69 Danis Fabricius, 1807 from West New Guinea (Indonesia), a tentative arrangement (Lepidoptera: Lycaenidae, Polyommatinae) Danis Fabricius, 1807 from West New Guinea (Indonesia), a tentative arrangement (Lepidoptera: Lycaenidae, Polyommatinae) Stefan Schröder Auf dem Rosenhügel 15, 50997 Köln, Germany email: [email protected] Suara Serangga Papua (SUGAPA digital) 13(1): 69-90. urn:Isid:zoobank.org:pub: 573CA25A-3C1A-4CB3-82B0-8F7F46670D5D Abstract: An overview is given of the species of the lycaenid genus Danis Fabricius, 1807, occurring in West Papua and Papua (Indonesia). All taxa are discussed with regard to their systematic position and - as far as possible - the type specimens of several species are shown for the first time. Particular emphasis is placed on the individual characters that are intended to help identify the often very similar species. Based on the species list of Hirowatari (1992), the following taxonomic changes are proposed: Danis helga (Grose-Smith, 1898) stat. rev., Danis horsa (Grose-Smith, 1898) stat. rev., Danis metrophanes (Fruhstorfer, 1915) syn. nov., Danis hermogenes (Fruhstorfer, 1915) stat. rev. Rangkuman: Peninjauan secara luas diberikan kepada spesies dari lycaenid genus Danis Fabricius, 1807, ditemukan di Provinsi Papua Barat (Indonesia). Semua taxa dibahas yang berhubungan dengan posisi sistematik sejauh mungkin. Type spesimen dari beberapa spesies ditunjukkan untuk pertama kalinya untuk semua spesies. Penekanan khusus diberikan untuk karakteristik individu yang ditujukan untuk membantu mengidentifikasi spesies yang sangat mirip. Berdasarkan list spesies Hirowatari (1992), perubahan taksonomi diusulkan sebagai berikut: Danis helga (Grose-Smith, 1898) stat. rev., Danis horsa (Grose- Smith, 1898) stat. rev., Danis metrophanes (Fruhstorfer, 1915) syn. nov., Danis hermogenes (Fruhstorfer, 1915) stat. rev. [translation by Daawia Suhartawan] Keywords: Polyommatinae, Thysonotis, Danina, Papua New Guinea Introduction The genus Danis Fabricius, 1807 includes comparably large lycaenid butterflies, representing a magnificent part of the butterfly fauna of New Guinea, where it shows the highest species diversity. The most common species is Danis danis (Cramer, 1775), which is also the one with the widest distribution, reaching from eastern Maluku to northern Australia. All other species are generally rare and much less is known about them, even though most of them are known to science since a long time. No recent revision of the genus is available and only few new taxa were described after the early twenties of the last century. Apparently, the lycaenid specialist G. E. Tite was working on the group in the 1960s, as he dissected specimens and also added comments on some, but nothing was published. There are only two papers by Yagishita (2000) and Tennent (2016), covering Danis danis, but not concerning other species. Nothing substantial was added after publication of Fruhstorfer’s „Thysonotis“ list in 1915. The most recent publication, illustrating many of the DOI: 10.19269/sugapa2020.13(1).05 70 Schröder, S., 2020. Suara Serangga Papua (SUGAPA digital) 13(1): 69-90 species, is the Australasian volume of D’Abrera`s Butterflies of the World series, dating back to 1971. Hirowatari (1992) gave a revised species list, including all taxonomical changes taken in the last century. Species diversity is apparently lower in PNG and accordingly, Parsons (1998) illustrates only a few species. Selected information concerning limited areas in West Papua were given by Gotts & Pangemanan (2010), van Mastrigt & Rosariyanto (2005), van Mastrigt & Warikar (2013) and Tim Redaksi KEP (2010). The genus name Danis has a long history, but was not always accepted. Just a few years after Fabricius introduced the name in 1807, Hübner (1819) described Thysonotis. Druce & Bethune-Baker (1893: 537) rejected the older name Danis as „a generic name must not be one which has been used as a specific name“ and placed all species known at this time in Thysonotis, dividing it into five groups. They included a danis-group to accomodate the species we know today as Danis danis. Due to the selection of Papilio danis Cramer, 1775 as type species of Danis by Hemming (1964), the well used name Thysonotis Hübner, 1819 became a synonym of Danis. Thysonotis had been used for many decades and became a kind of “waste basket” over the years to include various phenotypical similar species, many of them unrelated, in a group which is known as the Danis/Psychonotis mimicry complex (Parsons, 1998: 98). Subsequently, taxonomy of this complex was resolved and its members were correctly assigned to several other genera like Psychonotis, Nacaduba, Nothodanis and Perpheres (Hirowatari, 1992). Druce & Bethune-Baker (1893: 543) separated a “wallacei-group” from the more robust danis, remarking that they “have the cilia of both sexes pure white, not alternately black and white as in danis and allies”. Aside of T. perpheres Druce & Bethune-Baker, 1893 (now placed in Perpheres Hirowatari, 1992), T. melimnos Druce & Bethune-Baker, 1893 was included. However, the groups mentioned are not unambiguous and also depend heavily on preservation of specimens, as it is often not possible to observe this character in worn specimens. Far more important for determination, however, are the following characters: development of the white band on the hindwings (weak or very strong, its width and inclination), the width of the wing margins, presence of a white forewing patch, a tornal spot on the forewing underside and colour of space 7 of the hindwing upperside. Basic works include Grose-Smith (1894, 1897), Grose-Smith & Kirby (1895), Druce & Bethune-Baker (1893) and Fruhstorfer (1915). Hirowatari (1992) gave a very useful revised species-list and commented on some taxa, also giving new combinations, but did not figure any specimens. With the exception of Danis danis and its subspecies, determination of other Danis species is notoriously difficult. The reason for this lies undoubtedly in the complex situation that different species have a very similar wing pattern and are in most cases rare. Little is known about individual variation, especially concerning the different shades of blue, the width of the white bands on the hindwing underside and wing margins in specimens from different populations and altitudes. Except for the original descriptions and the Australian volume of D´Abrera (1971) there are almost no illustrations available, which could help with determination. Type specimens were never photographically figured and therefore misidentifications are prevalent. In addition, it may be very difficult to assign females to the corresponding males. Even the type series are inconsistent and males and females are not always correctly assigned to each Published on 17th December 2020 71 Danis Fabricius, 1807 from West New Guinea (Indonesia), a tentative arrangement (Lepidoptera: Lycaenidae, Polyommatinae) other, including different species. In several cases the authors of the last century included males and females from different localties in a type series, which do not belong together. It is possible to separate males from each other, but it is extremely difficult to match the correct females with them, if not collected at the same locality. To complicate matters, identical specimens from a given locality were assigned as allotypes to different species. For example, the allotype of Thysonotis helga belongs to the same species as the female holotype of melimnos (but maybe the male of helga has to be matched with the female of melimnos) and the metrophanes allotype belongs to the same species as the allotype of horsa. A similar situation is known from other lycaenid genera like Nacaduba or Jamides. Even if it is impossible to solve all problems at species level at this time, it is important to document species diversity known so far, so that a better assessment of the different forms can be made. Finally, a conclusive revision without DNA data might not be possible. However, this requires the availability of fresh, topotypic material for most of the species, which may also be problematic given the rarity of some species. Unfortunately DNA sequencing is very difficult is not yet possible for historical type specimens, which seems to be the only way to solve the systematic position of some taxa. For the time being, and as an aid for determination, phenotypic characters are used in this work to separate different species and most of the types are illustrated, so that a determination for the taxa occurring in western New Guinea should be possible. With few exceptions, names of synonyms are only given for taxa occurring in West Papua and Papua. Locality data are mainly based on the KSP collection. Abbreviations ANIC – Australian National Insect Collection, Canberra, Australia HT – holotype (in literature) NHMUK – The Natural History Museum, London, United Kingdom CSSK – Collection Stefan Schröder, Köln, Germany KSP – Koleksi Serangga Papua, Universitas Cenderawasih, Waena, Papua, Indonesia RMNH – Naturalis Biodiversity Center, Leiden, The Netherlands MFNB – Museum für Naturkunde, Berlin, Germany Scale bar in plates is 1 cm Systematical part Danis danis (Cramer, 1775) This species was recently subject of a very detailed review, including all known subspecies (Tennent, 2016), and there is no need to repeat the information presented in that publication. Of the presently known 24 subspecies only four occur in Papua and West Papua: apollonius C. & R. Felder, 1865 [mainland PNG, Waigeo, Salawatti and the islands in Teluk Cenderawasih = Geelvink Bay]; hermes Grose-Smith, 1894 [Schouten Islands], and two new DOI: 10.19269/sugapa2020.13(1).05 72 Schröder, S., 2020. Suara Serangga Papua (SUGAPA digital) 13(1): 69-90 subspecies described by Tennent: ssp. gebe Tennent, 2016 [Gebe Island] and ssp. kofiau Tennent, 2016 [Kofiau Island]. Tennent (2016) is illustrating numerous specimens, including detailed discussions and illustrations of the type species of apollonius, panaetius, herophilus and sophron. Danis danis apollonius (C. & R. Felder, 1865) (figs 1-6) Lycaena apollonius: C. & R. Felder (1865: 265, pl. 33, fig. 3). [HT ♀ “Nova Guinea ? type”, NHMUK] Thysonotis apollonius ab. plumbeus: Rothschild (1915: 140). [HT ♂ “Misol”] Thysonotis danis panaetius: Fruhstorfer (1915: 50). [HT ♀, Salawatti, RMNH] Thysonotis danis herophilus: Fruhstorfer (1915: 50). [LT ♂, “Waigiu”, NHMUK] ? Thysonotis danis sophron: Fruhstorfer (1915: 50). [HT ♀, “New Guinea”, NHMUK] [See Tennent (2016: 128)] ? Thysonotis lona: Röber (1927: 105). [“Waigeo”, type not located] Danis danis (apollonius) thinophilus: Toxopeus (1930: 129). [HT “Misool”, probably in RMNH] {nom. nov. pro apollonius Fruhstorfer nec Felder} Notes: Thysonotis lona Röber, 1927 is little understood. The translation of its description is as follows: “The available male from Waigeo differs from philostratus Feld. males from Halmaheira (Gilolo) in having a lighter colouration (more greenish than bluish); which is also reduced (because of the broadened white band), the black edge on front and margin is also narrower, the blue of the hindwing is reduced, but more present than in philostratus. Underside colouration is also more greenish and stronger. It is not known to me if typical philostratus [North Maluku !] occurs on Waigeo, but if this should be the case, lona could possibly be a seasonal form of philostratus.” Apparently Röber regarded lona as closely related to philostratus, which is the northern Maluku race (Halmahera, Ternate, Bacan and Kaioa) of Danis danis. Given the geographical distribution as presently known, lona is most likely a synonym of apollonius, which has already been proposed by Tennent (2016). The type material of lona is apparently lost, as neither the collections in Dresden or München hold any specimens labeled with that name. Distribution: Widely distributed in Papua and Papua Barat (Indonesia), including the Raja Ampat Islands and Schouten Islands, and in Papua New Guinea. Danis danis gebe Tennent, 2016 (figs 7-10) Danis danis gebe: Tennent (2016: 119, figs. 25-29). [HT ♂ Gebe Island, NHMUK] Notes: Description of this island race is based on a few specimens only. Tennent (2016) remarked “hindwing basal black area extending slightly along costa, leaving distal edge of black area distincly curved”. More material has to show if this is a diagnostic character of this subspecies, which otherwise does not show signifiant differences to ssp. apollonius, occurring on Waigeo, which lies only a few kilometres east of Gebe. Distribution: Only known from Gebe Island (Moluccas, Indonesia). Published on 17th December 2020 73 Danis Fabricius, 1807 from West New Guinea (Indonesia), a tentative arrangement (Lepidoptera: Lycaenidae, Polyommatinae) Danis danis kofiau Tennent, 2016 (figs 11-14) Danis danis kofiau: Tennent (2016: 121, figs. 47-51). [HT ♂ Kofiau Island, NHMUK] Remarks: Differs from other subspecies in having a broad costal forewing border in the males and a broader white median band in the females. Distribution: Only known from Kofiau Island (Papua Barat, Indonesia). Danis danis hermes (Grose-Smith, 1894) (figs 15-18) Thysonotis hermes: Grose-Smith (1894: 575). [HT ♂ Korrido (Supiori) and Biak, NHMUK] Thysonotis danis phoibides: Fruhstorfer (1915). [HT ♂ “Insel Mefor”, NHMUK] Notes: This taxon is characterized by the “duller and more greenish” blue wing colour and the rather broad dark borders. The species is already figured very well in Grose-Smith & Kirby (1895: pl. 1, fig. 5-8). Tennent (2016) explained the synonymy of phoibides Fruhstorfer, 1915 in detail. Distribution: Endemic to the Schouten Islands: Biak, Supiori, Numfor (Papua, Indonesia). Danis phroso (Grose-Smith, 1897) (figs 19-28) Thysonotis phroso: Grose-Smith (1897: 313). [HT ♀ “Etna Bay, Dutch New Guinea”, NHMUK] ? Thysonotis lygia: Grose-Smith (1897: 516). [HT ♂ “Dinner Island” = Samarai Isl., Milne Bay Province, PNG, NHMUK] Thysonotis melimnos scarpheia: Fruhstorfer (1915: 50). [Type ♀ “Dinner Insel, Ost-Neu Guinea”, Dep. ?] Danis phroso (Grose-Smith, 1897): Hirowatari (1992: 26, fig. 30A). [♀ genitalia] Identification: Most useful in determining phroso is the narrow subbasal band on the hindwing underside, which is blue in its lower and white in the upper part, a character which is not found in any other Danis, except for D. concolor (Rothschild, 1915). Males have a white discal patch on the forewing upperside, which reminds of the superficially similar looking males of the genus Perpheres Hirowatari, 1992. The patch is variable and may be very extensive as in the specimen figured by Parsons (1998: pl. 67, fig. 1891) or just faintly developed as in a specimen from Mimika (fig. 25, KSP 11861). Hindwing space 7 is white. Females do not show any traces of a white band on the hindwing upperside, which is present in all other species except D. concolor. Instead, there is a greenish-blue basal suffusion on the hindwing as well as at costa and dorsum of the forewing. Notes: Danis phroso was described from four female specimens and compared with D. wallacei (C. & R. Felder, 1865) from Waigeo, which is a very different species. The synonymy of lygia and scrapheia with phroso as proposed by D´Abrera (1971) is not yet settled as both occur in eastern PNG, while phroso is known from Papua. Characters of both taxa are not completely identical: lygia females from eastern PNG have the blue metallic scales on costa and dorsum of the upperside of the forewing much stronger than in females DOI: 10.19269/sugapa2020.13(1).05 74 Schröder, S., 2020. Suara Serangga Papua (SUGAPA digital) 13(1): 69-90 from Papua, and the marginal green metallic line along the forewing costa and termen on the underside is more complete (Parsons 1998: pl. 67 fig. 1891-93). In his description of lygia, Grose-Smith (1897) already mentioned the similarity with phroso, from which males differ in having a wider black hindwing border and an indistinctive white patch on the forewings. Additional material from PNG is needed to understand the specific variability of lygia and the proposed synonymy. It can not be excluded that populations from eastern PNG belong to a separate subspecies. Distribution: The type locality “Etnabaai” (Etna Bay, Papua, Indonesia) is located Southeast of Kaimana in Southwest Papua and is the southernmost of several large bays reaching far into the mainland on the southern coastline. Furthermore Timika, Yahukimo, Mimika (Papua, Indonesia) and Papua New Guinea. Danis concolor (Rothschild, 1915) (figs 29-32) Thysonotis phroso concolor: Rothschild (1915: 32). [Type ♂ “Snow Mountains, Base Camp”, Dep.?] Danis concolor (Rothschild, 1915): D’Abrera (1971: 325); Hirowatari (1992: 26, fig. 31A). [♀ genitalia] Danis phroso concolor (Rothschild, 1915): Gotts & Pangemanan (2010: 244-245). [♂ refigured herein, figs. 29-30]. Identification: Rothschild (1915) describes ssp. phroso concolor very briefly as follows: “♂. Differs from p. phroso in its larger size and the uniform blue of the forewings and absence of metallic green on fore- and hindwings”. ♀ has the base of wings greyish lavender blue, NOT bright blue.” Both species are very similar, but can be separated by the following characters: dark hindwing margins of the concolor male are much narrower than in phroso. The white forewing patch of the males is small and more or less restricted to space 2, in some specimens it is completely missing (fig. 29). The white patch on the forewing in females is less compact in concolor and usually divided by brown along the veins. In females the upper half of the cell is brown but there are white scales in the basal parts of spaces 4-6 so that the white seems to be cut off abruptly at almost a right angle. Females of both, phroso and concolor have a blue basal suffusion on the upper side of the hindwings, which is not known from any other species in this group, aside of horsa. They both have in common the underside subbasal band divided in white and blue-green. The upperside subcostal blue stripe in the females is variable in both species. Notes: Hirowatari (1992: 27, 25) mentions the unusual form of the valvae of this species, having a finger-like process at their ends, and therefore regards concolor as the “most disctinctive” taxon within Danis. Usually the distal margin of the valva bears a minute serration. Distribution: Snow Mountains, Mimika, Timika (Papua, Indonesia). Published on 17th December 2020 75 Danis Fabricius, 1807 from West New Guinea (Indonesia), a tentative arrangement (Lepidoptera: Lycaenidae, Polyommatinae) Danis glaucopis (Grose-Smith, 1894) (figs 33-36) Thysonotis glaucopis: Grose-Smith (1894: 575). [HT ♂ “Humboldt Bay”, NHMUK] Danis glaucopis (Grose-Smith, 1894): Parsons (1998: 431, pl. 67, fig. 1894-1896). Identification: A round winged species with a rather well developed white band on the hindwing upperside in males and females. Accordingly, in the females the basal part of the hindwing appears almost completely white but not brown as in many other species. The white markings on the forewing underside are reduced in extension to a small, sometimes rounded patch, leaving a wide dark brown area and according to Parsons (1998: 431) glaucopis is best identified by this character. This fits well with the original underside figure of Grose-Smith & Kirby (1895: pl. 1 fig. 10). Grose-Smith (1894: 576) compared glaucopis in particular with wallacei, which has a much better defined, solid white hindwing band and a wider hindwing margin in the males. Females of wallacei have a broader white underside band than glaucopis and the white forewing patch is much larger. Danis glaucopis also resembles D. metrophanes Fruhstorfer, 1915, described from Sorong, but according to Fruhstorfer (1915: 49), metrophanes differs in having the white band on the hindwing upperside almost completely reduced. Danis drucei has the white underside forewing patch significantly larger than in glaucopis or metrophanes. The blue of the males is deeper in drucei and wing margins are broader. Space 7 of the hindwing upperside is completely white in drucei. In addition, the white underside hindwing band is narrower, more inclined, so that the band almost touches the basal green- blue on the hindwing costa. The green costal stripe on the underside of the forewing is incomplete in metrophanes. Another species closely resembling glaucopis is the East Sepik (Papua New Guinea) endemic Danis regalis (Grose-Smith & Kirby, 1895). Notes: A rare species. The type series is from Wendesi (at the “neck” of the Birdshead Peninsula) and the Jayapura area (Humboldt Bay). According to Parsons, the holotype specimen is deposited in the NHMUK. Additional specimens are said to be in Staudinger collection in MFNB. Distribution: FakFak (Papua Barat) and Sarmi (Papua, Indonesia), and Papua New Guinea (Parsons, 1998: 431). Danis helga (Grose-Smith, 1898) (figs 37-46) Thysonotis helga: Grose-Smith (1898: 105). [HT ♂ “Ansus, Jobi Island”, NHMUK] Danis melimnos helga (Grose-Smith, 1898): Hirowatari (1992: 26). Identification: The upperside of the holotype of Danis helga (fig. 41) is bright blue with a small but distinctive white patch in the forewing spaces 1b-3 and a very broad white band on the hindwings. The dark wing margins are comparably broad. Grose-Smith (1898: 105) compares helga with horsa (Grose-Smith, 1898), which is now placed as a good species, but lacks the clear white band across the hindwing in the males. Compared with this species, helga males have a very prominent and broad white hindwing band. Grose-Smith also mentions: “Cilia of both wings narrowly white, crossed with black at the ends of the veins.” DOI: 10.19269/sugapa2020.13(1).05 76 Schröder, S., 2020. Suara Serangga Papua (SUGAPA digital) 13(1): 69-90 As the males, females are characterized by a very broad white band on the hindwing undersides, resembling wallacei females in this respect. D. melimnos appears to be much closer concerning male genitalia structure and therefore Hirowatari (1992: 26) and Parsons (1998: 431) regarded helga as the mainland Papua subspecies of melimnos. However, both melimnos and helga have Yapen as type locality, and thus can not be regarded as different subspecies of the same species. The situation is complicated by the fact that the allotype of helga (Figs 47-48) is conspecific with the female holotype of melimnos (Fig. 53). Further research and new material from Yapen is needed to ascertain that males and females of both taxa are correctly placed with each other. It can not be completely excluded that the male of helga has to be matched with the female of melimnos, but for now, both names are retained and applied to different species: This has also been followed by D’Abrera (1971: 326), mentioning helga being “deceptively like melimnos, but distinguishable by its larger size and by the rounder shape of the wings in both sexes.” D. melimnos has very broad white bands on the undersides of both wings (Grose-Smith, 1898: pl. 46, fig. 2). It is more likely that helga is conspecific with wallacei described from Waigeo. The pictured specimen from the KSP collection (figs 37-38) is quite similar to mamberano (Joicey & Talbot, 1916), but this species has a different wing shape, the blue on the hindwing appears jagged on the veins and the hindwing margin is wider. Distribution: Yapen Island (Papua, Indonesia). Danis mamberano (Joicey & Talbot, 1916) (figs 49-50) Thysonotis mamberano: Joicey & Talbot (1916: 82). [HT ♂: “River Mamberano, N. Dutch New Guinea”, NHMUK] Identification: According to Joicey & Talbot (1916: 82), mamberano is “nearest helga, Gr.- Sm., from Jobi”. In their short description it is said that wing margins and the white hindwing band are narrower than in D. helga. The forewing shows an indistinct white band from inner margin to vein 4. On the underside the costal blue is broader beyond the cell and the blue touches the white underside colour at tornus. Hindwing with white band as above. Notes: Described from a single male. The wing colour is a grey-blue with a diffuse discal band on the forewing and a compact white band on the hindwing. Hindwing margin is wide, with the blue-grey scaling indented between the veins. D. mamberano differs from helga concerning colouration as well as the extent and inclination of the white bands. Further research and new material is needed to confirm its taxonomic status. It can not be excluded that mamberano is a ssp. of helga, however, the blue wing colour of the male is darker than in any other known Danis species occurring in Papua. Distribution: Mamberano Area, Northern Papua (Papua, Indonesia). Danis melimnos (Druce & Bethune-Baker, 1893) (figs 51-53) Thysonotis melimnos: Druce & Bethune-Baker (1893: 544, pl.46, fig. 2). [HT ♀ “Dutch New Guinea, Jobi I. (A. B. Meyer)”, Dep.? “Type is contained in Staudinger coll.”, Berlin?] Published on 17th December 2020 77 Danis Fabricius, 1807 from West New Guinea (Indonesia), a tentative arrangement (Lepidoptera: Lycaenidae, Polyommatinae) Identification: The original description of T. melimnos is based on a single female, which was compared with wallacei C. & R. Felder, 1865, but already suggested to “prove quite distinct from wallacei” by the authors. The white bands at the underside follow more evenly the wing shape and appear broader in melimnos, and the white forewing upperside discal patch is not extending towards the termen as in wallacei. On the forewing underside the white does not reach the tornus. The white patch at the underside in females is elongated, very regularly outlined and almost parallel to the green-blue marginal markings, so that only a small band of brown remains in between. However, the variability of this character is not known. Notes: A very different interpretation of melimnos was given by Parsons (1989: 432), stating that the best way to determine melimnos is the underside colour of the bands, which is not plain white, but has a creamy-white to yellowish hue. This is the case in the female specimen designated as allotype of Thysonotis helga from Yapen, which clearly belongs to melimnos (figs 47-48), but in other specimens (fig. 52) the bands are pure white without any yellowish hue. The female specimen illustrated by D’Abrera (1971: 326) shows a short streak of greenish blue scales along the upperside forewing costa and Parsons (1998: 432) remarks that of all Danis species occurring in PNG only melimnos females have a broad subcostal band of blue scales on the forewing upperside. This is not the case in western New Guinea, where this character is also known from T. vidua Grose-Smith & Kirby, 1895 (= wallacei C. & R. Felder, 1865) and other unrelated species. Unfortunately, the underside of the specimen illustrated by Parsons (1998: pl. 67 fig. 1899) is not shown, but most likely it does not have the broad white bands, characterizing true melimnos. Druce & Bethune-Baker do not mention any metallic costal scales on the forewing and neither does the figure of the type show them. A female from Yapen (figs 51-52) has only a very weak streak of costal blue scales on the forewing. The female specimen figured as D. melimnos by Parsons (1989: pl. 67, fig. 1899) has much narrower white bands on the upperside of both wings, and such specimens were tentatively assigned to an undescribed mainland PNG race of melimnos by him. Possibly, the female from Yahukimo (figs 57-58) and the one from Vriendschap River (Kabupaten Asmat, 05°20’20” S - 138°52’72” E, figs 55-56) also belong to this new taxon. However, it is more likely they represent females of other species. Gotts & Pangemanan (2010: 244) mentioned that specimens from Mimika placed by them with melimnos are closer to this PNG race than to the specimens illustrated by D’Abrera. The rather narrow but regular yellowish bands on fore- and hindwing are also present in T. athanetus, which is regarded here as synonym of wallacei. Furthermore, the melimnos male Parsons figures on pl. 67 fig. 1897/1898 is very similar to the one figured by him as D. regalis (Grose-Smith & Kirby, 1895) and likewise, the male “melimnos” specimen in Gotts & Pangemanan (2010: 245) does not differ considerably from the figure of Grose-Smith & Kirby (pl. 2 figs 4-5) given for regalis. Both taxa show a much greater resemblance to regalis than to melimnos. However, T. regalis is little understood. The description gives “New Guinea” as distribution and later on Fruhstorfer (1915: 49) recorded it from “Deutsch-Neu-Guinea” where it is said to be the most common species at “Astrolabebai” (Madang Province of north-eastern PNG). Nothing is known about its distribution or individual variability and the whereabouts of the DOI: 10.19269/sugapa2020.13(1).05 78 Schröder, S., 2020. Suara Serangga Papua (SUGAPA digital) 13(1): 69-90 type are also unknown (“In the Collection of Dr. Staudinger”). Judging from the original illustrations (figs 59-60) it appears to be a species with a very conspicuous, broad hindwing margin and a rather narrow white band on the hindwings in the male. Currently there exists no valid definition of melimnos, especially concerning matching males, as only insufficient material for study is available from the type locality Yapen. Distribution: Yapen Island (Papua, Indonesia); possibly endemic. Danis wallacei (C. & R. Felder, 1865) (figs 61-72) Lycaena wallacei: C. & R. Felder (1865: 265, pl. 33 figs 8-10). [Type “Waigiou”, Dep] Thysonotis vidua: Grose-Smith & Kirby (1895: 30, pl. 2, f. 9-10). [Type ♀: “Waigiou”] Thysonotis albomarginata: Rothschild (1915: 140). [HT ♀: Misool, NHMUK] Thysonotis melimnos athanetus: Fruhstorfer (1915: 50). [HT ♂: „Salawati“, RMNH] Identification: The illustrations of Felder & Felder (1865) in the “Reise der österreichischen Fregatte Novara um die Erde” are generally accurate if compared with recently photographically illustrated type specimens (e.g apollonius, or philostratus; see Tennent, 2016), so there is no reason to doubt the reliability of the illustration of the male holotype on pl. 33 fig. 8. Most of the taxa described by the Felders are now in the NHMUK (Rothschild Bequest B.M. 1939) and it is most likely that the type of wallacei is in the British Museum. Unfortunately, it was not possible to locate this specimen. Both, the vidua and wallacei type- specimens are not preserved in the MFNB, Berlin. Based on the illustrations, D. wallacei males are apparently characterized by a solid, slightly curved and very prominent white hindwing band, which appears to be even broader than in drucei. Variation of the conspicious white forewing patch (spaces 1b-3) in the males is insufficiently known; it is well visible (see D’Abrera, 1971: 327), and may be as strong as known from helga. A large white forewing patch is present in the females, expanding outwardly along spaces 2 and 3; basal area is brown (not white as in drucei or other Danis females). On the underside, there is a small black spot in space 1b close to the tornus of the forewing as in hengis and drucei. A faint line of green-blue metallic subcostal scaling is present in the females, which is also found in melimnos. Grose-Smith & Kirby (1895: 30) mentioned differences in the width and extension of this green marginal line on the forewing between vidua (= wallacei) and drucei, but these characters seem to vary. Grose- Smith & Kirby (1895) had already mentioned the close resemblance of vidua to wallacei: “Underside almost exactly like that of the female of T. Drucei [(postea)], except, that in the anterior wings the curved outer part of the blue band is broader, and extends to the white portion of the wing…” Notes: Both, melimnos athanetus Fruhstorfer, 1915, as well as wallacei hermogenes Fruhstorfer, 1915 are from Salawati (Bernstein collection, RMNH) and closely resemble each other. Additional labels of Van Eecke attached to both types give a determination as wallacei, which may have been the reason for D’Abrera (1971: 326) to include both in wallacei. However, both type specimens are not conspecific: they are very similar, but there are some differences concerning the underside markings. In hermogenes (Fig. 93-94) the white patch Published on 17th December 2020