67 Australian hepatic types at the National Herbarium of New South Wales (NSW) and the Ray Herbarium (SYD) E.A. Brown, H.P. Ramsay and J. Seur Abstract E.A. Brown, Ramsay, H.P. & Seur, /. (National Herbarium of Nexv South Wales, Royal Botanic Gardens, Sydney, NSW, Australia 2000) 1992. Australian hepatic types at the National Herbarium of New South Wales (NSW) and the Ray Herbarium (SYD). Telopea 5(1): 67-90. The National Herbarium of New South Wales and the Ray Herbarium in Sydney, New South Wales hold important collections of type specimens of Austalian hepatics. A total of 99 names are listed here, together with details of the protologue, complete label data for each specimen, other herbaria known to hold part of the type, and recent information concerning the status of the name. Included are many Stephani and Carrington and Pearson names as well as more recent ones described by Hewson and Na-Thalang. Introduction While the Hepatic Collections in the two herbaria in Sydney, New South Wales, are not large [6000 specimens, 100 types (NSW) and 2284 specimens, <10 types (SYD)] (Ramsay & Seur 1990), type specimens of many Australian and exotic species are present. Particularly significant collections held at the National Herbarium of New South Wales [NSW] are those of T. Whitelegge, W.W. Watts (including some from W. Gunn), and H.J. Hewson; and at the Ray Herbarium, University of Sydney [SYD], those of Hewson and O. Na-Thalang. The main geographical regions represented by the types are Australia including Lord Howe Island (Whitelegge, Watts, Na-Thalang), presented in this paper, and Vanuatu [New Hebrides] (W. Gunn) and Papua New Guinea (Hewson) to be reported separately. The earliest descriptions of New South Wales hepatics are those of B. Carrington & W.H. Pearson (1888) who determined and published a list of Whitelegge's collections. R.H. Bastow's work on Tasmanian hepatics (1888, 1914) also included reference to many New South Wales collections. Watts published several papers on Australian hepatics (1902, 1903, 1905) and left a manuscript (1912), as part of a Census of New South Wales Plants, that he had intended publishing in the same way as he had done for the mosses (Watts & Whitelegge 1902, 1906). In this unpublished manuscript he listed species, localities and collector's names for 205 Hepatics and 10 Anthocerotes. The names contained in this unpublished manuscript include many described by F. Stephani based on specimens sent to him for identification (Stephani 1884, 1889; Stephani & Watts 1914). The cryptogamic collection at NSW was increased greatly in the late 19th and early 20th century by W.W. Watts. After his departure to Victoria in 1916 (Ramsay 1980) curation and addition to the collections was very limited. Conditions at NSW were extremely cramped until 1982 when a new building opened and the W.W. Watts room was set aside for cryptogams. A period of inactivity in collecting and describing hepatics in New South Wales persisted from 1920 until the mid 1960s, when taxonomic research was undertaken 68 Telopea Vol. 5(1): 1992 by G.K. Berrie and R. Carolin's students Hewson and Na-Thalang, at Sydney Univer¬ sity. In 1963 Berrie published Haplomitrium intermedium, the first member of the Calobryales recorded in Australia, from a type collected in the Blue Mountains (Ber¬ rie 1963). Hewson (1970a, b) revised the family Aneuraceae in Australia and New Guinea while Na-Thalang (1980) revised the genus Riccia in Australia. Hewson and Na-Thalang types are located either at SYD or NSW and in some cases at both. There is still much to be learnt about hepatics in New South Wales; we do not even have a complete species list for this State in which both subtropical and temperate taxa occur. Some recent publications (Ramsay et al. 1990, Downing et al. 1991) and further studies being undertaken by H.P. Ramsay et al. list the hepatics for specific areas in N.S.W. and form the basis for the eventual production of a Census of New South Wales hepatics. Locating types is essential for taxonomic revisions and a register providing details and locations of type specimens can be useful. Ramsay & Seur (1990) and Ramsay et al. (1990) outlined the methods for collection and recording of data for a register of type collections of mosses. Their investigations highlighted the often incomplete in¬ formation in many type citations in taxonomic literature (e.g. date of collection and collector's number). Since most descriptions of species of hepatics published prior to the 1970s do not cite the herbarium holding the type the holotype cannot be clearly defined. Examining the type specimen and its label data may give important details not included in the protologue. Method In this paper we deal only with those hepatic type specimens of Australian species held in the two herbaria named NSW and SYD. The studies were carried out in a series of steps: 1. All data on each herbarium packet containing specimens designated as types or sp. nov. (i.e. putative types) were recorded on index cards. 2. The type description for each taxon was obtained and the label data compared with that in the protologue. Where these matched, the specimen is recorded as a probable type. 3. Where possible, reference is made to the herbarium holding the major collections of the author of the basionym if this is not NSW or SYD. In most cases, except for species named after the mid 1970s, the holotypes are held in European herbaria and specimens in Australia usually are isotypes. For a number of species a series of specimen numbers were listed with the type description and are syntypes and it may be desirable, therefore, when taxonomic revisions are carried out, to designate a lectotype at that time. 4. References are given for recent studies, name changes, lectotypification etc. for each taxon. 5. To ensure that we have detected all relevant type specimens held at NSW and SYD, we checked publications and searched through all collections at NSW and SYD for specimens not labelled as types but carrying the original label data. As the status of quite a number of specimens had been overlooked in the repackaging at NSW during the early 1980s these further checks, although time-consuming, proved rewarding. 6. The information is now stored on disk and will remain available for future updating. Brown, Ramsay & Seur, Hepatic types 69 The register of type specimens of Australian hepatics presented here has been ar¬ ranged alphabetically by genera and species based on the name for which the specimen is a type. The list contains the original name for each taxon and reference, the pro- tologue [Proto] (collection details as cited in the protologue), full label data [Label] for each separate specimen (label 1, 2 etc.) at NSW and/or SYD if more than one was present. Also listed are location and status of all types [Types] where known (from examination or from literature), synonymy where applicable, as well as any other recent information such as lectotypification [Comment]. At some time in the past many of Watts' collections have been broken up, repacket- ted and relabelled; the duplicates may all be contained within the one specimen folder or sometimes they have been separated. Where label information does not differ significantly between packets it is usually listed only once, with a note that there are 2 or more specimens; sometimes information on packets within the same specimen folder differs markedly and this has been recorded. The labels on the outer specimen folders has not been recorded as it is only a more recent copy of the information on the Watts' packets. No taxonomic decisions other than those already in the literature are presented here, nor is lectotypification of any species implied. This is particularly important for taxa described by Stephani; in his later years he broke up many of his specimens and sold duplicates to BM, FH and other institutions. Some of these duplicates, when they come from mixed collections, are not the taxon referred to in the protologue and great care must be taken when typifying. It is probably unwise and inaccurate, therefore, to cite holotypes and this has not been done, even when Stephani specified a collector's number. Herbaria, other than NSW and SYD, holding types have been listed from Index Hepaticarum. Index Hepaticarum VIII (Bonner 1976) and prior vol¬ umes were used to obtain putative label information for many of the types held at G. E.A. Brown has examined the type specimens from overseas herbaria for the genera Riccardia and Aneura in the course of other studies (Brown & Braggins 1989). Register of names Aneura aequicellularis Stephani (Stephani & Watts 1914: 95) Proto: 'Australia, New South Wales (Wentworth Falls): Watts legit No. 117 [1117].' Label: New South Wales: Wentworth Falls, cave, W.W. W[atts] 1117, Sep 1912. Types: G (lectotype), NSW (isolectotype). Comment: = Riccardia aequicellularis (Stephani) Hewson fide Hewson (1970b: 79). Lectotypified by Brown & Braggins (1989: 26). Aneura athertonensis Hewson (1970a: 188) Proto: 'Holotype — Charmillan Creek, Tully Falls Road, Atherton Tableland, on basalt rock on creek bank in disturbed rain-forest, Qld. Hewson, 422, 8.1964 (NSW): Isotype (BRI).' Label: Queensland: Charmillan Creek, rock, rainforest, basalt, H.j. Hewson H.64.71.422, 22 Aug 1964. Types: NSW (179239: holotype), BRI (isotype). Comment: Name in current use. 70 Telopea Vol. 5(1): 1992 Aneura coriacea Stephani (1917: 23) Proto: 'Australia: New South Wales (Watts legit.)-' Label: New South Wales: Katoomba Falls, W.W. Watts 567, 15 Jan 1902. Types: G (11036: lectotype), NSW (H241: isolectotype). Comment: No particular specimen cited in protologue. The NSW specimen has been annotated by Hewson: 'Isotype of A. coriacea = Riccardia crassa, published Hewson 1970b'. Lectotypified by Brown & Braggins (1989: 85). Specimen in G seen by Steph¬ ani is Watts 567 (leones Hep. 474). Aneura eachamensis Hewson (1970a: 184) Proto: 'Holotype — Vision Falls, Lake Eacham, Atherton Tableland, on basalt rock in rain-forest, Queensland. Hewson, 398, 8.1964 (NSW): Isotype (BRI).' Label: Queensland: Vision Falls, Lake Eacham, H.]. Hewson H.64.68.398, 20 Aug 1964. Types: NSW (179240: holotype), BRI (isotype). Comment: Name in current use. Aneura gigantea Stephani (Stephani & Watts 1914: 95) Proto: 'Australia, N.S. Wales, (Cambewarra): Watts legit No. 920.' Label: New South Wales: under cliff, above Baldy's, Cambewarra, W.W. Watts 920, 18 Oct 1907 [3 packets in one specimen folder]. Types: G (11042), NSW. Comment: = Aneura alterniloba (Hook. f. & Taylor) Taylor & Hook. f. in Gottsche, Lindenb. & Nees var. gigantea (Stephani) Hewson; G 11042 and '920 NSW' are cited as the holotype and isotype respectively by Hewson (1970(7: 187). Aneura pusilla Stephani (Stephani & Watts 1914: 96) Proto: 'Australia, (Old Railway Cutting, Blackheath): Watts legit, 1051.' Label: New South Wales: old railway cutting, Blackheath, W.W. Watts 1051, Jan 1911 [3 packets in one specimen folder]. Types: G (11049: lectotype), NSW (H242: isolectotype). Comment: Listed as a synonym of Riccardia crassa (Schwaegr.) Carrington & Pearson (Hewson 1970a: 104). Taxon reinstated and transferred to Riccardia; lectotypified by Brown & Braggins (1989: 82). Aneura rodwayi Hewson (1970a: 188) Proto: 'Holotype—Hartz River, Mt. Hartz National Park, on clay soil near river bank, Tas., Hewson, 215, 9.1963, (NSW.).' Label: Tasmania: Hartz River, Mt Hartz, on soil near river, H.J. Hewson H63.33.215,10 Sep 1963. Type: NSW (179241: holotype). Comment: Name in current use. Aneura rufescens Stephani (Stephani & Watts 1914: 96) Proto: 'Australia, New South Wales, (National Pass, Wentworth Falls): Watts legit, 1124/ Brown, Ramsay & Seur, Hepatic types 71 Label: New South Wales: National Pass, Wentworth Falls, W.W. Watts 1124, Sep 1912 [2 packets in one specimen folder]. Types: G (11050: lectotype), NSW (H243: isolectotype). Comment: = Riccardia crassa fide Hewson (1970b: 104). Lectotypified by Brown & Braggins (1989: 85). Aneura rupicola Stephani (1917: 41) Proto: 'Australia: New South Wales (Watts legit.).' Label: New South Wales: 1. Shaws Bay, log, W.W. Watts 579, 20 Sep 1901; 2. Shaws Bay, Ballina, Richmond] R[iver], log, W.W. W[atts] 579, 20 Sep 1901 [both packets are in one specimen folder]. Types: G (11051 ex Hb Levier 3388), NSW (H248). Comment: G specimen annotated 'Original. Shaw's Bay, east Ballina. leones Hep. 520’. = Riccardia rupicola (Stephani) Hewson; G 11051 cited as holotype and 'NSW 579' as isotype by Hewson (1970b: 99). Aneura walesiana Stephani (Stephani & Watts 1914: 97) Proto: 'Australia, New South Wales, (Blackheath): Watts legit, 1023.' Label: New South Wales: Horseshoe Falls, Blackheath, wet cliff, W.W. W[atts] 1023, 7 Jan 1911 [3 packets in one specimen folder]. Types: G (lectotype), NSW (H244: isolectotype). Comment: = Riccardia crassa fide Hewson (1970b: 104). Lectotypified by Brown & Braggins (1989: 85). Aneura wattsiana Stephani (1917: 46) Proto: 'Australia, New South Wales (Watts legit.).' Label: New South Wales: 1. E Ballina, [Richmond River], on tree fern, wet places, W.W. Wlattsl 237, 2 Mar 1901 (NSW H252); 2. label identical, no NSW number. Types: G (11057 ex Hb Levier 3018), NSW. Comment: No specific locality or specimen number cited in protologue. Specimen in G (11057) has the following data 'W.W. Watts 237, 1901, Ballina, New South Wales, leones Hep. 299'. = Riccardia wattsiana (Stephani) Hewson; G 11057 cited as holotype and 'NSW 237' as isotype by Hewson (1970b: 91). The un-numbered NSW specimen has a second Watts collection attached to it (Watts 346). Archilejeunea australis Stephani (1911: 734) Proto: 'Australia, New South Wales.' Label: New South Wales: back of Wickham's, nr. Ballina, tree, W.W. Watts 415, 3 June 1902. Types: G (ex Hb Levier 3483), NSW. Comment: = Thysananthus australis (Stephani) Thiers & Gradstein; specimen in G (ex Hb Levier 3483-Watts 415) cited as the holotype by Thiers & Gradstein (1989: 66); no reference is made to the G specimen being labelled as the type by Stephani. Telopea Vol. 5(1): 1992 72 Balantiopsis decurrens Stephani (Stephani & Watts 1914: 98) Proto: ‘Australia, (Wyong): Watts, 985.' Label: New South Wales: gully south of house, 'Kingwell', Wyong, W.W. Watts 985, Nov 1907 [3 packets in one specimen folder]. Types: G, NSW. Comment: = B. diplophylla (Hook. f. & Taylor) Mitten fide Engel (1968: 109, 111). Balantiopsis hastatistipula Stephani (Stephani & Watts 1914: 98) Proto: 'Australia, (Blackheath), Watts, 1052.' Label: New South Wales: Blackheath, old railway cutting, W.W. Watts 1052, Jan 1911 [2 packets in one specimen folder]. Types: G, NSW. Comment: = lsotachis sp. fide Engel (1968: 118). Balantiopsis kingwella Stephani (Stephani & Watts 1914: 99) Proto: 'Australia (Kingwell, Wyong): Watts 943.' Label: New South Wales: gully south of house, 'Kingwell', Wyong, W.W. Watts 943, Nov 1907 [3 packets in one specimen folder]. Types: G, NSW. Comment: = B. diplophylla fide Engel (1968: 109, 111). Balantiopsis subkingwella Stephani (Stephani & Watts 1914: 99) Proto: 'Australia (Kingwell, Wyong): Watts 945, 971a Label: New South Wales: 1. 'Kingwell', Wyong, W.W. Wlattsl 945, 20 Aug 1909; 2. 'Kingwell', Wyong, gully south of house, W.W. Watts 971a, Nov 1907 (NSW H285) [2 packets in one specimen folder]. Types: G, NSW. Comment: = B. diplophylla fide Engel (1968: 109, 111). Watts 971b is the type of Mas- tiogobryum corbieri Stephani, see below. Brachiolejeunea robusta Stephani (1912: 141) Proto: 'Australia. New South Wales.' Label: New South Wales: 1. Hopkins Creek, Stanwell Park, rocks, W.W. W[atts] 830, May 1905; 2. Hopkins Creek, Stanwell Park, [W.W.I Watts 830, May 1905. Types: FH, G (ex Hb Levier 4882), NSW, U. Comment: Specimen in G (Hb Levier 4882-Waffs 830) cited as holotype and FH as isotype by Thiers & Gradstein (1989: 64) but there is no supporting discussion. Re¬ duced to synonymy under Spruceanthus thozetianus (Gottsche & F. Muell.) Thiers & Gradstein. Chiloscyphus maximus Stephani (Stephani & Watts 1914: 103) Proto: 'Australia (Etta's Glen, Black Spur, Viet.): Watts, 968.' Label: Victoria: Etta's Glen, Black's Spur, Healesville, W.W. Watts 968 & 968a, 21 Dec 1906 [5 packets in one specimen folder]. Types: G, NSW. Brown, Ramsay & Seur, Hepatic types 73 Comment: The NSW specimen consists of 5 small packets variously labelled Watts 968, Watts 968a or Watts 968 + 968a. All are annotated 'Chiloscyphus maximus St. sp. n. 1913' in Watts' hand. Apparently a large variety of C. coalitus (Hook.) Nees in Gottsche, Lindenb. & Nees (Scott 1985: 133). Chiloscyphus montanus Stephani (Stephani & Watts 1914: 103) Proto: 'Australia (Neate's Glen, Blackheath): Watts, 927.' Label: New South Wales: Rotunda, Neate's Glen, B[lack]heath, on damp ground base of cliff, W.W. Watts 927, 7 Apr 1903 [3 packets in one folder]. Types: G, NSW. Comment: = Geocalyx caledonicus Stephani fide Grolle (1979). Fimbriaria dioica Stephani (Stephani & Watts 1914: 104) Proto: 'Australia, N.S. Wales (near Gladesville): Watts 1095.' Label: New South Wales: off Pittwater Road, Gladesville, W.W. Watts 1095 & [J.F.] Shirley, Nov 1911 [3 specimens in one folder]. Types: G, NSW. Comment: Fimbriaria is an illegitimate name but this taxon has not been formally transferred to Asterella, where it belongs. This group is under revision. Fossombronia grossepapillata Stephani (1917: 73) Proto: 'Australia (Watts legit.).' Label: New South Wales: park, Young, (mining holes), W.W. Watts 988, Sep-Oct 1906 [3 packets in one folder]. Types: G (22171), NSW. Comment: G 22171 (Watts 988) designated as type specimen by Scott & Pike (1988: 196). Fossombronia integrifolia Stephani (1917: 73) Proto: 'Australia orientalis (N.S. Wales).' Label: New South Wales: 1. Parramatta, E. Cheel (Hb Watts 799), 1 Oct 1910 [2 pack¬ ets]; 2. Rookwood, Sydney, E. Cheel (Hb Watts 812), 5 Aug 1900 [2 packets]. Types: FH (812), G (22174), NSW. Comment: Specimens of Watts 812 held at FH and G designated as the type by Scott & Pike (1988: 197). Fossombronia wattsii Stephani (1917: 75) Proto: 'Australia, New South Wales (Watts legit.).' Label: New South Wales: The Gap, near Young, W.W. W[atts] 881, 26 Sep 1905. Types: G (Hb Levier 5155), NSW. Comment: No locality or specimen number cited in protologue but G specimen of Watts 881 cited as type in Bonner (1965: 221). Type material (at NSW) checked and studied by Pike: 'abnormal spores present 8/3/82'. 74 Telopea Vol. 5(1): 1992 Frullania asperifolia Stephani (Stephani & Watts 1914: 105) Proto: 'Australia (Yarrangobilly Caves & Mount Wilson): Watts 1093, etc).' Label: New South Wales: 1. Yarrangobilly Caves, W.W. Watts 923, Jan 1906; 2. 'Yar- rawa', Mt Wilson, W.W. Watts 1063,10 Jan 1911; 3. Denmarque, Mt Wilson, W.W. Watts 1078,13 Jan 1911; 4. Falls beyond Kiandra road, Yarrangobilly Caves, W.W. Watts 926, Jan 1906; 5. Yarrangobilly [Caves], gully beyond hostel, W.W. Watts 922, Jan 1906; 6. Avenue, Mt Wilson, W.W. Watts 1093, 11 Jan 1911. Types: G (18193), NSW. Comment: Although two localities were cited in the protologue only one number (Watts 1093) was given and this is cited as the type by Bonner (1965: 239). Hattori (1979a) cites the type as 'G18193, the Avenue, Mt Wilson, 3500 ft on tree Jan 11,1911, leg. W.W. Watts (Hep. Aust. ex Herb. Watts 109 [= 1093])' and considers this taxon conspecific with F. falciloba Taylor ex Lehm. Frullania australis Stephani (1911: 659) Proto: 'Australia orientalis. New South Wales.' Label: New South Wales: E Ballina, tree, W.W. Watts 117, 14 Aug 1900 [3 packets in one folder]. Types: G (18194 ex Hb Levier 2638), NSW. Comment: G 18194 is cited as the type by Bonner (1965: 241), without the Watts collection number. Known only from the type collection (Hattori 1979b: 123). Frullania belmorensis Stephani (Stephani & Watts 1914: 106) Proto: 'Australia (Belmore Falls): Watts, 931.' Label: New South Wales: Belmore Falls, W.W. Watts 931, 5 Oct 1908 [3 packets in one folder]. Types: G, NSW. Comment: Known only from the type collection (Hattori 1979b: 126). Frullania difficilis Stephani (1889: 160) Proto: 'Australia, New South Wales.' Label: New South Wales: Park, Gray's Island, Richmond River, W.W. Watts 125,6 May 1900. Types: G (ex Hb Levier 2632), MEL 61558, NSW. Comment: No specimen number or specific locality is cited in the protologue but the type is cited by Bonner (1965: 288) as 'New South Wales, Richmond River, Pearce's Creek, 2 VIII 1900, W.W. Watts s.n., Hb Levier 2632 in Hb G'. Hattori (1987: 553) lists a number of other possible syntypes (including the above NSW specimen) and places F. difficilis in synonymy under Frullania (Trachycolea) seriata Gottsche ex Stephani. Frullania elongata Stephani (1910: 423) Proto: 'Australia orientalis.' Label: New South Wales: Pearce's Creek nr R. Trimble's, bark of old tree stump, W. W. Watts 96, 2 Aug 1900. Types: G, NSW. Brown, Ramsay & Seur, Hepatic types 75 Comment: Watts 96 is cited by Bonner (1965: 296) as the type. = F. rubella fide Hattori (1979n: 354); = F. rubella Gottsche ex Stephani var. elongata (Stephani) Hattori fide Hattori (1983: 165). Frullania excisula Stephani (Stephani & Watts 1914: 107) Proto: 'Australia, N.S. Wales (Mount Wilson and Blackheath): Watts 1030, 1071/ Label: New South Wales: 1. Horseshoe Falls to Jungle, Blackheath, (rock by track), W. W. Watts 1030, 7 Jan 1911 [2 packets]; 2. Yarrawa, Mt Wilson, W.W. Watts 1071,11 Jan 1911 [3 packets]. Types: G, MEL, NSW. Comment: Bonner (1965: 229) cited Watts 1017 [= 1071] as the type. The NSW speci¬ mens of both Watts 1030 and 1071 are annotated by S. Hattori Apr. 1986: 'not type of F. squarrulosa but syntype of F. excisula'. F. excisula seems to be conspecific with Frullania falciloba (Hattori 1979a: 336). Frullania ferdinandi-muelleri Stephani (1910: 417) Proto: 'Australia, New South Wales.' Label: New South Wales: Tintenbar(?), Richmond] RJiver], tea tree bark, W.W. Watts 153, 2 Aug 1900. Types: G (18211 ex Hb Levier 2628), NSW. Comment: No information on locality, date, collector or specimen number(s) appears in the protologue but Hattori (1979a) describes this species as known only from the type collection and cites G 18211 as the type. Frullania filipendula Stephani (1910: 419) Proto: 'Australia, Queensland, New South Wales.' Label: New South Wales: E Ballina, base of saplings and on ground, heath, W.W. Watts 468, 17 June 1902 [2 packets in folder]. Types: G (18212 ex Hb Levier 3416), NSW. Comment: No information on locality, date, collector or specimen number(s) appears in the protologue but Bonner (1965: 304) cites this specimen (G 18212) as the type. Seems to be conspecific with F. rubella (Hattori 1979a: 340). Frullania howeana Stephani (Stephani & Watts 1914: 107) Proto: 'Lord Howe Island. (Watts 62).' Label: New South Wales: near Face, east side of Mt Lidgbird, L[ord] HJowe] IJsland], W.W. Wlatts] 62, 27 July 1911 [2 packets in folder]. Types: G (18215), NSW. Comment: Endemic, known only from the type collection (Hattori 1979a: 346). Frullania minutistipula Stephani (Stephani & Watts 1914: 108) Proto: 'Australia (Rodriguez Pass): Watts 1001b,1001c.' Label: New South Wales: Rodriguez Pass, Blackheath, W.W. Watts 1001b & c, 4 Jan 1911. Types: G, NSW. Comment: NSW specimen has been annotated (by Watts?): 'mixed with Frullania gracilliina sp. n. and Radula wattsiana'. = F. rostrata (Hook. f. & Taylor) Hook. f. & Taylor ex Gottsche, Lindenb. & Nees fide Hattori (1979a: 352). 76 Telopea Vol. 5(1): 1992 Frullania simmondsii Stephani (Stephani & Watts 1914: 109) Proto: 'Australia (near Brisbane): Simmonds leg. (Watts 1110).' Label: Queensland: near Brisbane, J.H. Simmonds, Hb Watts 1110,27 Aug 1887. Types: G, NSW. Comment: NSW specimen annotated as isotype by Hattori. Known only from the type collection (Hattori 1979b: 143). Frullania subtropica Stephani (1910: 416) Proto: 'Australia, New South Wales.' Label: New South Wales: off Tintenbar road, 3 miles [5 km] from Ballina, Richmond River, tree, W.W. Watts 107, 30 June 1900. Types: G (18230 ex Hb Levier 2602), NSW. Comment: Protologue does not include locality, date, collector or specimen informa¬ tion but G 18230 is cited as the type by Bonner (1965: 440). Frullania variabilis Stephani (1910: 420) Proto: 'Australia, New South Wales.' Label: New South Wales: Cambewarra Mtn, W.W. W[atts] 628, 22 May 1903. Types: G (ex Hb Levier 4416), NSW. Comment: Protologue does not include locality, date, collector or specimen informa¬ tion but the above specimen in G (Hb Levier 4416) is cited as the type by Bonner (1965: 462). Known only from the type collection; probably a form of F. pycnantha (Hook. f. & Taylor) Taylor ex Gottsche, Lindenb. & Nees (Hattori 1979b: 149). Haplomitrium intermedium Berrie (1963: 191) Proto: 'Between one and two miles along the road from Bilpin to Mount Irvine, Blue Mountains, N.S.W. Holotype: Sydney University Herbarium, Bryophyte No. 2/62. Cotype: Professor R.M. Schuster Collection No. 50,616.' Label: see below Types: SYD (holotype; not located). Comment: Probably on loan, but record not found. Isotachis grandis Carrington & Pearson (1888: 1041) Proto: 'On wet rocks, Lawson, Blue Mountains, June, 1884.' Label: New South Wales: 1. Lawson, Blue Mountains, on wet rocks, ]T. Whitelegge], June 1884; 2. Lawson, no 14 liverwort, [T. Whitelegge], June 1884. Types: BM, MANCH?, NSW. Comment: The former NSW specimen has been further annotated by Whitelegge: '17 Isotachis grandis n. sp. (small portion of stem) plXXVI 34 TW W.H.P.' and the latter has the additional information: '17 N. sp. No 34 P. Isotachis grandis'. Probably no. 34 of a batch sent to Pearson by Whitelegge. Isotachis terricola Stephani (Stephani & Watts 1914: 10) Proto: 'Australia (Blackheath): Watts, 1016.'