A new species of the genus Lucasioides (Crustacea: Isopoda) from Ashiu, Miyama, Nantan-shi, Kyoto, central Japan journal or Bulletin of the Toyama Science Museum publication title number 33 page range 39-45 year 2010-03-15 URL http://repo.tsm.toyama.toyama.jp/?action=repos itory_uri&item_id=921 富山市科学博物館研究報告第33号，pp39-45；（2()10） ANewSpeciesoftheGenusL"c“j"火s（Crustacea：Isopoda）fromAshiu， Miyama，Nantan-shi，Kyoto，CentralJapan＊ NobomNunomura ToyamaScienceMuseum l-8-31Nishinakano-machi，Toyama939-8()84,Japan 京都大学芦生研究林から発見されたハヤシワラジムシの1新種 布村昇 富山市科学博物館 939-8084富山県富山市西中野町1-8-3： 京都府南丹市の京都大学フィールド科学教育研究センター森林ステーション芦生研 究林内から渡辺弘之博士により採集された陸生等脚目の1種を調査したところ、オオ ハヤシワラジムシ属の新種であることが判明し、新種L"casjoj昨sash奴e"sjs(和名： アシュウハヤシワラジムシ：新称)として記載した。本種は宮崎県から知られている ニチナンハヤシワラジムシL師c方加α"e"”Nunomuraと最も類似するが、(1)はっき りした背面の模様があること、(2)第2触角の第2節が第1節より明らかに長いこと、（3） 胸肢に多数の2叉剛毛が見られること、(4)第1触角の剛毛数が多いこと、(5)第1〜4胸 節の剛毛が側縁からより離れた位置にあること、(6)第1および第2胸節の長節や腕節 の剛毛は比較的密生しないことなどで区別される。本種のホロタイプは富山市科学博 物館（TOYACr-l9907）に保管される。 キーワード：等脚類、新種、芦生、分類学、アシュウハヤシワラジムシ Keywords：：Isopod，newspecies，Ashiu，taxonomy，L"c“joj昨s“〃"e"sjs Hitherto，11speciesofthegenusL"c“/or昨sintheworld（Shmalfilss、2003）and7speciesinJapan(Nunomura， 1987,1991,1992,2000,2003,2008）havealreadvbeenknownasvaledRecently,ProfEmeritusDrWatanabehappened tocollectstrangeten･estrialisopodcrustaceansinAshiuForestResearchStation，FieldScienceandResearchCenter， Nantan-shi，KyotoThevweresenttomefbridentiflcation，Atthecloserexaminationsofmine，theyprovedtorepresent tobeasnewspeciesofthegenusZ,"c“joj昨s、 Orderlsopoda FamilyAgnaridae L"c妬加i“s“ん加E"”，n.Sp. （Japanesename:Ashiu-hayashi-warajimushi,new） （Figs，1−3） Mnre"α／exα'"j"“：2早早(l早holotype，70mminbodylengthandl早paratype，5．4mminbodylength)： Nodahatadani，Ashiu，Miyama，Nantan-shi，Kyoto，31July2009,colLHiroyukiWatanabe；1J(allotype，70mmln bodylength)andl早paratype,55mminbodylength),Kamitani,AshiuMiyama,Nantan-shi,Kyoto,May26,2009,colL HiroyukiWatanabeTypeseriesisdepositedasfbllows：holotype（TOYACr-19907)，allotype（TOYACr-19908）and ＊Contributions廿omtheTovamaScienceMuseum・No.38重 ヅ qC J里』 NobomNunomura aparatype（TOYACr-l9909）atToyamaScienceMuseum,paratypesandaparatype（OMNHAr‐7776）atth運 Osaka MuseumofNaturalHistory、 Descr""o〃げんo/o”eた"，α/e･Bodyellipsoid(Fig.1A),2ltimesaslongaswide・Colorblackish,withlongltu- dinaldarkerpattems，symmetrywithrespecttoaline，itlooksasifanlmageofRorschachtest・Medialareao萱 anterlormarginofcephalontrlangular(FiglB).Anterolaterallappetlateralborder､Nodulilateralisonpereonalsomlte皇 l-4（Fig.1R）remote廿omthelateralborder､Posterlormarginofpleotelsontrlangular． 一 一 一 /ぐずで ､#・ルー I・ 一員伽 E ﾙｩｰ ’ 〆一一 ､(cid:631) ー1mm (、一一、 一一一一一一一一一〜 、− {◎ 門一 、 ‐〜＝ごー(cid:631)、 が 工菖 ８ 別 P ０ご ００４０３ざ５４５ 0．営2一5 一・一・・》一 01胃 ’0．05 が R 】 Ⅱ ､ Ⅳ V Ⅵ Ⅶ Fig.1．L"“s"j此s“ん加e"sjsn.§p、 A，DorsalviewラB，AnteriorviewofcephalonラC，Antennule；D､AntennaラE，Rightmandible；F，Le仕mandible； G，Innerlobeofmaxillula；H，OuterlobeofthesameラI，Maxilla；J，Maxilliped；K−N，Pleopodsoffemalel-4＆O Endopodofpleopod5P，Exopodofthesame；Q,Uropod;R,PositionofnodulilateraIis（All,Holotypefemale） 4も AnewL"c“/o/火sfromAshiuJapan Antennule(FiglC）3−segmented；terminalsegmentwith6aesthetascsatthetipAntenna（FiglD)，reachingthe boundaxybetweenpereonitesland2・TenninalHagellarsegments1．6timeslongerthanthebasalone・Rightmandible (FiglE):parsincisiva3-headed；laciniamobilisnotchitinizedweakly3-toothedラ3penicils；processusmolarisrepre‐ sentedbyatufiofsetae・Leftmandible（FiglF):parsincisiva3-headed；laciniamobilis3-toothed；2penicilsぅprocessus molarisrepresentedbyamfiofsetae・Maxillula：innerlobe（FiglG)with2plumosesetaeasmallacuteprQjection outerlobe（FiglH）withlOsimpleteethondistalmarginMaxilla（FiglI）relativelyslenderラapicallybilobed,the mnerwiderandbearingafieldofdenselyspacedsensillaMaxillipe。（FiglJ)：enditewith3spursondistalmargm； palpalsegmentlwithastrongtooth Pereopodl（Fig2A):basis3．6timesaslongaswide,withlO-l2setaeonlnnermarginand5setaeonoutermar‐ gin；ischium45％aslongasbasis，with6setaeonlnnermarginand2setaeonoutermargln；merusalmostaslong asischium，withlOsetaeonlnnermarginandsomeshortsetaeonoutermargln；calpusaslongasmerus，withabout 20setaeincluding6-7bifUrcatedonesonlnnermargln；propodusalittlelongerthancalpus，with81ongsetaeand morethanl8shortsetaeonlnnermargln，andllsetaeonoutermargln、 Pereopod2（Fig2B):basis33timesaslongaswide,with7setaeonlnnermarginand4setaeonoutermargln§ ischiumO､6timesaslongasbasis，withlOsetaeonmnermarginand2−3setaeonouterdistalareaぅmerus0．8times aslongasischium，with8setaeonmnermarginandasetaatouterdistalareaラcarpusl3timeslongerthanmeruso with41ongbifUrcatedsetaeand4-5shortersetaeonlnnermarginll-l2setaeonoutermarglnand4setaeatouter distalangleうpropodusalittlelongerthanca中us，with2bihlrcatedsetaeand3-4setaeonmnermarginand9−10setae onoutermargln・ Pereopod3（Fig2C)：basis2．8timesaslongaswide,withlO-l2setaeonbothmargins；ischium45％aslong asbasis，withlOsetaeonmnermargmandasetaatouterdistalangle；merus0．65timesaslongasischium，with 9-lOsetaeonlnnermarginandasetaatouterdistalangleうcalpusl､stimeslongerthanmerus，withaboutl2setae including3-5bifUrcatedonesonlnnermargin，2relativelylongandsomeshortsetaemcludingbifnrcatedonesondis- taImargln，and8-9setaeonoutermargln；propodusalittlelongerthancaIpus，withlOsetaeonbothmargins・ Pereopod4（Fig2D)：basis27timesaslongaswide,withl3-l4shoItsetaejischiumhalfthelengthofbasiS with5setaeonmnermarginand2−3setaeonoutermargln；merus0．7timesaslongasischium，with7setaeonlnner margln；camus1．3timeslongerthanmerus，withlOsetaeincluding4bifUrcatedonesonlnnermargin，4−5setaeon distalmarginand6-8shortsetaeonmnermargm；propodusalmostaslongascalpus，with5setaeonbothmarglns。 Pereopod5（Fig2E)：basiswide,basis2．9timesslongaswide,with9-lOshortersetaeonmnermarglnanda longersetaatlnnerdistalangle；ischiumO､7timesaslongasbasis，withlO−12setaeonmnermarginand2setaeon outermargln；merus0．4timesaslongasischium，with9−lOsetaeonlnnermarglnand2setaeonouterdistalarea』 carpus1．4timeslongerthanmerus，with7-9setaeonlnnermargin，2−3setaeondistalmarglnandl5-l6setaeon outermargln；propodusalittlelongerthancamus，withlOsetaeonmnermarglnand6setaeonoutermargln･ Pereopod6(Fig2F）：basis27timesaslongaswide,withasetaatmnerdistalangleandlOsetaeonlnnermar‐ gin；ischium65％aslongasbasis，with8setaeonlnnermarginand3setaeonouterdistalarea；merus0．55times aslongasischium，with7setaeonbothmargln；carpus1．7timeslongel･thanmelus，withlOsetaeonlnnermargm， 3setaeondistalmarglnandl3-14setaeonoutermarglnぅpropodusalittlelongerthancarpus，withl31ongerand shortersetaeonlnnermarginll-12setaeonoutermargln Pereopod7（Fig2G):basis3．7timesaslongaswide,with7-8setaeonlnnermarginand3-4setaeonoutermar‐ ginうischiumS5％aslongaswide，with3setaeonoutermargln；merus0．8timesaslongasischium，with5setae onlnnermargin，4setaeonmnerdistalarea，2setaeonouterdistalareaand5−6setaeonoutermargl､；carpusl,Z tlmeslongerthanmerus，withllsetaeonlnnermargin，2setaeondistalmarglnand7-8setaeonoutermargln号 propodusaslongascarpus，with3weakersetaeonbasalareaand3strongersetaeondistalareaofinnermargm， aboutll-l2shortersetaeonoutermargln． 41 NobomNunomura Pleopodl（FiglK)：exopodroundedtriangular,with2setaeonoutermargln Pleopod2（FiglL)：exopodwith3-4setaeonoutermargl、 Pleopod3（FiglM):exopodwith2setaeonoutermargln Pleopod4（Fig｣N)：exopodtriangular,with6setaeonoutermargln Fig.2．〃“αsmj火s“〃"e"姉，、｡sp， A−G；Pereopodsl-7(All：Holotypefemale)： ４ AnewL"c“jo〃たs廿omAshiuJapan PleopodS:endopod（FiglO）rectangularうexopod（FiglP）roundedtriangular,with5setaeonoutermargln Uropod(Fig｣Q）:basistrapezoid；endopodaslongasbasis,withmorethanlOsetaeonbothsidesうexopodl4times longerthanendopod，withmorethanl5setaeonlnnermarginand6−7setaeonoutermarglnandatuftofsetaeon apicalmargln． A姓J/e：Astopereopods，pleopodsandantenna，therearesomedifferenceslnmale・ Antenna（Fig3A）terminalnagellarsegments2ltimeslongerthanthebasalone；mumallengthoftenninalsegment slightlyshorterthanfbmale・ Peropodl（Fig3B)：basis29timesaslongaswide；ischiumhalfthelengthofbasis；merusO8timesaslong aswide，withdenselvsetaeincludingbihlrcatedandtrifUrcatedsetaeonlnnermargln；carpus1．2timeslongerthan merus，withdensesetaeincludingsomebihlrcatedandtrifilrcatedonesonmnermargln；propodusO､8timesaslong ascamus，withaseriesofsetaeonbasalhalfofinnermar2inand2setaeondistalhalfofinnermargln・ Pereopod2(Fig3C):basis33timesaslongaswide;ischiumO,55timesaslongaswide,withl2setaeonmner marginand2setaeonouterdistalarea；merusabouthalfthelengthofischium，withmanvsetaeincludingbihlrcated andtrihlrcatedonesonlnnermargln；camuswithmanysetaeincludingbifilrcatedandtrifUrcatedsetaeonlnnermargln setae；propodusO､8timesaslongascarpus，with9-lOsetaeonlnnermargln・ Peropod3（Fig3D):basis29timesaslongaswide;ischiumQ6timesaslongasbasis,with21ongsetaeondis‐ talareaandabout4setaeonlnnermarglnanddistalangle；merus2/3aslongasischiumwithmorethan20Setae oninnermarginand2setaeonlnnermarglnぅcarpus1．3timeslongerthanmerus，withmorethanl5setaeincluding bifilrcatedonesonlnnermarglnand2-3setaeonlnnermarglnぅpropodus1．2timeslongerthancalpus，with8setae onlnnermargln、 Pereopod4（Fig3E)：basis28timesaslongaswide；ischium45％aslongaswide,with8-10setaeonlnner marginand3-4outerdistalarea；meruswithmorethan20setaeonmnermarginincludingbifUrcatedonesラcamusl,S timeslongerthanmems，withmorethan20setaeincludingbihlrcatedonesonlnnermargm；propodusalittlelonger thancarpus，with6-7bifilrcatedormfilrcatedsetaeonlnnermarginandl7-l9setaeonoutermargln Pereopod5（Fig3F):basis32timesaslongaswide;ischiumO6timesaslongaswide,with8-9setaeonlnner marginand4-5setaeonouterdistalarea；merus0．6timesaslongasischium，withl2−13setaeonlnnermarglnand 3setaeonouterdistalareaラcaIpusl・stimeslongerthanmerus’withl3-14setaeonlnnermargln；propodusalittle longerthanlOsetaeonmnermargmandmorethan20shortsetaeonoutermargin Pereopod6（Fig3G):basis3.ltimesaslongaswide,withl5-16setaeonmnermarginand8-llsetaeonouter margin：ischiumO,6timesaslongasbasis，withll-l2setaeonlnnermarginand21onEbihlrcatedsetaeonouterdis- talareaうmems2/3aslongasischium，withlO-l2setaeincludingbifUrcatedonesonlnnermarglnand2setaeon outerdistalarea；calpus1．2timeslongerthanmerus，withlO-llsetaeincluding21ongbihlrcatedonesonlnnermar- gln；propodusaslongascarpus，withl2−l4setaeonlnnermarginandl3-l4setaeonoutermargln， Pereopod7（Fig.3H)：basis23timeaslongaswide,with2setaeatlnnerdistalangle；ischiumO､8halfthe lengthofbasis，with3setaeonlnnermargln；merus85％aslongasischium；carpusaslongasmerus，posterlorparT protrudedroundly，with8-91ongerand7-8sholtersetaeonlnnermargm，andasetaatouterdistalangle；propodus aslongascalpuswith8setaeonlnnermarglnandl5−l6setaeonoutermargm・ Penes（Fig31）fUsifbnn Pleopodl：endopod（Fig31）straightandapicalpartslightlybentsouterwards,withaseriesofdenticles；exopod (Fig3J）semicircular,innermarginslightlysinuateandoutermarginwith2setae Pleopod2：endopod（Fig3K）longandtapermgtowardthetipandslightlyexceedingbeyondthetipofexopod； exopod(Fig3L）triangular,withroundedtipand5setaeonoutermarginPleopod3（Fig3M):endopodwith3setae onoutermarginPleopod4（Fig3N）exoopodwith3setaeonoutermarginPleopod5（Fig30)：endopodwith3 setaeonoutermargln． 4苫 NobomNunomura Fig.3．L"casmj火s“〃"e"卵sn.sp． A，Flagellumofantennaofmale；B−H，Pereopodsl-7ofmale；I，PenesandPleopodl；J，Exopodofthesame；K， Endopodofmalesecondpleopod;L,Exopodofthesame；M-O，Exopodofmalepleopods3-5（All：AⅡotypemale) E〃"20/ogy:ⅧAshiu"isthenameoftypelocality・ 姓"'”肺：ThepresentnewspeciesismostcloselyalliedtoL〃jc〃加α"e"sjs（Nunomura，2003)，recorded廿om Miyazaki，southemKyushu,Butthefbnnerlsseparated廿omthelatterinhefbllowingfeamres:(1)distinguishedcoler pattem,（2）longerterminalnagellarsegment,（3）presenceofmanybifilrcatedsetaeonpereopods,(4)moreaesthetascs ofantennule,（5）remotepositionofpereonalsomitesl−4and（6）lesssetoseonmerusandcamusofpereopodsl-2 AcknowledgementS lwouldliketoexpressmysmceregratimdetoDr，HiroyukiWatanabefbrhiskindnesslngivingmeachancetoex− ammesuchinterestmgmaterials． 44 AnewL"c“roi‘たsけomAshiuJapan RefErenc“ Nunomura，N､，1987．SmdiesontheterrestrialisopodcrustaceansinJapanlV・TaxonomyofthefamiliesTrachelipidae andPorcellionidae．β"//・Zhya脚α此/、M/s”11：l-76 Nunomura，N､，1991．Smdiesonthete画estrialisopodcrustaceansinJapa、Ⅵ、Furthersupplementstothetaxonomvo B""､7byα'"αSaMt/s・’14：l-26 Nunomura，N,、2000．TelTestrialisopodandamphipodCrustaceansfomthelmperialPalace，Tokyo，Mｾ脚．MJ”Sa M“’7b勺0，5：79-97． Nunomura，N・’2003．TerrestrialisopodcrustaceanshomsouthemKyushu，SouthemJaDan．B""．Zbva耐aLSt?/，〃"s,‐ 26:25-45． Nunomura,N,2008.TwonewspeciesofthegenusL"c“joj“s（Isopod）廿ommoleistunnelsofNanto-shi,Toyama， middleJapanB""“7byα"7aSciM"&’31:3-ll Nunomura，N､，2009．TerrestriallsopodCnlstaceans廿omDaitolslands，SouthemJapan．B""．7bva碗αSα，MiJJ，．32：75 87． Nunomura，N，2010ANewspeciesofthegenusL"c“joj火s（Crustacea：Isopoda）伽mnestmaterialofMりge7α 〃"αjz""'/,Toyamaken,centralJapanB""、7byα"'α此』M“，33（inpress)． SCIⅡnalhlss，H，2003.WorldCatalogofterrestrialisopod（Isopods：Oniscidea）Sr""垣α'7eγβe/".M""rkSerA,654 l-34L 45