PROC. ENTOMOL. SOC. WASH. 93(3), 1991, pp. 709-718 TAXONOMIC REVIEWS OF LIOONCASEY AND LISTEMUSCASEY, WITH DESCRIPTIONS OF TWO NEW SPECIES (COLEOPTERA: BYRRHIDAE) Paul Johnson J. % Mailing address: Department of Entomology, University of Wisconsin, Madison, Wisconsin 53706. Abstract.—ThegeneraLioonCaseyandListemusCaseyare redescribed; Lioon nezperce, new species, and Listemus kootenai, new species, are described. The following new syn- onyms are presented: Lioon simplicipes (Mannerheim) is the senior synonym ofL. specu- lareCasey and L. puncticeps Casey, and Listemus acuminatus (Mannerheim) is the senior synonym ofL. satelles Casey and L.formosus Casey. Lectotypes are designated for Lioon puncticeps Casey, L. speculare Casey, Listemus acuminatus (Mannerheim), and L. for- mosus Casey. Key Words: Coleoptera, Byrrhidae, taxonomy, nearctic Continuing studies by me on the Byrrhi- plicipes, for which the type could not be daehaveledtothediscoveryofundescribed located (H. Silfverberg, in litt.). species o^Lioon Casey and Listemus Casey Casey's (1912) original description of from northern Idaho and northeastern Lioon is detailed, but critical characters for Washington states, U.S.A. Additionally, an adequatecontemporarycomparison with several synonymies are updated and four other genera were not included. Thus, with lectotypes are designated. Thus, this report the discovery ofa new species attributable is intended to briefly review the species of to this genus the following generic rede- Lioon and Listemus, and to describe one scription is presented: new species in each genus. Form small, length 2.5-4.2 mm, ovoid- globose, highly convex dorsally; shallowly Genus Lioon Casey 1912 todeeplydepressedatpronotal-elytraljunc- Type species: Amphicyrta simplicipes ture; integument shining, piceous with Mannerheim 1852: 342, here designated. olivaceous reflections to subviridescent, Lioon was established by Casey (1912) to venter without viridescent reflections; ev- include Amphicyrta simplicipes Manner- anescently microreticulate; shallowly punc- heim (1852), Simplocaria inflata LeConte tate, each puncturebearingasinglepaleseta. (1868), and his own species L. speculare Head not reposable, mouthparts always (Casey 1912)andL.puncticeps(Casey 1912). visible. Eye ovoid, narrowing anteriorly, Curiously, Casey failed to note a type spe- facetsindividuallyconvex. Antenna 1 1-seg- cies for Lioon even though he did so for mented, clavate; segment 1 short, fusiform; other genera newly described in the same segment 2 short, narrowing distally; seg- article. Types or type series have been ex- ment 3 long, slender, cylindrical; segments amined foralldescribedtaxaexceptA. sim- 4-5 short, cylindrical; segments 6-1 1 pro- 710 PROCEEDINGS OFTHE ENTOMOLOGICAL SOCIETY OFWASHINGTON gressively widening and compressed. Fron- tarsus with simple paired claws, each with toclypeal margin carinate; clypeus obsoles- 2 empodial setae. cent, shortest medially, lateral angles Abdomenslightlylongerthanbroad, pro- minutely triangular; anteclypeal region co- portionately shorter and broader in male; riaceous. Labrum quadrate, emarginate an- sutures linear, suture 1 obsolescent, ster- teriorly. Mandibles asymmetrical in molar nites 1 and 2 connate; intercoxal process of region, rightside mandible shallowly exca- stemite 1 broad, width greater than length vated and with a stout premolartooth, left- of a metacoxa; visible stemite 5 broadly sidemandibledeeplyexcavatedbutwithout rounded apically. Aedeagus trilobed, me- premolar tooth; ectal surface evenly round- dianlobewithapexsubangular,laterallobes ed laterally with a broad flange extending elongate, basal piece incompletely sclero- dorsally; dorsum flattened with a short an- tized, dorsal sclerotization reducedwithdisc terior tubercle; carinate on ventral margin. membranous. Female gonocoxite lobate, Maxillary palpus 4-segmented, ultimate compressed, moderately setose; stylus short segmentpyriform,attenuate;laciniaandga- andcylindricaltoconical, with 2 longapical lea lobate, densely setose apically; lacinia setae. reduced,0.5x lengthofgalea. Labiumlight- ly sclerotized, ligula membranous, shallow- lyemarginateand densely setose anteriorly; Lioon simplicipes (Mannerheim) palpus 2-segmented, ultimate segment pyr- Amphicyrta simplicipes Mannerheim 1852: iform. Posterior tentorial pits shallow. 342; LeConte 1854: 1 16; Hamilton 1894: Pronotum trapezoidal, narrowing ante- 27; Dalla Torre 1911: 18. Type: not seen, riorly, strongly convex, anterior angles not apparentlylost(Silfverberg, inlitt.). Orig- visible from dorsal aspect; posteriormargin inal repository: Zoological Museum, Hel- with4-8 peg-likecrenulaesetslightlybelow sinki. Type locality: "insula Sitkha," margin of dorsal surface; hypopleura tri- Alaska. angular, broad and shallowly concave pos- Simplocaria inflata LeConte 1868: 62; syn- teriorly; prostemal sutures deeply sulcate. onymy by Henshaw 1882: 230; Dalla Prostemum with disc T-shaped, transverse Torre 1911: 13; Leng 1920: 193. Holo- armsbroad; median processbroadlyround- type: sex notconfirmed, probably female. edapically. Mesostemumobsoletelaterally, Labelling: "Cala.; Type 8150; S. simpli- deeply excavate medially for reception of cipes (Mann.), inflata LEC." Repository: prostemal process. Scutellum small, trian- Museum ofComparative Zoology, Cam- gular, acuminate posteriorly. Elytra con- bridge. Type locality: California. nate, without ventral preapical flange; epi- Lioon simplicipes (Mannerheim), of Casey pleura broad basally, narrowingapicallyand 1912: 69; Leng 1920: 193; Hatch 1961: terminating shortly before apex. Metano- 301. tum apterous; axillary sclerites obsolete. Lioon injlatum (LeConte), of Casey 1912: Metastemum quadrate, 3.5x wider than 69. long, anterior intercoxal process wide; met- LioonspeculareCasey 1912: 68; Leng 1920: endostemite obsolete; metepistemum and 193; Hatch 1961: 302. New Synonym. metepimeron obsolete. Legs not retractile; Lectotype here designated: sex not con- procoxae and mesocoxae globular; meta- firmed, probably female. Labelling: coxaenarrow, transverse; trochantersshort, "Metlakatla, B. Col., Keen; Casey be- triangular; femora fusiform; tibiae slender, quest 1925; Type USNM 48388; specu- apex briefly excavate to receive tarsomere lare Csy." Respository: U.S. National 1; tarsi 5-segmented, filiform, simple; pre- Museum of Natural History. Type lo- VOLUME 93, NUMBER 3 711 Figs. 1-2. Aedeagi ofLioon spp., dorsal aspect; Fig. 1, L. simplicipes (Mann.), right paramere detached to show variation; Fig. 2, L. nezperce, n. sp. cality: Metlakatla, British Columbia. Mannerheim(1852)evidentlyhadatleast Paralectotype with same data. two specimens before him at the time of LioonpuncticepsCasey 1912:69; Leng 1920: description; the locations or existence of 193; Hatch 1961: 302. New Synonym. these specimens are at present unknown Lectotype here designated: sex not con- (Silfverberg, in litt.). The type locality was firmed, probably female. Labelling: "Cal; originally given as "insula Sitkha." This lo- Caseybequest 1925; TypeUSNM 48389; calityispresumedtobewhatisnowBaranof puncticeps Csy." Repository: U.S. Na- Island in the Alexander Archipelago, on tional Museum ofNatural History. Type which is sited the previous settlement and locality: Areata, Humboldt Co., Califor- now town ofSitka, Alaska. nia. Threeparalectotypeswith samedata. LeConte (1868) stated that Simplocaria 712 PROCEEDINGS OFTHE ENTOMOLOGICAL SOCIETY OF WASHINGTON inflata "... is easily known by its very in- wide, smooth, impunctate, asetose. Elytra flated form, resembling in outline Amphi- with lateral margins carinate, shallowly re- cyrta simplicipes,"' and briefly noted some flexed. characters of punctation and form. Later, Aedeagus (Fig. 2) with median lobe flat- apparently through advice to Henshaw tened, truncate, and laterally expanded at (1882), LeConte provided the synonymy of apex; lateral lobes with apices narrowly 5. inflataandA. simplicipes. AlthoughDalla rounded; basal piece contorted. Female Torre (1911) listed these species separately gonocoxites short, broad, lightly sclero- as S. inflata and A. simplicipes, they were tized; styli short, cone-shaped. reassociated generically by Casey (1912). This species is known only from the state CaseyrecognizedS. inflata, butwithoutdis- ofIdaho, U.S.A., in the counties ofClear- cussion, and included both species in his water, Latah, and Shoshone. new genus Lioon. Thespecificepithetispresentedasanoun Both LeConte and Casey based their rec- in apposition, and is in recognition of the ognition ofthesetaxa ontraitsofcoloration Nez Perce, an aboriginal American tribe in- and punctation which can be observed to habitingtheregioninwhichspecimenswere varyconsiderably within any series ofspec- collected. imens. All specimens in the type series of Holotypedepositedonpermanentloanto L. speculareare teneral; thus the pallid col- the California Academy of Science, San oration that primarily defines this species Francisco, through the University ofIdaho, in Casey's key is not valid for species rec- Moscow, and bearing the labelling: ognition. Genitalic characteristics of spec- ID[AHO], Clearwater Co., Aquarius, 1700 imens fittingthedescriptionsand tentative- ft., T40N R7E s5 NW'A, 5 April 1986; P. ly attributable to any ofthe synonyms are J. Johnson collector; in epixylic mosses; not significantly different between popula- Holotype: Lioon nezperce Johnson 1990. tion samples. Additionally, 4 paratypes bearing the same datawillbedonatedtotheCaliforniaAcad- Lioon nezperce, New Species emyofSciences. Seventeenadditionalpara- Charactersasforthegenus,except: length types bearing the same data will be distrib- 2.5-3.5 mm, width 1.8-2.4 mm, subglo- uted among the W. F. Barr Entomology bose, oval in dorsal aspect, highly convex Museum, University of Idaho; U.S. Na- in lateral aspect, pronotal-elytral juncture tional Museum of Natural History, Wash- shallowly depressed; piceous with oliva- ington, D.C.;TheNatural HistoryMuseum, ceous reflection, shining, without metallic London; and my personal collection. sheen, piceous ventrally, appendages rufo- Additional material examined with the piceous; punctures small, shallow, equidis- following collecting data: IDAHO, Clear- tant, setigerous. Pubescence pallid, short to water Co., Isabella Creek, 31.X.1984 (4), moderately long. 5.V.1985 (26), 14.VI.1985, P. J. Johnson; Head with punctures moderately sparse, BeaverCreek, 23.X.1984, P. J. Johnson (3); separatedbygreaterthanowndiameter; an- IdahoCo., 1 mi. ESlateCreek, 28.1V.1983, teriormarginoffronsnarrowlycarinate; an- F. W. Merickel (2), 10.IX.1984, P. J. John- tenna with segment 3 one-third length of son (8), 1.VII.1984, A. Allen (35); 20 mi. segment 2; labrum transverse, short, shal- ESE Lowell, Meadow Creek, 17.V.1983, F. lowly emarginate. W. Merickel (3), 1.IX.1984, P. J. Johnson Pronotum with lateral margins finelycar- (5); DeVoto Memorial Cedar Grove, inate, shallowly reflexed; posterior margin 1.X.1982, F. W. Merickel (1); Grangeville, NE bearing 4 tuberculate crenulae each side of 27.V.1985, A. Allen (14); 7 mi. Lowefl, median line. Scutellum slightly longer than 27.IV.1985, P. J. Johnson (17); Latah Co., VOLUME NUMBER 93, 3 713 N 7.2 mi. Harvard, 1 1.XI.1984, P. J. John- ture evenly contoured, not depressed. son (1). Specimens from these additional Dorsalintegument piceous, shining, usually locationsarecurrentlylocated incollections with aeneous and viridescent vittate reflec- ofthe W. F. BarrEntomological Collection, tionson elytra. Pubescence moderately long, University ofIdaho; A. Allen, Boise; or my cinereous with rufous to rufotestaceous own. patches. Punctation moderate to coarse, sparse to dense, often with rugose areas on Key to Lioon Species elytraespeciallyinthehumeralregion. Ven- 1. Larger, 3.2^.2mm;aedeagusasin Fig. 1;dis- tral integument rufous to piceous, shining; tribution as in Fig. 5 L. simplicipes coarsely, moderately to densely punctate. - Smaller, 2.5-3.5 mm; aedeagus as in Fig. 2; Head partially retractile, mandibles vis- known only from north-central Idaho ible when head reposed. Eye oval, moder- L. nezperce ately convex, shallowly emarginate dorso- Determination of Lioon species is diffi- anteriorly; facets confluent. Antennae cult without reliance upon aedeagal mor- 1 1-segmented, clavate; segment 1 asym- phology (Figs. 1, 2) and distribution (Fig. metrically swollen; segment 2 transverse, 5). Lioon simplicipes is found only in mesic 0.35 X length ofsegment 1; segment 3 sub- conifer forests west of the Cascade Range cylindrical, slightly expanded distally, 2x in Oregon and Washington, andcoastal for- length ofsegment 2; segments 3 and 4 short, estsoftheCoast Range mountainsin north- cylindrical, progressively widening distally. ern California, British Columbia, and Fronto-clypeal margin rounded; clypeus southeastern Alaska. Contrarily, L. nezperce obsolescent, narrow medially, triangular is only known from mesic conifer forests in laterally; densely, coarsely punctate. La- north-central Idaho. Its distribution east of brum quadrate; densely, coarsely punctate; the Cascade Range is quite disjunct from shallowly emarginate on anterior margin. that of L. simplicipes due to habitat sepa- Mandibles tridentate at apex, asymmetrical ration by the arid intervening regions. proximally; with setose prostheca; rightside mandible with venter sloping to sharp, lin- Genus Listemus Casey 1912 ear incisor blade, premolar tooth small, Typespecies:ListemusformosusCasey 1912: blunt; mola shallowly concave; leftside 6, by original designation. mandible concave ventrally, incisor blade ListemuswasestablishedbyCasey(1912) broadly emarginate; premolar tooth dis- to include Morychus acuminatus Manner- tinct, blunt; mola with oval, shallowly de- heim (1852), and his own new species L. pressed grinding surface. Labium moder- satelles and L. formosus. Type specimens ately to darkly sclerotized; palpus havebeenexamined forboth Caseyspecies, 4-segmented, ultimate segment narrowly butMannerheim'stypecouldnotbelocated pyriform; galea and lacinia broadly lobate, (Silfverberg, in litt.); however, an apparent densely setose apically; submentum rect- cotype in the LeConte collection was ex- angular, 2X widerthan long, coarselypunc- amined. tate, laterally with deep sutural pits; men- Casey's description ofListemus is inad- tum transverse, 3 x wider than long, equate for proper comparison with other moderately punctate; prementum trapezoi- byrrhid genera, thus with the discovery of dal, narrowing distally, sparsely punctate; a new species the following redescription is palpus 3-segmented, ultimate segment presented: broadly pyriform; ligula bilobate, densely Form small, length 3.3^.4 mm, mod- setose distally. erately convex dorsally, ovoid to elongate- Thorax with pronotum transverselytrap- oval in dorsal aspect; pronotal-elytraljunc- ezoidal, moderately convex; hypopleura 714 PROCEEDINGS OFTHE ENTOMOLOGICAL SOCIETY OFWASHINGTON Figs. 3-4. AedeagioiListemusspp., ventralaspect; Fig. 3,L. kootenai. n. sp.; Fig.4,L. acuminatus(Mann.). broad and shallowly concave posteriorly, approximate, broad medially, sinuate at narrowing anteriorly; prosternal sutures midlength, gradually narrowing laterally; closed. Prostemum T-shaped, lateral arms excavate posteriorly for reception of fem- narrow, medial process wide, broadly ora; posterior intercoxal process narrow. rounded apically. Mesostemum obsolete Scutellum small, narrowlytriangular; elytra laterally, deeply excavated medially for re- notconnate; ventral flange subapical. Meta- ception ofprosternal process; coxae round- notum micropterous, wings scale-like, ed. Metastemum transverse, 2.5 x wider <0.5 X length ofmetanotum; axillary scler- than long; anterolaterally excavated as cru- ites obsolescent. Legs retractile into sternal ral depressions for mesolegs; coxae nearly fossae; procoxaglobular; mesocoxaconical. VOLUME 93, NUMBER 3 715 slightly transverse; metacoxa transverse to History. Type locality: Areata, Hum- metepistemum,narrowlaterally;trochanter boldt Co., California. narrow, elongate triangular; femora wide, ListemussatellesCasey 1912: 13; Leng 1920: compressed fusiform, ventrally flattened to 191. New Synonym. receive tibiae; tibiae dorsally expanded, ca- Holotype: sex not confirmed. Labelling: USNM riniform, set with comb-like arrangment of "Cal; Casey bequest 1925; Type shortspines;tarsi 5-segmented, segments 1- 48326; satelles Csy." Repository: U.S. 4 progressively shorter and segment 5 sub- National Museum of Natural History. equal in length to 2-^ together, segments 1 Type locality: Hoopa Valley, Humboldt and 2 densely pubescent ventrally, segment Co., Califomia. 3 with fleshy ventral lobe; pretarsus with a pair ofsimple claws and 2 empodial setae. Mannerheim'stypecannotbelocated,and Abdomen with stemites coarsely, mod- maybelost(H. Silfverberg, in litt.)The spe- erately-densely punctate; stemite 1 broadly cies was evidently described from at least excavated to receive metalegs; stemites 2- two specimenscollectedby Frankenhaeuser 4 subequal, sutures shallowly arcuate; ster- and Pippingskold on "insula Sitkha" (Man- nite 5 broadly rounded to subtruncate api- nerheim 1852). A specimen bearing Man- cally, 2X length of stemite 4, medio-api- nerheim'sstyleoflabelandhandwritinghas callydepressed. Malewithaedeagussimple, been examined from the LeConte collec- trilobed, ostium ventral and at midlength tion, and may be treated as a cotype as it is of median lobe. Female with gonocoxites known that LeConte had received dupli- compressed, broadly arcuate laterally, catesfrom Mannerheim ofNorthAmerican densely setose; styli short, peglike, apically species (Darlington 1961). Based on the in- setose, moderately sclerotized. definite status of Mannerheim's retained specimen(s) and the label notation "!" in- Listemus acuminatus (Mannerheim) dicating that Mannerheim had examined Morychus acuminatus Mannerheim 1852: thisspecimen,theLeContespecimenishere 341. designated the lectotype ofMorychus acu- Lectotype here designated: sex unknown minatusMannerheim andhasbeen soiden- [abdomen missing]. Labelling: "84.; 106; tified with my label. P. acuminatus IMannerh., Sitkha; Type The type specimens ofL. satelles and L. 2294." Repository: Museum ofCompar- formosus are nearly indistinguishable con- ative Zoology, Cambridge. Type locality: specificforms. Casey(1912) separatedthese "insula Sitkha," Alaska. putative species on highly variable sculp- Pedilophorusacuminatus(Mannerheim), of tural, coloration and pubescence character- LeConte 1857: 30 (not 1854: 115); Ham- istics. Variations in these latter two species ilton 1894: 27; Wickham 1903: 181; Dal- and L. acuminatus overlap considerably. la Torre 1911: 16. Specimens attributable to either L. acu- Listemus acuminatus (Mannerheim), of minatus, L. satelles, or L. formosus have Casey 1912: 12; Leng 1920: 191; Hatch been examined from throughout the known 1961: 296. range ofListemus. Specimens expressing a Listemus formosus Casey 1912: 12; Leng pattem of relatively denser and coarser 1920: 191; Hatch 1961: 296. New punctation and pubescence occur at more Synonym. coastal sites in Califomia and Oregon, with Lectotype here designated: male. Label- specimens expressing sparser and finer ling: "Cal; Casey bequest 1925; Type punctation and pubescence occurring more USNM 48325; formosus Csy." Reposi- northward and along the west side of the tory: U.S. National Museum of Natural Cascade Range. In addition, the apices of 716 PROCEEDINGS OFTHE ENTOMOLOGICAL SOCIETY OFWASHINGTON Fig. 5. Generalized distribution ofLioon spp. in Fig. 6. Generalized distribution ofListemus spp. western United States and Canada. Lioon simplicipes in western United States and Canada. Listemus acu- is coastal and open; L. nezperce is inland and cross- minatusiscoastalandopen; L. kootenaiisinlandand hatched. crosshatched. the parameres are slightly more elongate in males from northern British Columbia and Listemus kootenai^ New Species adjacent Alaska. This morphological trend Characters of the genus, except: length appears to coincide with a trend ofcoarser 4.0-4.4 mm, width 2.3-2.5 mm; pronotal rugosity ofhumeral areas and with the re- and head punctures large, coarse; elytral mainder of the elytra being sparsely and punctation coarse, evenly distributed, not finelypunctate. In addition, thelateral mar- condensed to form rugose patches and gins ofthe parameres vary in depth ofmid- smooth,impunctateareas,piceouswitheven length sinuosity, from nearly linear to con- aeneous reflections, without alternating ev- cave, but no geographically related trends anescent rufous and viridescent vittae; pu- are apparent. A coloration trend is also ap- bescence silvery-greythroughout, notform- parent with more northerly and Cascadian ingpatchesorvittae. Aedeaguswith median specimenstendingtobeevenlyaeneousand lobeapexevenlyrounded; laterallobesslen- more coastal specimens often possessing der, apices distinct and attenuate (Fig. 5). distinct viridescent vittae. This species is known onlyfrom northern There are no marked or regular separa- Idaho State, Kootenai and Boundary coun- tions ofany variations towarranttaxonom- ties, and extreme northeastern Washington ic separation, and in fact large population State, Pend Oreille and Spokane counties. samples will frequently contain most pos- The species epithet is a regional place- sible variations. Interestingly, neither the name treated as a noun in apposition, and types nor other specimens in the Casey col- referstotheKootenaiNativeAmericantribe lection show any indication that he was which once inhabited the general region in- aware ofthis degree ofvariation. volving northern Idaho, northeastern VOLUME NUMBER 93, 3 717 Washington, northwestern Montana, and British Columbia, to southeastern Alaska, southeastern British Columbia. eastward only to the western slopes of the Holotypedepositedonpermanentloanto Cascade Range. the California Academy of Sciences, San Francisco, through the University ofIdaho, Acknowledgments Moscow, and with the labelling: IDAHO, My thanks are extended to the following Kootenai County, Farragut State Park, 7 personsforeithercollectingorlendingspec- May 1985, C. L. Campbell; Holotype Lis- imens, or providing field notes: J. B. John- temus kootenai Johnson 990. Three para- 1 son, C. L. Campbell, and F. W. Merickel, types with same labelling as holotype, and University of Idaho; R. E. Nelson, Colby anadditional 7 paratypeswiththe following College; W. L. Turner, Washington State locality labellings: IDAHO, Kootenai County, 4 mi. W Athol, 9.VI.1971, W. F. University;J. D. LattinandG. Parsons, Or- egon State University; D. H. Kavanaugh, NBaonr-dm(a1)n;,Ha7.nVnIa.h198C6e,daPr. HJ.atJ,oh5n.8sonmi.(1)S; CKailnigfsoorlnviear,AUc.Sa.deNamtyionoaflSMcuiesneceusm; oJf.NaMt.- kWmASNHSIuNllGivTaOnNLa,keP,e1n7d.VIOr.e1i9l8le1,CRo.unE.tyN,el-1 uMrualseHuismtoorfy;NaAtlufrraeldHFi.stNoeryw;toann,d JAr..,AFlileenl,d Psaornk(,2);neSaprokBaanledCKonunotby,caMmt.pgSrpoouknadn,eS5t2a0t0e Boise. J. B. Johnson, D. K. Young, and J. M. Kingsolverreadthemanuscriptandpro- ft., 10.VI.1986, W. J. Turner(2), BaldKnob, vided a number ofbeneficial comments. 5500 ft., 3.VI.1986, J. B. Johnson (1). Para- types are located in the collections of the Literature Cited UniversityofIdaho, Washington StateUni- versity, R. E. Nelson, and my personal col- Casey,T.L. 1912. DescriptivecatalogueoftheAmer- ican Byrrhidae. Memoirson theColeoptera 3: 1- lection. 69. Key to Listemus Species Dalla Torre, K. W. von. 1911. Fam. Byrrhidae, pp. 5-36. InW.Junk,ColeopterorumCatalogus,pars 1. Dorsalpunctationfinetomoderate,frequently 33, Berlin. condensedintorugulosepatchesonelytra;col- Darlington, P. J., Jr. 1961. Letter from LeConte to oration bright aeneous, usually with distinct Alexander Agassiz. The Coleopterists Bulletin viridescentvittaeonelytra;aedeagusasinFig. 15(4): 128. 3; distribution as in Fig. 6 L. acuminatus Hamilton, J. 1894. Catalogue ofthe Coleoptera of - Dorsal punctation coarse, evenly distributed; Alaska, with the synonymy and distribution. coloration evanescent aeneous, without viri- Transactions ofthe American Entomological So- descent vittae; aedeagus as in Fig. 4; known ciety 21: 1-38. only from northern IdahoandadjacentWash- Hatch,M. H. 1961. TheBeetlesofthePacificNorth- ington L. kootenai west, Part III: Pselaphidae and Diversicomia I. UniversityofWashington Press, Seattle. Listemus kootenai differs from L. acu- Henshaw,S. 1882. IndextotheColeopteradescribed minatus by the coarser punctation of the byJ. L. LeConte,M.D.TransactionsoftheAmer- pronotum and elytra, the lack of aeneous ican Entomological Society 9: 197-272. andviridescent vittae, and shapeofthe par- LeConte,J. L. 1854. SynopsisoftheByrrhidaeofthe amere apices (Figs. 5, 6). The allopatric dis- AUnciatdeedmyStaotfesN.atPurraolceSecdiienngcsesof7:t1he13-P1hi1l7a.delphia tribution will separate these species as well. . 1857 (1860). Report upon the insects col- Listemus kootenai is known only from me- lected on the survey. In Zoological Report, Ex- sic conifer forests in extreme northeastern plorationsandSurveysforaRailroad Routefrom WashingtonStateandnorthern IdahoState. the Mississippi RivertothePacificOcean. Wash- ington, D.C. Contrarily, Listemus acuminatus is found in mesic conifer forests from northwestern sur.ve1y86t8o.AlCaoslkeao.ptTerraansoafctthieonUsniotfetdheStAatmeesrciocaasnt California, through Oregon, Washington, Entomological Society 2; 59-64. 8 PROCEEDINGSOFTHE ENTOMOLOGICALSOCIETY OFWASHINGTON 71 Leng, C. W. 1920. Catalogue ofthe Coleoptera of Wickham, H. F. 1903. The North American species AmericaNorthofMexico.Mt.Vernon,NewYork. ofPedilophoms.TheCanadianEntomologist35(7): Mannerheim,C.G. 1852. ZweiterNachtragzurKae- 179-182. fer-fauna der Nord-Amerikanischen Laender des RussischenReiches.BulletindelaSocieteImperi- ale Naturalistes Moscou 25(2): 283-387.