215 FLORA OF SANDY COAST OF GANJAM Table 1: Plant species found in the strand flora of Ganjam district, Orissa, India (contd.) Family Name of the taxa Flowering period Growth form Chenopodiaceae Suaeda maritima (L.) Dumort; 393 Apr.-Dec. H Polygonaceae Antigonon leptopus Hook. & Arn.; 24 Throughout year L Aristolochiaceae Aristolochia indica L.; 17 Jul.-Oct. L Lauraceae Cassytha filiformis L.; 08 Oct.-Nov. H Euphorbiaceae Acalypha indica L.; 47 Jul.-Dec. H Breynia vitis-idaea (Burm.f.) Fischer; 41 Mar.-Feb. S Croton bonplandianus Baill.; 64 Throughout year H Euphorbia hirta L.; 276 Throughout year H Euphorbia rosea Retz.; 1015 Feb.-Oct. H Euphorbia tirucalliL.; 520 Jul.-Oct. S Jatropha curcas L ; 83 May-Oct. S Jatropha gossypi folia L.; 79 Jul.-Oct. S Micrococca mercurialis (L.) Benth.; 35 Aug.-Sep. H Phyllanthus fraternus Webster; 501 Apr.-Jan. H Phyllanthus reticulatus Poir.; 512 Most part of year S Phyllanthus rotundifolius Klein ex Willd.; 1016 May-Oct. H Phyllanthus urinaria L.; 370 Jul.-Oct. H Phyllanthus virgatus Forst. f.; 336 Jul.-Feb. H Ricinus communis L.; 181 Most part of year S Sebastiania chamaelea (L.) Mull.-Arg.; 581 Most part of year H Moraceae Ficus benghalensis L. var. benghalensis-, 72 Apr.-Jun. T Ficus religiosa L.; 80 Jul.-Oct. T Streblus asper Lour.; 372 Mar.-Apr. T Casuarinaceae Casuarina equisetifolia L.; 1031 Apr.-May T Agavaceae Agave americana L.; 534 Apr.-May S Liliaceae Asparagus racemosus Willd.; 1527 Sep.-Oct. H Gloriosa superba L.; 51 Sep.-Nov. H Commelinaceae Commelina benghalensis L.; 196 Jul.-Jan. H Arecaceae Borassus flabelliferL.; 15 Mar.-May T Cocos nucifera L.; 46 Throughout year T Phoenix sylvestris (L.) Roxb.; 332 May-Oct. T Pandanaceae Pandanus fascicularis Lam.; 116 Jul.-Sep. S Typhaceae Typha angustata Bory & Chaub.; 1067 Most part of year H Cyperaceae Bulbostylis barbata (Rottb.) C.B.CI.; 124 Jul.-Nov. G Bulbostylis subspinescens C.B.CI.; 465 Jul.-Dec. G Cyperus arenarius Retz.; 20 Jul.-May G Cyperus compressus L.; 40 Jul.-Dec. G Cyperus iria L.; 43 Aug.-Jan. G Cyperus paniceus (Rottb.) Boeck.; 390 Jul.-Aug. G Cyperus polystachyos Rottb.; 29 Feb.-Oct. G Cyperus rotundus L. subsp. rotundus\ 50 Jul.-Dec. G Cyperus stolon iferus Retz.; 52 Oct.-Feb. G Cyperus triceps Endl.; 55 May.-Dec. G Eleocharis atropurpurea (Retz.) Presl.; 58 Aug.-Dec. G Fimbristylis acuminata Vahl; 583 Jul.-Nov. G Fimbristylis aestivalis (Retz.) Vahl; 548 Apr.-May G Fimbristylis ferruginea (L.) Vahl; 592 Apr.-Oct. G Fimbristylis ovata (Burm.f.) Kern; 575 Jun.-Dec. G Poaceae Aristida setacea Retz.; 164 Aug.-Feb. G Arundo donax L.; 1007 Oct.-Dec. G Bambusa arundinacea (Retz.) Willd.; 266 ? T Bothriochloa bladhii (Retz.)S.T. Blake; 273 Aug.-Feb. G Brachiaria distachya (L.) Stapf; 19 Sep.-Dec. G Chloris barbata Sw.; 119 Sep.-Jan. G Cynodon dactylon (L.) Pers.; 91 Most part of year G 216 J. Bombay Nat. Hist. Soc., 107 (3), Sep-Dec 2010 FLORA OF SANDY COAST OF GANJAM Table 1: Plant species found in the strand flora of Ganjam district, Orissa, India (contd.) Family Name of the taxa Flowering period Growth form Dactyloctenium aegyptium (L.) P. Beauv.; 42 Sep.-Mar. G Digitaria ciliaris (Retz.) Koeler; 27 Jun.-Apr. G Digitaria longiflora (Retz.) Pers.; 156 Jul.-Dec. G Eragrostis ciliaris (L.) R.Br.; 162 Most part of year G Eragrostis coarctata Stapf; 38 Sep.-Apr. G Eragrostis japonica (Thunb.) Trin; 148 Oct.-Jan. G Eragrostis tremula (Lam.) Hochst. ex Steud.; 23 Jul.-Dec. G Oplismenus burmannii (Retz.) P. Beauv.; 215 Oct.-Nov. G Perotis indica (L.) Kuntze; 1514 Aug.-Nov. G Spinifex littoreus (Burm.f.) Merr.; 1080 Sep.-Feb. G Sporobolus indicus (L.) R.Br. var. diander(Retz.) Jovet & Guedes; 26 Jun.-Feb. G G-Grass, H-Herb, L- Liana, S-Shrub, T-Tree, ?-Not observed (Table 1). These taxa belong to 134 genera distributed among The foreshore ranges from 0.5 to 1.0 km from the water 61 Angiospemi families. The monocots constitute 24% of line along the coast. The width of the backshore ranges the total species (Table 2). between 0.5 and 1.0 km from foreshore towards interior. The 10 dominant families of the coastal flora were Backshore flora consists of a mixture of native, naturalized, represented by 96 species (55% of the total species ). The most exotic and some cultivated native species. Exotic species such dominant family was Poaceae (18 species) followed by as Anacardium occidentale, Casuarina equisetifolia and Euphorbiaceae (16 species), Cyperaceae (15 species), Cocos nucifera are planted in the backshore area. Pandanus Fabaceae (11 species). Asteraceae (9 species) and Rubiaceae fascicularis is also often cultivated in the area. Annona (8 species). squamosa, Cleome viscosa. Cassia torn, Opuntia strictci, The floristic components are classified into five growth Chromolaena odorata, Parthenium hysterophorus, Lantana forms (i) grasses and sedges, (ii) herbs, (iii) shrubs, (iv) woody camara var. aculeata, Hyptis suaveolens, Croton climbers (lianas) and (v) trees. The highest number of species bonplandianus, Jatropha gossypifolia, Jatropha curcas are (49%) was grouped under herbs; grasses and sedge were some of the examples of naturalized ahen species that invaded represented by 18% of the species. Trees and shrubs were the coast. represented by 13% and 12% of species, respectively. The woody climbers constituted 8% of species. DISCUSSION The part of the beach that lies between the ordinary low and high tide is included under foreshore. All the species Rao (1971) has divided internal distribution pattern of observed in this region are native. The foreshore consists of the Indian maritime strand flora into three types - one type stabilized and unstabilized sand dunes. The heights of the includes those plants showing complete fidelity to inner stabilized sand dunes ranged between 10 to 15 m and harbour strand, the second type encompasses plants of mid/outer strand Cyperus arenarius, Spinifex littoreus, Ipomoea pes-caprae under the maritime influence, and the third type consists of and Launaea sarmentosa. The unstabilized sand dunes are plants from strand to inland extension. Out of the first type mostly covered with dense patches of Hydrophylax maritima, only the sand strand flora is observed in the district and rock Sesuvium portulacastrum and Digitaria longiflorci. strand is absent. The interesting elements of this strand flora are Cyperus arenarius, Ipomoea pes-caprae, Hydrophylax Table 2: Floristic analysis of strand flora of Ganjam district, maritima, Sesuvium portulacastrum and Spinifex littoreus Orissa, India (Table 1). Rao (1971) while reporting strand flora of India cited Taxa Monocotyledons Dicotyledons Total 75 species of angiosperms for Orissa coast. Most of these species mentioned by Rao are from the northern part of Orissa Families 8 (13%) 53 (87%) 61 (100%) Genera 27 (20%) 107 (80%) 134(100%) and 21 species reported in the paper have also been included Species 42 (24%) 133 (76%) 175 (100%) in Rao’s list of 75 species. The taxa reported by Subudhi et J. Bombay Nat. Hist. Soc., 107 (3), Sep-Dec 2010 217 FLORA OF SANDY COAST OF GANJAM al. (2002) for Orissa coast does not cover Ganjam coast. Ceropegia candelabrum, a climber reported many years ago from the interior part of the state (Khurda and Dhani area of Khurda district) by Haines and Mooney, is now recorded from the vicinity of Casuarina equisetifolia plantations of the sandy coast of Ganjam district. Bulbostylis subspinescens and Micrococca mercurialis were reported from Puri and Konark sandy coast by Haines (1921-1925) and are new records for Ganjam sandy coast. These species have been considered as threatened taxa by Saxena and Brahmam (1994-1996), but on the coast of Ganjam these species are common. The coastal flora is an admixture of native as well as exotic species. Out of the total 175 species, 155 species (89%) are native including 3 (2%) species which are planted. The exotic species in the area are represented by 20 species (11%) of which 16 species (9% of the total) have already been naturalized in the area (Table 1). Fig. 1: Map showing the coastal Ganjam district, Orissa, India Although the exotic species are very few in number, they have invaded the coastal areas to an extent that they ecologically and economically very important. Different have now become a threat to the growth of native species. components of Screw pine Pandanus fascicularis, a semi¬ Antigonon leptopus, Chromolaena odorata, Hyptis suaveolens natural plant, are used by the local people (Panda et al. 2000- and Parthenium hysterophorus are the most notorious invasive 2001) for various purposes. The male flowers of the kewda aliens. The loss of native flora in the coast may be due to plant yield perfume (kewda scent, kewda oil and kewda habitation and privatization of common property resources water), worth millions of rupees (Sahu 2004; Sahu and Misra of the coastal villages (Sahoo and Misra 1994) and 2007). Coconut Cocos nucifera and Cashew nut Anacardium industrialization. The industries such as Indian Rare Earth occidentale are important economic resources of the area. Limited, hatcheries, Gopalpur port and such other activities The creeping stem of Cyperus arenarius, a native species on in the Ganjam coast pose a threat for the very survival of the sands, is utilized to prepare traditional rope. Stem, leaf strand flora. and branches of many of the species, such as Calotropis The aged beefwood (Casuarina equisetifolia) gigantea, Cassia occidentalis, Chromolaena odorata, plantations in the coast provides a congenial environment Lantana camara var. aculeata, and Screw pine Pandanus for the growth of many species such as Aristolochia indica, fascicularis and Palmyra palm Borassus flabelifer are used Asystasia gangetica, Ceropegia candelabrum, Coccinia as fuel. grandis, Emilia sonchifolia, Hybanthus enneaspermus, Micrococca mercurialis, Pergularia daemia, Sida cordifolia. ACKNOWLEDGEMENTS Solatium trilobatum and Tylophora indica. Azadirachta indica, Lannea coromandelica and Morinda pubescens are We thank the Ministry of Earth Sciences, Government also common within the beefwood plantation and seeds of of India for financial assistance and the Head, PG Department these species are usually dispersed by birds. of Botany, Berhampur University, Berhampur for laboratory The plant resources of the coast of Ganjam district are facilities. Bentham, G & J.D. Hooker (1862-1883): Genera Plantamm. 3 vols, London. Reprinted 1961, Botanical Survey of India, Calcutta. 1372 pp. Brahmam, M. & H.O. Saxena (1980): Flora of Ganjam, Orissa (India). Jain, S.K. & R.R. Rao (1977): A Hand Book of Field and Herbarium J. Econ. Tax. Bat. 1: 119-125. Methods. Today and Tomorrow’s Printers & Publishers, Champion, H.G. & S.K. Seth (1968): Revised Survey of Forest Types New Delhi. Pp. 157. of India. Publication Division, Government of India, New Delhi. Mooney, H. (1950): Supplement to the Botany of Bihar and Orissa. Gamble, J.S. 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Misra (1994): Village ecocomplex functioning Regional Research Laboratory and Orissa Forest Development with common property resources: a case study on coastal Orissa. Corporation Ltd., Bhubaneswar, Orissa. 2918 pp. Env. Cons. 21: 57-61. Subudhi, FI.N.. B.P. Choudhury & B.C. Achary (2002): Systematic Sahu, D. (2004): Ecology, energetic and economics of kewda {Pandanus enumeration of plants from Orissa coast. J. Eon. Tax. Bot. 26( 1): fascicularis Lam.) industries in Ganjam district, Orissa. M. Phil. 185-192. J. Bombay Nat. Hist. Soc., 107 (3), Sep-Dec 2010 219