Bull. nat. Hist. Mus. Lond. (Geol.)56(l):21-24 Issued29June 2000 A preliminary of arthropod families list Burmese amber present in the collection at The Natural History Museum, London RASNITSYN A. P. ArthropodLaboratory, PaleontologicalInstitute, RussianAcademy ofSciences, Profsoyuznaya Str. 123, Moscow 117647, Russia ROSS A. J. DepartmentofPalaeontology, TheNaturalHistoryMuseum, CromwellRoad, South Kensington, London SW7 5BD SYNOPSIS. Thepresentaccountisbasedonthe 117piecesofBurmeseamberinthecollectionsofTheNaturalHistoryMuseum, London.Thematerial wasfoundtobeveryrichinfossils yieldingalmost 1200individual arthropodspecimens. INTRODUCTION Insecta OrderArchaeognatha (APR) Below is a list offamilies of arthropods identified in the pieces of Family Machilidae Burmeseamberin the DepartmentofPalaeontology ofThe Natural In.20188 History Museum, London. The collection was initially identified to Order level by AJR, then APR made a more thorough study with OrderZygentoma (APR) 4 additional identifications by scientists at the Paleontological Insti- Family Lepismatidae 3+1? tute, Moscow and other institutions. This list provides, for thNe fHirMst In.19104-6,In.19107- 16,In.19117-22(T),In.20149? time,adetailedaccountoftheBurmeseambercollectionatthe andhopefullywillstimulatefurtherresearchbyspecialistentomolo- OrderEphemeroptera (N. D. Sinitshenkova) gists andpalaeoentomologists. Family Prosopistomatidae The list follows the format ofRoss & Jarzembowski (1993). For In.20173 (T) each order and family, the identifier(s) is given in brackets, along withthetotalnumberofspecimens. Foreachfamily,theregistration OrderBlattodea (APR) 58 numbersofthepiecesofamber(pre-fixedbyIn.)aregiven. Foreach Family Blattellidae (AJR) 3 piece of amber, the number of individual specimens is given in In.19102-3, In.20149, In.20172 bracketsifthereismorethanone(abbreviatedspm), andifthepiece Family Polyphagidae (AJR) 1 contains atypeorfiguredspecimenthis is alsoindicatedinbrackets In.20194 (abbreviatedTorFrespectively).SpecimensIn.19102-23,In.20149- Undetermined (mainly nymphs, isolated legs orbody parts) 54 50andIn.20195allcamefromthesameblockthatwasslicedupinto In.l9102-3(9spm),In.l9104-6(2spm),In.l9107-16 7 slabs (see Cockerell, 1917). The 7 slabs contain a total of 458 (5 spm), In.19117-22, In.19123 (6 spm), In.19128, inclusions, whichmakes the wholeblocktherichestpiece ofamber In.19132 (2spm), In.19136(2spm),In.20149(9spm), ever recorded. For further information on the type and figured In.20150 (7 spm), In.20151, In.20152, In.20164, & specimens in the collection, see the comprehensive list by Ross In.20171 (2 spm), In.20174, In.20175, In.20182, York, this volume. The specimens of copal that Cockerell (1922) In.20192,In.20200 recordedarenotincludedbecausethereis uncertainty asto whether they are from Burma. Order Dermaptera The total numberofspecimens in the list is 1198. Family Labiduridae In.20146(T) Collembola Order Embioptera Order Collembola (APR) 57+1? Family Burmitembiidae Family Sminthuridae 32+3? In.19132 (T) In.19098?, In.19100, In.19123 (4 spm), In.20150? (2 spm), In.20151, In.20186 (10 spm), In.20188 OrderIsoptera (APR) 91? (16 spm) Family Hodotermitidae (K. Krishna) 2 Undetermined 22+1? In.19123, In.20160 In.19100, In.19102-3, In.19123 (2 spm), In.19128, Family Kalotermitidae 89? In.19132, In.20150 (4+1? spm), In.20151, In.20171 In.19096 (2 spm, 1 T), In.19102-3 (11 spm, 1 T), (9 spm), In.20180, In.20183 In.l9104-6(3spm),In.l9107-16(15spm),In.l9117- ©TheNatural HistoryMuseum,2000 3 ) 22 A.P. RASHITSYN AND A.J. ROSS 22, In.19123 (many, 50? spm), In.20149 (5 spm), OrderThysanoptera (APR 14 In.20174,In.20699 Family Lophioneuridae (V. V. Zherikhin) 1 In.20 94 (T) 1 OrderMantodea (P. Vrsansky,APR) 3+1? Undetermined 13 Undetermined 3+1? In.19133^1, In.19135, In.19098, In.20149, In.20168, In.19100-1 (nymph), In.20150, In.20152 (nymph), In.20172 (8 spm) In.20162? (ootheca?) Order Coleoptera (APR,V. G. Gratshev, N. B. Nikitsky, P. OrderOrthoptera (APR) 3+2? Hammond) 195 Superfamily Grylloidea 2+1? FamilyAderidae 1? In.20152? (nymph), In.20171 (2 spm) In.20150? Undetermined 2? FamilyAnthicidae 1 In.19107-16?, In.20161? In.19107-16 (T) Superfamily Cantharoidea 1+1? OrderPhasmatodea (APR) 3? In.19123 (larva), In.20169? Undetermined 3? Family Carabidae 1 In.19117-22? (3 spm, eggs) In.20150 (larva) Family Cerambycidae 1 OrderHemiptera (D. E. Shcherbakov) 82 In.20188 FamilyAchilidae 28 Family Chrysomelidae 1? In.19094 (2 spm), In.19102-3 (8 spm), In.19104-6 (2 In.20150? (larva) spm, 1 T), In.19117-22 (5 spm), In.19123 (2 spm), Family Cisidae 1? In.20149(4spm),In.20150(2spm),In.20168,In.20186, In.19136? In.20191 Superfamily Cleroidea 1? FamilyAleyrodidae 3 In.20169? In.19133-4 (T), In.20193 (T), In.20703 Family Colydiidae 3+3? FamilyAphrophoridae 1? In.19107-16? (T), In.191 17-22?, In.19123 (2 spm), In.19094? In.20149?, In.20174 Family Cixiidae 2 Family Cucujidae 4? In.19094 (2 spm, IT) In.19102-3?, In.19104-6?, In.20150?, In.20174? Superfamily Coccoidea 16+1? Family Dermestidae +1? 1 In.19098, In.19102-3, In.19125?, In.19132, In.20155, In.19107-16 (1+1? spm, IT) In.20160, In.20168 (2 spm), In.20172, In.20174, Family Elateridae 19 In.20175, In.20186, In.20188, In.20191, In.20193 In.19102-3 (8 spm, 1 T), In.19107-16 (5 spm, 1 T), (2 spm), In.20708 In.191 17-22 (2 spm), In.19123 (2 spm), In.20186, Family Coreidae s.l. 1 In.20187 In.20186 Family Eucinetidae 1 Family Enicocephalidae 6 In.20186 In.19095 (T), In.19107-16 (2 spm, 2 T), In.19123 (2 Family Eucnemidae 2 spm, IT), In.19128 In.20168 (2 spm) Superfamily Fulgoroidea undetermined 1 Family Helodidae 1? In.19098 (2 spm), In.19130, In.20160, In.20167 (3 In.20150? spm), In.20171, In.20172, In.20174, In.20175, Family Lathridiidae 1 In.20191, In.20195 In.19136 Family Ochteridae Family Melandryidae 1+1? 1 In.20160 In.19136, In.20706? Undetermined Family Melyridae 1 1 In.19107-16(5spm),In.19117-22,In.19125,In.20152, In.20160 In.20158,In.20185 Family Microsporidae 1 In.19132 OrderPsocoptera 47 Family Mordellidae 1 Family Compsocidae 1 In.20160 In.20193 Family Nitidulidae 1 Family Pachytroctidae 2 In.20178 (larva) In.l9136(2spm, IT) Family Oedemeridae 2? Family Psyllipsocidae 4 In.19123?, In.20190? In.19097 (T), In.20174, In.20193, In.20706 Family Pselaphidae 6 Undetermined 40 In.19104-6 (2 spm), In.19123, In.20150, In.20171, In.19096,In.19098(2spm),In.19102-3,In.19107-16, In.20207 In.19177-22 (2 spm), In.19136 (18 spm), In.20150, Family Ptiliidae 3 In.20151 (2 spm), In.20163, In.20172, In.20173, In.19132, In.19160, In.19107-16 In.20174,In.20175(2spm),In.20178(2spm),In.20186, Family Rhipiphoridae 1 In.20187, In.20199, In.20707 In.19092 (T) 6 , LIST OF ARTHROPOD FAMILIES IN BURMESE AMBER 23 Family Salpingidae 1? In.19097?, In.19098, In.19100-1 (2 spm, 1 T), In.20199? In.19133-4?,In.20147(T),In.20155(2spm),In.20156, Family Scraptiidae 1 In.20164,In.20167(5spm),In.20168(2spm),In.20171 In.19102-3, In.19107-16, In.19117-22, In.19123 (11 In.20172(2spm),In.20187(2spm),In.20189,In.20198, spm), In.l9126,In.20150 In.20707 (4 spm), In.20710 (7 spm) Family Staphylinidae 23 Family Rhagionidae 1+1? In.19107-16, In.19136 (2 spm), In.20150 (13 spm), In.20175, In.20709? In.20152, In.20158, In.20160 (2 spm), In.20169, Family Scatopsidae 1 In.20751,In.20194 In.20150 Superfamily Staphylinoidea undetermined 2 Family Sciaridae 5 In.19123, In.20194 In.19096, In.20151, In.20152, In.20188, In.20199 FamilyThroscidae 1? FamilyTherevidae 1 In.20150? In.20148 (T) Undetermined 89 Superfamily Tipuloidea undetermined 1 In.19092 (2 spm), In.19094 (2 spm), In.19098, In.20171 In.19100-1, In.19102-3 (9 spm), In.19104-6, Undetermined 34 In.l9107-16(12spm),In.l9117-22(12spm),In.19123 In.19097, In.19099, In.19102-3 (2 spm), In.19107- (7 spm), In.19131, In.20149 (10 spm), In.20150 (3 16 (2 spm), In.19117-22 (3 spm), In.19123 (larva), spm),In.20151,In.20152 (5 spm),In.20158,In.20160, In.19136, In.20149 (3 spm), In.20150, In.20168, In.20164, In.20173, In.20176, In.20183 (2 spm), In.20172, In.20173 (2 spm), In.20183, In.20184 (2 In.20188, In.20189 (3 spm), In.20192, In.20193, spm), In.20188, In.20192 (2 spm), In.20194 (2 spm), In.20194 (4 spm), In.20195, In.20196, In.20200, In.20199 (2 spm), In.20208 (3 spm), in.20708 (2 In.20205, In.20708 spm) OrderDiptera (V. A. Blagoderov, M. B. Mostovsky) 203 OrderHymenoptera (APR) 102 FamilyApsilocephalidae Family Bethylidae 31 1 In.20167 (T) In.19107-16 (5 spm, 2 T), In.191 17-22 (3 spm, 1 T), Family Cecidomyiidae 14+1? In.19123 (9 spm), In.19136 (2 spm), In.20149 (7 spm, In.l9096(3spm),In.l9107-16(T),In.l9125,In.20151, IT), In.20150 (5 spm) In.20160, In.20167, In.20168 (2 spm), In.20173, Superfamily Chalcidoidea 3 In.20174, In.20194 (1+1? spm), In.20210 In.20208, In.19133^1, In.20194 Family Ceratopogonidae 12 Family Diapriidae 1? In.19133-4 (T), In.20168 (5 spm), In.20172, In.20173 In.19124? (2 spm), In.20190, In.20208, In.20700 Family Embolemidae 2 Family Chaoboridae (E. D. Lukashevitch) 3 In.20151, In.20194 In.20157 (T), In.20168 (2 spm, 2T) Family Evaniidae 1 Family Chironomidae 25+6? In.20192 (T) In.19098 (2 spm), In.19117-22 (2 spm), In.20148?, Family Formicidae 5 In.20153 (7 spm), In.20168 (2 spm), In.20173?, In.19125 (T), In.19132, In.20182 (3 spm, 1 T, IF) In.20183 (1+2? spm), In.20189, In.20194 (1+1? spm), Family Gasteruptiidae 6 In.20207, In.20704?, In.20705 (2 spm), In.20707 (4 In.19091 (T),In.19098(T),In.19117-22(T),In.19129, spm), In.20708 (2 spm) In.20169, In.20174 Family Diadocidiidae 5 Family Megaspilidae 1 In.20175 (4 spm), In.20188 In.20159 Family Dolichopodidae 1? Family Mymarommatidae 2 In.20197? In.20193, In.20208 Family Empididae 38+4? Family Pelecinidae 1 In.19096(8spm),In.19099(3spm, 1 T),In.19102-3(4 In.20174 spm),In.19104-6(F),In.19107-16(3?spm),In.19117- Family Scelionidae 23 22 (5 spm, 1 T), In.19123 (7 spm), In.19136 (2 spm), In.19092, In.19094, In.19096, In.19102-3 (2 spm), In.20149?, In.20151, In.20173 (2 spm), In.20178, In.19107-16, In.19117-22 (5 spm), In.19136 (3 spm), In.20192 (3 spm), In.20209 In.20158(2spm),In.20167,In.20183(2spm),In.20193, Family Keroplatidae 2 In.20197, In.20706 (2 spm) In.19104-6 (T), In.20179 Family Serphitidae 2 Family Limoniidae 1 In.20151, In.20190 (F) In.20155 Family Sphecidae 17+1? Family Mycetophilidae (P. Chandler) 1 In.19093 (T), In.19123 (2 spm, 2 T), In.19125 (T), In.19100-1 (T) In.19136 (9 spm, 9 T), In.20150 (T), In.20164?, Superfamily Mycetophiloidea undetermined 7 In.20181 (T), In.20197 (T), In.2071 1 (T) In.19136, In.20171 (4 spm), In.20175 (2 spm) Family Tiphiidae 2? Family Phoridae 3 In.20150? (2 spm) In.19102-3, In.19123, In.20193 Undetermined 4 Family Psychodidae 33+2? In.19102-3, In.19104-6, In.20149, In.20208 24 A.P. RASHITSYN AND A.J. ROSS (8 spm), In.20158, In.20171 (3 spm), In.20179, In.20183, In.20186 (3 spm), In.20188, In.20189 (7 spm), In.20699 OrderAraneae (K.Y. Eskov) 36 OrderLepidoptera (APR, G. Robinson) 7 Family Eusparassidae 1 Family Micropterygidae 3+1? In.20197 (juvenile) In.19135 (T), In.20167, In.20168, In.20204. Family Myrmeciidae 3 Undetermined In.20193, In.20197, In.20208 In.l9123,In.20151,In.20172 Family Oonopidae 2 In.20210, In.20168 Order Neuroptera (A. G. Ponomarenko, M. Engel) Family Pisauridae (D. Penney) 1 Family Berothidae In.19132 In.20193,In.20177,In.20197 Family Tetragnathidae 1? Family Osmylidae In.20208? In.20150 (larva) Family Theridiidae 1 Superfamily Osmyloidea undetermined In.20193 In.19107-16? (larva) Family Thomisidae 1 Undetermined In.20174 (juvenile) In.19102-3 Undetermined 26 In.19104-6, In.19117-22 (2 spm), In.19136 (3 spm), Order Raphidioptera (A. G. Ponomarenko) In.20151, In.20152 (4 spm), In.20154, In.20172 (2 Undetermined spm),In.20174,In.20178, In.20183,In.20186(3 spm), In.20150 (larva) In.20189(2spm),In.20191 (2spm),In.20194,In.20700 OrderTrichoptera Order Chelonethi (APR, K.Y. Eskov) 38 Family Hydroptilidae Family Cheiridiidae (M. L. I. Judson) 3 In.20180(T) In.191 17-22 (2 spm, 1 T), In.19123 (F) Family Olpiidae (M. L. I. Judson) 1 OrderUndetermined (APR) 13 In.20149 (T) In.19102-3(3spm),In.19104-6,In.19107-16(2spm), Undetermined 34 In.19117-22, In.20149 (2 spm), In.20150 (2 spm), In.19102-3 (6 spm), In.19107-16 (5 spm), In.19117- In.20167 (2 spm) 22, In.19123 (11 spm), In.20149 (2 spm), In.20150 (6 spm), In.20171, In.20189, In.20195 Arachnida Order Scorpionida (APR) Undetermined OrderAcarina (A. L. Kartsev, O. L. Makarova) 164 FamilyAnystidae In.20174 (F) 1 In.20167 Diplopoda Family Bdellidae In.l9128,In.20193 Superfamily Bdelloidea undetermined 1? Order Polyxenida (APR, S. I. Golovach) 60 In.20203? Family Synxenidae 60 Family Cheyletidae In.19102-3 (4spm),In.19104-6(6spm), In.19117-22 In.19107-16 (T), In.19123 (3 spm), In.20149 (2 spm. 1 T), In.19123 (24 spm), In.20149 (5 spm), Family Erythraeidae 25 In.20150 (18 spm), In.20169 In.19102-3, In.19107-16 (2 spm), In.19123 (3 spm), In.19132, In.20149, In.20150 (11 spm), In.20188, Order Undetermined (S. I. Golovach) 1? In.20193 (5 spm) In.19123? Family Eupodidae 5+2? In.20150?, In.20193 (5+1? spm) Family Ixodidae REFERENCES In.19125 (juvenile) Undetermined 122 In.19095,In.19100-1,In.19102-3(14spm),In.19104- Cockerell,T.D.A. 1917.ArthropodsinBurmeseamber.AmericanJournalofScience. 6 (6 spm), In.19107-116 (25 spm), In.19117-22 (8 Ser.4,44: 360-368. 1922. Fossilsin Burmeseamber.Nature. 109: 713-714. spm), In.19123 (25 spm), In.19128 (2 spm), In.19132, Ross,A.J.&Jarzembowski,E.A. 1993.Arthropoda(Hexapoda:Insecta).In:Benton. In.20149(4spm),In.20150(8spm),In.20151,In.20152 M.J. (editor) Thefossilrecord2: 363^26. Chapman& Hall.London.