Elytra,Tokyo,35(1):194-204,May30,2007 A Revision of theGenusFo,rmosotoxotus (Coleoptera, Cerambycidae,Apatophyseinae),withDescription ofaNewSpecies fromSikkim Nobu0OHBAYASHI EntomologicalLaboratory,FacultyofAgriculture,EhimeUniversity, Tarumi,Matsuyama,790-8566Japan Abstract ThecerambycidgenusF,ormosotoxotus isrevised. For,nosoto:xotus masataka1sp nov. isdescribed fromSikkim, andF fulvopi1osus HAYAsHI, 1979 is synonymizedwithFmalayanusHAYAsHI,1977. Thesystematicpositionofthegenus isdiscussedanditisplacedinthesubfamilyApatophyseinae. Introduction OnmyshorttriptoParismadein2005, I foundaveryoldspecimenbelongingto thegenusFormosotoxotusinthecollectionoftheMuseumnationald'Histoirenature11e. Afteracarefulexamination, itwasconcludedthatthespecimenbelongedtoaspecies new to science. Alongwith descriptionofthenewspecies, I takethisopportunity to revisebrieflythegenusFormosotoxotusandbestowsomeconsiderationsonitssystematic position. Taxonomicplacement of thespeciesofFormosoto:x;otus hasfrequentlychanged. KANO (1933) describedthefirstspecies 「oxotinusauripi1osusfromTaiwanasaclose relativeofTo:)cotinusrein1i HEYDEN. Inthesameyear,MATSUSHITA(1933) described Artelidaaslatica fromTaiwan(thegenusArtelidahadbeenuseduntil thenexclusively forMadagascanspecies). GRESSITT (1951) placedT1oxotinusauripi1osus in the genus To:x;otus (subgenus To:xotinus)oftheStenocorini. SimultaneouslyheerectedanewgenusParanthophylax (in Xy1osteini) foranewspecies,P.sericeus (designatedasthetypespecies) andP aslaticus, transferredtothisgenusfromArtelida. Paranthophylaxiscurrentlyregarded as a juniO「 Synonym of the genus Trypogeus LACORDAIRE. In l960, HAYAsHI synonymizedArtelidaaslatlcawithToxotinusauripi1osusandestablishedanewgenus F,ormosotoxotusfor lt. Sincethen,fivemorespecieswereaddedtothegenusF,ormoso- t();)cetus, viz., F malayanusHAYAsHI, 1977and F fulvopi1osusHAYAsHI, 1979, from Malaysia, F. uenoi N. 0HBAYAsHI, l995, from Kalimantan, Indonesia, F takaoi NIISATo, 1996, from North Vietnam, and F nobuoi VIvEset NIIsATo, 2006, from Nepal. Up tonow,all theauthorsregardedthosespeciesasmembersof thesubfamily RevisionoftheGenus.Formosotoxot11s 195 Lepturinae except JENIS (2001) who illustrated an unidentified Malayan species of Fo, rmosoto:ofetus and placed it, without proper explanation, in the Apatophyseinae, togetherwithmanyMadagascanspecies, theAfricanDorcasomusSERvILLEandCapo- te;xotus rugosus TIPPMANN, and theOriental generaProta-x;isGAHANand Trypogeus LAcoRDAIRE. The subfamily Apatophysinae LAcoRDAIRE was erected by DANILEvsKY (1979) forasinglegenus,ApatophysisCHEvRoLAT,mainlybasedon the larval morphology (original LAcoRDAIRE's Apatophysides included also theSouth AfricangenusPachytt'conTHOMSONandtheOrientalTrypogeuswhoselarvaewereand still areunknown). Thecorrectspellingof thesubfamilynameapparentlyshouldbe Apatophyseinae, first used by LoBANov et a1. (1981). So far there have been few external adult characters distinguishing theApatophyseinae from other subfamilies. However, thesubfamilycouldbecharacterizedbyacombinationofstridulatoryfileof mesonotum, hind wing venation andmale genitalia (personal unpublished results). Basedonmystudyofhindwingvenationandmalegenitalia,Iconcludethat thegenus Fo,rmosoto:x:otus isclosely related toApatophysls, and thusit belongsto thesubfamily Apatophyseinae. Thisshortpaper isdedicatedtothememoryof thelateProf.Dr.MasatakaSAT0, whowasoneofthemostprominentJapanesecoleopterist,andalsohadbeenmyclosest seniorlikeabrother sincefiftyyearsago. IwishtothankPetrSvAcHA(CzechAcademyofScience,CeskeBudejovice) for hisusefuladviceandcritical reviewof themanuscript. My thanksarealsoduetothe followingentomologists for their kindhelp, usefulsuggestionsorofferof invaluable specimens for study: GianfrancoSAMA, Mikhail L. DANILEvsKY, EduardVIVESand Tatsuya NIIsATo. I was also much indebted to Thierry DEuvE and Azadeh TAGHAvIAN of the Museumnational d'Histoire nature11e, Paris for their kind help extended tomeatmyvisittothemuseum. SubfamilyApatophyseinaeLAcORDAIRE,1869 ApatophysidesLAcoRDAIRE,1869,234. Apatophysinae:DANILEvsKY,1979,827. Apatophyseinae:LoBANoveta1.,1981,786,794. - SvAcHA&DANILEVSKY,1988,125. - SV CHA et al., 1997,364, fig 55. Notes. After DANILEvsKY (1979) erected the subfamily ApatophySinae LAcoRDAIRE for receiving the genus Apatophysls, SVACHA (in SVACHA & DANILEvsKY,1987)madethefollowingcomment:“ThegenusApatophyslshasnodi「cot relatjonto theLepturinae,andithasbeenplacedinaseparatesubfamilyApatOphy- sejnae(DANILEvsKY,1979). _ DespiteDUFFY(1957,1980), thegenusDO「CaSOmuS canbeonnoaccountretainedinLepturinae,and,althoughdifferinginsomecharacters fromApatophysis, itshouldbeprovisionallyincludedintheApat,ophyseinae. ...The wrjtersuspects all theLepturine-1ikeformsoccurringin theEthiopianRegion (inCl. Madagascar) tobeofasimilar taxonomicposition” AccordingtoP.SVACHA (Pe「S・ 196 Nobuo OHBAYASH] Figs.1-8. HindwingvenationofthesubfamilyApatophyseinae. - 1,ApatophysisbarbaraLucAs 2,ApatOphySSiSlnCiaSEMENOVTIAN-SHANsKIJ;3,Artelidasp;4,Mastododersnodjco11jsKLUG;5 Formosoto)cetusaurtpi1osa;6,Fmalayanus;7,F nobuoi;8,TripogeuscabigasiVIvEs. . Comm),afterinclusionofDorcasomustheApatophyseinaeshouldberenamedDorca_ SOminaeforpriorityreasons(DorcasomideswereestablishedinLAcoRDAIRE,1869a, 456).AfterdiscussionswithsomeEuropeanentomologists,itisbecomingclearthatthe majorityof theso-ca11edlepturine-1ikeformsofMadagascarshouldbeincludedjnthjs Subfamily,buttherearedifferentopinionsconcerningthepositionofthegenusDorca_ somus distributed in continental Africa. Not havingexaminedspecimensof I)orca_ SOmuS, ICannot contribute tosolving theproblemandtherefore I retain thename ApatophyseinaeuntilthechangetoDorcasominaeisformallypublished. ConcerningtheAsiangeneraof thissubfamily,VIvEs (2006) describedanew 9enuSBO「neophystsfromSabah,Malaysia,andalsoupgradedtwoformersubgeneraof APatOPhySSi togenericstatus:ParatophysisGREssITTetRoNDoN,1970,andEpjtophysls GRESSITTetRONDON,1970. Inaddition,thegenusTrypogeusshouldbeincludedinthis Subfamily(E.VIVES,pers.comm). Astheresult,sixgeneraofApatophyseinaeoccur Revision of the GenusF,ormosotoxotus 197 in Asia: atopayszs, formosotoxotus, Paratophysls,◆pftophysls, orneop ysisand 「rypo- geus. HindwingvenationofthegeneraApatophysis,Artelida,Mastododera,Formosotoxo- tusandTripogeusareillustratedinFigs.1-8forcomparison. GenusFlormosotox:otusHAYAsHI,1960 Fo,mosotoxotusHAYAsHI,1960,1;typespecies:ArtelidaaslatlcaMATSUSHITA,1933=ToxotitltSl au「lP11oSuS KANO,1933. - HAYAsHl&VILLIERS,1985,27,31. Bodystout,short,entirelycoveredwithadpressed1onghairs.Headshort;mandi- bleslargeandsharp;genaaslongasthewidthacrosseye;antennainsertedlaterallyat somedistancebeforeeye;eyecoarselyfacetedandemarginatebehindantennalinsertion; temperagraduallynarrowedposteriorly,notconstricted. Pronotumwithconicallateral tuberclesandtwopairsofdiscaltubercles,theanteriorpairofwhichislarger thanthe posterior one; presternal processnarrow andslightly dilated at apex; acetabula of procoxaeclosedornarrowlyopenedposteriorly;mesosternumwithoutstridulatoryfiles andwithacompletesubcuticular longitudinalmedianblack linevisiblethrough the translucentcuticle;surfaceofcuticlewithouttransversestriation, indistinctlyreticulate. Elytrabroaderthanpronotum,almosttwiceaslongasbasalwidth,almostparallel-sided orslightlynarrowedposteriad. Hindwingvenationasin Figs 5-7. Legs rather long; tibiadilated andusually flattened apically; hind tarsuswith first segment as longas secondandthirdcombined. Medianlobeofmalegenitalialongerthantegmen,widened frommedianstruts indorsal view;medianstrutslonger thanhalf of total lengthof median lobe;median foramenroundlyopened. Notes. ThisgenushasverycloserelationwiththegenusApatophysls, inparticular bythefeaturesofhindwingvenation, thestructureofmalegenitalia,laterally inserted antennalscape,andtwopairsofdisca1tuberclesofpronotum,but it isdistinguishableby rather short antennae, irregularlyarrangedpubescenceon dorsalsurface, hind tarsus with first segment not longer than second and third combined, abdomen of female entirelycoveredbyelytra, andsoon. Fo,rmosotox:1otusauripi1osus (KANO,1933) (Figs5,9,13-14) T,oxotinusaltripi1os1lsKANO,1933,263; typelocality:Tattaka(7,300ft.inalt)nearMusha,CentralFormosa (Cerambycinae,sensulate). - TAMANUKI,1939,69, fig 25. (Senocorini) Arte11daasiatlca MATSUSHITA,1933,172;typelocality: Hozan,Formosa. (Toxotini) Paranthophylaxaslattcus:GREssITT,1951,50,51. (Xy1osteini) T,o_xot11s(T1oxotinus)atlriplIos11s: GREssITT,1951,56,58. (Stenocorini) Specimensexamined. ld1',Sziuaniakou,Taichung,Taiwan,25-V-1999, M.SAT0 leg; l Anmashan, Taichung, Taiwan, 25~28-IV-2002, N. 0HBAYAsHI leg; l , , Meifeng,Nantow,Taiwan,25-IV-1976,K.MAsuMoT01eg;1早,Sungkang - Meifeng 198 Nobu0OHBAYASHI Figs 9-12. Male genitaliaof Formosoto;xlotusspp. - 9, F aurlpi1osus (Sziuaniakou, Taichung Taiwan);10,F malaJ,anus (TanahRata,CHL,Malaysia);11,F takao1(Mt.Phu-Pan, Laos) 12,F masatakaisp nov. (Sikkim). (2,044-2,127m), Nantow, Taiwan,19-V-1969,S.HIsAMATsuleg; l早,PiluShenmu (2,200m), Hualien,Taiwan,25-VI-1989,M.SAT01eg. F1ormosotoxotusmalayanusHAYAsHI, 1977 (Figs 6,10,15-20) FormosotoxotusmalayanusHAYAsHI,1977,95; typelocality:CameronHighlands,Pahang,Malaysia.(Xy- Iosteini). - HAYAsHI&VILLIERS,1985,33,pl 5, fig.13. (Xylosteini) F1ormosotoxotus fulvopi1osus HAYAsHI, 1979, 3; type locality: Cameron Highlands, Tana Rata, Pahang, Revision of the GenusF1ormosoto)cetus 199 Malaysia. (Xy1osteini). - HAYAsHI&VILLIERS,1985,33,pl 5,fig.14.(Lepturinae). Sy'l・'lOV・ Formosotoxotltssp:JENlS:,2001,34(pl 20),fig.137/101.(Apatophyseinae) HAYAsHI (l977) firstdescribedF.' malayanus fromtheCameronHigh- Notes. lands. Twoyearslaterin1979,headdedanothernewspecies,F fulvopi1osusfromthe samelocality. Accordingtotheoriginaldescriptions,bodylengthsoftheformerspecies are10mm(theholotypemale)and15mm(aparatypefemale),andthatofthelatter speciesare17mm(holotypemale)and21mm(aparatypefemale). In1985HAYASHI &VILLIERSre-describedthesetwospecies,anddistinguishedthembythefollowingkey: 1. Bodybrownishred,coveredwithfulvoushairs; fronsseparatedfromclypeusbyan arcuategroove; antennaeshorter than body inmale, gena shorter than eye- diameter;elytraabouttwiceaslongasbasalwidth,discrathersimple.10-15mm F malayanus - Bodydark reddishbrown,coveredwithpalefulvoushairs; fronsseparated from clypeusbyastraighttransversegroove;antennaefairlylongerthanbodyinmale, genaaslongaseye-diameter;elytraalittlelongerthantwiceaslongasthebasal width disc with four disca1costae. 17-21 mm F fulvopiiosus My examinationof aseries ofspecimenscollectedon theCameron Highlands showedthat thedifferencesbetweenthetwospeciesdescribedbyHAYASHIareintra- specificvariations. Thedifferenceofantennallengthandelytra1proportionindicatedin the key is not a specific featurebut asexual difference, because theholotype of Folrmosoto:xlotusmalayanusdescribedasmaleinhisoriginaldescriptionwasafemale. Other differential characteristics also depend on thesexual dimorphism, or on the differenceofbodysize. Body lengthsof thespecimensexaminedarequitevariableas follows: 6.8, l l.2, 11.5, lt.9, 12.0, 12.7, 16.0, 17.1mm in male and 7.5, 8.6, 9.9 (holotypeofF malayanus),11.0,11.2mminfemale,respectively. Disca1costaeofthe elytraaredistinctinlargespecimens,andtheconditionof fronto-clypea1suturediffers withindividuals. Asaresult,F fulvopi1osusishereplacedasajuniorsynonymofF malayanus. Specimensexamined. 1早 (Holotype), Tana Rata, Malaysia, 22-I-1976, Y. KIYoYAMAleg. (FolrmosotoxotusmalayanusHAYASHld'l) (M.HAYASHICo1l.0MNH [98-32]); 6 4早早, Mt. Jasar, Cameron Highlands, Pahang, Malaysia, III-1985, , nativecollector; l TanaRata,C.H.L.,Malaysia,16-I-1980,coll.H.DETANI; l , , TanaRata, Malaysia, 14-I-1980, coll.N.NIsHIKAwA. F,ormosotox::otusuenoi N. OHBAYASHI, 1995 (Figs 21-22) F,ormosoto;x;otus ue,101 N.OHBAYAsHl,1995,437, figs. l-6; typelocality: Mt. Berangin,W. Kalimantan, Indonesia. (Lepturinae) Specimenselcamined. 1 (Holotype), Mt Berangin,W. Kalimantan, Indonesia, 200 NobuoOHBAYAsHI VIII-1992,Collectedbyanative;1早(Allotype),Mt.Saran,W.Kalimantan, Indonesja, IX-1992,collectedbyanative. Formosoto;c'etustakaoiNIIsATo,1996 (Figs.11,23) Fo'「mOSOto:ofetustakao1NIISATo,1996,101,figs.1&2;typelocality:NearSapa,LaoCalProvince,North Vietnam. (Stenocorini) Notes. ThisspecieswasfirstdescribedfromasinglespecimencollectedinNorth Vietnam,andasecondspecimenisnewlyrecordedfromnorthernLaos. Thespecimen f「omLaosshowssomedifferencesintheproportionofpronotumwhichisslenderer than thatoftheVietnamesespecimen. Forexample,pronota11engthis0.90timesaslongas maximumwidthofpronotuminsteadof0.84,andalsopronota11engthIsl27timesas longasapicalwidthof pronotum, insteadof 1.14. Until further specimensbecome available,thisdifferenceisconsideredintraspecificor localvariation. Specimensexamined. 1 (Holotype),nearSapa, LaoCalProvince,NorthViet- nam,25-III-l995,N.KATsURAleg; ld'l,Mt.Phu-Pan,BanSaleui,XamNeuaProv., Laos,28-III-2005,J.YAMAsAK01eg. Fo,rmosoto;x:otusnobuoiVIvEset NnsATo,2006 (Figs7,24-25) Flo「mOSOtoC)etusnobuo1VIVESetNIIsATo,2006,273,figs.1&2;typelocality:Taplejung,ca2,000minalt NechiProvince,E.Nepal.(Xy1osteini) Specimensexamined. 8 39-9 (Paratypes), Taplejung, ca 2,000m in alt. , NechiProvince,E.Nepal, l3~18-VI-2000,localcollectors. Formosotox;:otusmasatakaisp nov. (Figs.10,26) Male. Lengthl3.5mm,width4.5mm. Bodyrobust, rathershort;colorchest- nutbrown;maxillarypalpusreddishbrown;mandibles,lateralmarginoflabrum,and marginal areaofscutellumblackish;antennayellowishbrownexcept for scapeand pedicel,whicharedarkerthantheremainingsegments;legsyellowishbrownexcept for darkenedapicalareaof femoraand first tarsalsegments.Bodyentirelyclothedwith adpressedpaleyellowpubescence,ratherlongonhead,shortandfineonpronotum,and onelytraarrangedinwhorlscausingmarbledappearance. Headincludingmandibles longer thanpronotum, gradually narrowedposteriad behindeyes;mandiblesstout,long,glossydorsally,protrudingmediallyandprovided withhairsonoutersidesofbasalhalf;labrumtwiceaswideaslong,gentlyarcuateat sideandsubtruncateat apicalmargin,clypeusnarrowlytransverse; fronsdepressed RevisionoftheGenusFormosotoxotus 201 / / f 、 本 20 、 , , 、 / / l 1 21 22 , Figs.13-26. Habitus of F,ormosotoxotus spp. - 13, F auripi1osus (Taiwan), male; 14, ditto, female;15-17,Fmalaya川ts (CHL,Malaysia),male;18-20,ditto,female(20=holotype);21,F. uenoj(w.Kaljmantan,Borneo),holotypemale;22,ditto,paratypefemale;23,F takaoi (Laos); 24, F nobuo1(Nepal), paratype male; 25, ditto,paratype female;26, F masatakal SP nov. (Sikkim), holotype. 202 NobuoOHBAYASHI mediallywithelevatedlateralridges;agrooveextendingfrombaseoffronstovertexbut not reachingocciput; antennal tuberclesstronglyelevatedandextending forwards to lateral ridgesof frons;vertexslightlyswollenonbothsidesofmediangroove;occiput transverselydepressed. Eyesverycoarselyfaceted,large,distinctlyprominent laterad, almostovalandslightlyemarginatebehindantennalinsertions. Antennaslenderthough eachsegmentisthickenedapically,flattenedfromfifthtothelastsegments,notreaching theapexofelytra;scapegentlycurved, thickenedapicadandproducedocto-apically; thirdsegmentslightlyshorterthanscapeandlongerthan fourth,seventhsegment the longest;relativelengthofeachsegmentasfollows:26:10:24:21:28:27:29:25:25: 21:27. Maxillarypalpuswithlastsegmentwidestnear themiddle,2.6timesaslongas wide,slightlynarrowedtoward theroundlytruncateapex. Pronotumtrapezoidalwithconicalandnearlyrectangular lateral tubercles,widest acrossthetubercles,1.42timeswider thanlong,narrowlymarginedatapexandbase; basel20timesaswideasapex;discprovidedwithtwodistinctpairsofobliqueanterior androundedposteriorswellings. Presternalprocessstronglynarrowedbetweencoxal cavities, then triangularly dilatedapically; acetabulaofprocoxaeclosedposteriorly; mesosternumwithoutstridulatoryfiles,withablackmedianlongitudinal line;scutellum nearly tongue-shaped. Elytrawidestathumeri, twiceaslongaswide,slightlynarrowed fromhumeri to apicalfifth,thengentlyroundedtobluntsuturalapex;discslightlyconvexatbothsides behindscutellum, then attenedaroundbothsidesalongsuture,denselycoveredwith shallowandminutesetigerouspuncturesthroughout. Legs stout andmoderate in length; femora clavate; all tibiaecompressed and widenedapicadwithroundedouterangles;hindtarsuswiththefirstsegmentaslongas second and third combined. Malegenitalia:- Medianlobe127timesaslongastegmen,slightlycurved,ventral sidebisinuateinlateralview;dorsalplatewiderthanventralplate,graduallynarrowed apicadwithroundedapexinapicalthird,notreachingtheapexofventralplateindorsal view; median struts 0.62 times as longasthe total lengthofmedian lobe;median foramenopenedasroundemargination. Tegmenwithlateral lobesshort,one-sixthas Ion9asitstotallength;apicalareaoflaterallobesprovidedwithmoderatelylongsetae; roofoncedeeplyemarginatebehindthebaseoflaterallobes;ringedpartthjck,almost parallel-sided, thennarrowed towardunconnectedbase. Holotype. ,Sikkim,1890,HARMAND(MUS.HIST.NAT). Nofurtherdata are available. The holotype ispreserved in the collection of the Museum natjona1 d'Histoirenature11e,Paris. Etymology. Thespeciesisdedicated tothelateProf.Dr. MasatakaSAT0 for the memoryofhisgreatcontributiontotheentomology. Notes. This new species can bedistinguished from the knownspecies by the followingkey. RevisionoftheGenusFo,rmosoto;xotus 203 Key to theSpeciesoftheGenusF,ormosoto;x:otus 1. Elytradistinctlyconvergent posteriorly f auripifosus - Elytraalmostparallel-sidedorslightlynarrowedposteriad 2 2. Elytra provided with several large deeply impressed punctures, the pubescence around thosepuncturesradiallyarranged F. uenoi - Elytrawithoutlargepunctures;elytra1pubescenceirregularlyarranged. - - - -3. 3. Apicesofeighthtotenthantennalsegmentsangulateexternally. __- -Fnobuoi - Apicesofeighthtotenthantennalsegmentsnotangulate 4 4. Vertexdeeplyandwidelyconcave F malayanus - Vertexshallowlyandnarrowlyconcave 5. Alltibiaestronglycompressedandarcuatelydilatedoutwards(femaleunknown) F takaoi - A11tibiaeslightlycompressedandnotdistinctlydilatedoutwards(femaleunknown) f masata al'sp nov 要 約 大林延夫: Fo,rmosotoxotus属の再検討とシッキム産1新種の記載. - 台湾のキンケカタビロ ハナカミキリを基準種として創設されたFlormosotoxotus属には, これまで6種が知られていた が, このうちマレーシアのキャメロンハイランドから記載されたF fulvopi1osusHAYAsHI, 1979 は, 同じ産地から記載されたFmalayanusHAYAsHI,1977と同種と認め, 前者を後者のシノニム とした.また,パリの自然史博物館に所蔵されていたシッキム産の1新種を,故佐藤正孝博士に 献名してFmasatakaiとして記載した. また, 従来ハナカミキリ亜科に含められていた本属の分 類学的位置について検討し, 後翅翅脈や雄交尾器などの形態がApatophysls属にきわめて近いこ とから, ニセハナカミキリ亜科Apatophyseinaeに含めるべきであるとした. References DANILEvsKY,M.L.,1979. Descriptionofthefemale,pupaandlarvaofApatophySSi paV1oVSkiipLAV・and djscussionofsystematicpositionofthegenusApatophysisCHEvR.(Col.,Cer).Ent.0bOZ「.,58:821-828・ (InRussian) GREsslTT,J.L.,1951. LongicombeetlesofChina. Longicomia,2:667pP.,22PIS・ - &J.A.RoNDoN,1970. CerambycidaeofLaos. Pacif Ins.Mon., (24):1-314. HAYAsHl,M.,1960. ThestudyofLepturinae(Col:Cerambycidae). Nipontus,TakamaStu,1(6):1-26・ - 1977.studyonCerambycidaefromWestMalaysia.Bu11.0sakaJona'tWolmet1s' J「.Colte9e,(12):93- 128. - 1979. studyonCerambycidaefromWestMalaysia(Col).Part11.Ibid., (13):51-87・ Revjsjonof theAsianLepturinae(Coleoptera:Ce「ambyCidae)WithSpecial - & A. VILLIERS, lg85. referencetothetypespecimens'inspection.Part1.Ibid., (19/20):1-75・ JENIs,1.,2001. Long_hornedbeetles,Distenidae,0xypeltidae,Vesperidae,Anop1odermatidae&Ce「ambyCi- dae1. vesperidae&CerambycidaeofEurope,1.333pp.AtelierRegulus,Zlin・ KANO. T、lg33. Newandunrecordedlongicorn-beetlesfromJapananditsadjacentte「「itO「ieS・ KOntyu,