THE RAFFLES BULLETIN OF ZOOLOGY 2010 Supplement No. 23: 109–129 Date of Publication: 31 Oct.2010 © National University of Singapore ANNOTATED CHECKLIST OF ANOMURAN DECAPOD CRUSTACEANS OF THE WORLD (EXCLUSIVE OF THE KIWAOIDEA AND FAMILIES CHIROSTYLIDAE AND GALATHEIDAE OF THE GALATHEOIDEA) PART II – PORCELLANIDAE Masayuki Osawa Research Center for Coastal Lagoon and Environments, Shimane University, 1060 Nishikawatsu-cho, Matsue, Shimane 690-8504, Japan Email: [email protected] Patsy A. McLaughlin Shannon Point Marine Center, Western Washington University, 1900 Shannon Point Road, Anacortes, WA 98221-4042, USA Email: hermit@fi dalgo.net INTRODUCTION Porcellanidae Haworth, 1825 = Porcellaniens H. Milne Edwards, 1837 Ng et al. (2008) and McLaughlin et al. (this volume) referred = Porcellaniadae Randall, 1840 to the “windows” to the literature and the “springboards” for = Porcellanodea Henderson 1888 associating species with their scientifi c names that provided = Porcellainea Holmes, 1900 the foundations for subsequent brachyuran, paguroid and lithodoid research. For the porcellanids, one treatise in particular has provided a similar base upon which virtually all DESCRIPTIVE TERMS AND CURRENT STATUS subsequent porcellanid reports have been patterned. Despite its regional focus, Haig’s (1960) monograph of eastern General morphology. – The general body shape is crab- Pacifi c species included 87 of the 225 species estimated to like and the carapace is well calcifi ed. Regions of the dorsal be present worldwide at the time (Chace, 1951). During the integument are not usually well defi ned. The anterior margin last half century the number of genera has increased from of the carapace is produced into a short rostrum or rostral 14 prior to Haig’s (1960) monograph to 30 and the number lobe. The eyes are pigmented; ocular orbits are sometimes of valid species from 225 to 277. well developed and ocular acicles are absent. The antennules have broad basal segments. The antennal peduncles are four-segmented, with the distal three moveable; antennal HISTORY OF CLASSIFICATION acicles are absent. The third maxillipeds each have a broad and fl attened ischium; no crista dentata is developed. Gills A. Milne-Edwards & Bouvier (1894) reviewed the early consist of 14 pairs of biserial phyllobranchiae, paired placements of the family and despite their superficial arthrobranchs are present on each third maxilliped and on resemblance to brachyuran crabs, the classifi cation of the pereopods 1–4; one pleurobranch is developed on body wall Porcellanidae has had a much less convoluted path than seen above each pereopod 2–5. for the lithodids and Lomis. By the time that Boas (1880) completed his studies of the Decapoda, there was general The chelipeds are symmetrical or asymmetrical, usually agreement that the Porcellanidae was correctly assigned to broad and dorsoventrally fl attened, but sometimes infl ated. the Galatheoidea. That relationship has not been challenged, Pereopods 2–4 are developed as ambulatory legs; pereopod 5 even though other changes in the superfamily Galatheoidea is reduced, modifi ed, and frequently carried under carapace. have been proposed (e.g., McLaughlin et al., 2007). The pleon is well developed, symmetrical, carried bent under cephalothorax, and held against thorax. It is composed of six pleomeres and a telson. INFRAORDER ANOMURA MACLEAY, 1838 Adult males usually have well developed, paired pleopods Superfamily Galatheoidea, extant family Porcellanidae on pleomere 2 that are modifi ed as gonopods; occasionally Superfamily Galatheoidea Samouelle 1819 these are reduced, rudimentary or absent. Females usually 109 Osawa & McLaughlin: Checklist of world Porcellanidae have paired pleopods on each pleomere 3–5, but those of Aliaporcellana telestophila (Johnson, 1958) [Polyonyx] pleomere 3 are sometimes reduced or absent. The telson Allopetrolisthes Haig, 1960 and uropods are well developed; the telson is composed of = Allopetrolisthes Haig, 1960 (type species 5 or 7 segments. Porcellana angulosa Guérin, 1835, by original designation; gender masculine) Development. – Although early carcinologists considered Allopetrolisthes angulosus (Guérin, 1835) [Porcellana] porcellanid larvae to be quite similar to brachyuran zoeae, = Porcellana carinata Kinahan, 1857 it would appear that the only characters shared by the two = Petrolisthes reissi Ortmann, 1892 groups are the reduced and somewhat rod-shaped antennal Allopetrolisthes punctatus (Guérin, 1835) [Porcellana] endopod and the absence of a functional exopod on the = Porcellana cristata H. Milne Edwards, 1837 third maxilliped. Porcellanid zoeae are very distinctive and Allopetrolisthes spinifrons (H. Milne Edwards, 1837) immediately recognized by their very elongate rostral and [Porcellana] posterolateral carapace spines. Only two zoeal stages are Ancylocheles Haig, 1978 passed through prior to the molt to megalopa, except in = Ancylocheles Haig, 1978 (type species Porcellana Petrocheles Miers, 1876 where fi ve zoeal stages have been gravelei Sankolli, 1963, by original designation; documented. It is also only in this genus that uropods develop gender masculine) in a zoeal stage. In all other porcellanids studied to date, Ancylocheles gravelei (Sankolli, 1963) [Porcellana] uropods develop in the megalopal stage. Development of Capilliporcellana Haig, 1978 larval pleopods occurs in the second zoeal stage and these = Capilliporcellana Haig, 1978 (type species pleopods become function in the megalopal stage, except Porcellana murakamii Miyake, 1942, by original in Petrocheles, where pleopods never develop. However, designation; gender feminine) pleopods are reduced or lost with the molts to early crab Capilliporcellana murakamii (Miyake, 1942) stages. And in the cases of males, pleopods do not reoccur [Porcellana] unless the second pair develops as gonopods, except in Capilliporcellana wolffi Haig, 1981 Novorostrum Osawa, 1998, and some species of Petrolisthes Clastotoechus Haig, 1960 Stimpson, 1858, where reduced or moderately developed = Clastotoechus Haig, 1960 (type species Petrolisthes pleopods are seen on pleomeres 3–5. Egg-bearing pleopods do diffractus Haig, 1957, by original designation; redevelop on pleomeres 3–5 or 4 and 5 in adult females. gender masculine) Clastotoechus diffractus (Haig, 1957) [Petrolisthes] Current status. – Among the phylogenetic analyses that have Clastotoechus gorgonensis Werding & Haig, 1983 been reported in the past quarter century, the Porcellanidae Clastotoechus hickmani Harvey, 1999 have remained aligned with the Galatheoidea; however, Clastotoechus lasios Harvey, 1999 the sister taxon varied between the Galatheidae (Martin & Clastotoechus nodosus (Streets, 1872) [Petrolisthes] Abele, 1986; McLaughlin et al., 2007; Ahyong et al., 2009) Enosteoides Johnson, 1970 or the Chirostylidae (Pérez-Losada et al., 2002; Ahyong & = Enosteoides Johnson, 1970 (type species O’Meally, 2004; Macpherson et al., 2005), with Morrison et al. Porcellana corallicola Haswell, 1882, by original (2002) placing the Porcellanidae as sister to the Galatheidae- designation; gender masculine) Chirostylidae pair. Although previous data indicated that Enosteoides lobatus Osawa, 2009 the Porcellanidae was correctly placed in the Galatheoidea, Enosteoides melissa (Miyake, 1942) [Porcellana] Ahyong et al. (2009) found that the Porcellanidae nested Enosteoides ornatus (Stimpson, 1858) [Porcellana] in a paraphyletic Galatheidae. Thus the true phylogenetic = Porcellana corallicola Haswell, 1882 relationship of this family to other galatheids is not as well Enosteoides palauensis (Nakasone & Miyake, 1968) understood as heretofore believed. [Porcellana] = Enosteoides hainanensis Yang & Sun, 2005 Euceramus Stimpson, 1860 CHECKLIST = Euceramus Stimpson (type species Euceramus praelongus Stimpson, 1860, by monotypy; gender Family Porcellanidae Haworth, 1825 masculine) Euceramus panatelus Glassell, 1938 Aliaporcellana Nakasone & Miyake, 1969 Euceramus praelongus Stimpson, 1860 = Aliaporcellana Nakasone & Miyake 1969 (type Euceramus transversilineatus (Lockington, 1878) species Porcellana suluensis Dana, 1852, by [Porcellana] original designation; gender feminine) Eulenaios Ng & Nakasone, 1993 {2} Aliaporcellana kikuchii Nakasone & Miyake, 1969 = Eulenaios Ng & Nakasone, 1993 (type species Aliaporcellana pygmaea (De Man, 1902) [Porcellana] Polyonyx cometes Walker, 1887, by original = Polyonyx pugilator Nobili, 1905 designation; gender masculine) Aliaporcellana suluensis (Dana, 1852) [Porcellana] Eulenaios cometes (Walker, 1887) [Polyonyx] {1} = Polyonyx denticulatus Paul’son, 1875 = Polyonyx hexagonalis Zehntner, 1894 110 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Heteropolyonyx Osawa, 2001 = Porcellanides Nobili, 1901 (type species = Heteropolyonyx Osawa, 2001 (type species Porcellanides festae Nobili, 1901, by monotypy; Heteropolyonyx biforma Osawa, 2001, by original gender feminine) (preoccupied name) designation; gender masculine) = Porcellanopsis Rathbun 1910 (type species Heteropolyonyx biforma Osawa, 2001 Porcellana festae Nobili, 1901, by monotypy; Heteroporcellana Haig, 1978 gender feminine) (replacement name) {5} = Heteroporcellana Haig, 1978 (type species Megalobrachium erosum (Glassell, 1936) [Pisosoma] Porcellana corbicola Haig, 1960, by original Megalobrachium festae (Nobili, 1901) [Porcellanides] designation; gender feminine) {6} Heteroporcellana corbicola (Haig, 1960) [Porcellana] = Porcellanides festae Nobili, 1901 Liopetrolisthes Haig, 1960 = Porcellanopsis festai (Nobili, 1901) = Liopetrolisthes Haig, 1960 (type species Porcellana Megalobrachium garthi Haig, 1957 mitra Dana, 1852, by original designation; gender = Pachycheles rotundus A. Milne-Edwards & masculine) Bouvier 1894 (nomen nudum) Liopetrolisthes mitra (Dana, 1852) [Porcellana] Megalobrachium mortenseni Haig, 1962 = Porcellana spinosa Philippi 1860 Megalobrachium pacifi cum Gore & Abele, 1974 {7} = Porcellana patagonica Cunningham, 1871 Megalobrachium peruvianum Haig, 1960 = Porcellana pulchellula Cano, 1889 Megalobrachium poeyi (Guérin-Méneville, 1855) Lissoporcellana Haig, 1978 [Porcellana] {8} = Lissoporcellana Haig, 1978 (type species = Megalobrachium granuliferum Stimpson, 1858 Porcellana quadrilobata Miers, 1884, by original = Porcellana robusta Torralbas, 1917 designation; gender feminine) Megalobrachium roseum (Rathbun, 1900) [Porcellana] = ? Porcellanides Czerniavsky 1884 (type species Megalobrachium sinuimanus (Lockington, 1878) Porcellana kriczagini Czerniavsky, 1884, by [Petrolisthes (Pisosoma)] original designation; gender masculine) {3} Megalobrachium smithi (Glassell, 1936) [Pisosoma] Lissoporcellana fl agellicola Osawa & Fujita, 2005 Megalobrachium soriatum (Say, 1818) (Porcellana] Lissoporcellana furcillata (Haig, 1965) [Porcellana] {9} Lissoporcellana miyakei Haig, 1981 = Megalobrachium walteri Rodrigues da Costa, Lissoporcellana monodi Osawa, 2007 1968 Lissoporcellana nakasonei (Miyake, 1978) [Porcellana] Megalobrachium tuberculipes (Lockington, 1878) (replacement name) [Pachycheles] = Porcellana maculata Miyake, 1957 (preoccupied Minyocerus Stimpson, 1858 name) = Minyocerus Stimpson, 1858 (type species Lissoporcellana nitida (Haswell, 1882) [Porcellana] Porcellana angusta Dana, 1852, by original Lissoporcellana pectinata Haig, 1981 designation; gender masculine) Lissoporcellana quadrilobata (Miers, 1884) = Porcellina Müller, 1862, (type species Porcellina [Porcellana] stellicola by monotypy; gender feminine) = Porcellana streptochirus White, 1847 (nomen Minyocerus angustus (Dana, 1852) [Porcellana] nudum) = Porcellina stellicola Müller, 1862 = Porcellana streptochira Miers, 1884 = Porcellana rosamondae Boone, 1930 = Porcellana gaekwari Southwell, 1909 Minyocerus kirki Glassell, 1938 = Lissoporcellana paraquadrilobata Yang & Sun, Neopetrolisthes Miyake, 1937 1992 = Neopetrolisthes Miyake, 1937 (type species Lissoporcellana spinuligera (Dana, 1853) [Porcellana] Neopetrolisthes ohshimai Miyake, 1937, by (replacement name) original designation; gender masculine) = Porcellana armata Dana, 1852 (preoccupied Neopetrolisthes alobatus (Laurie, 1926) [Petrolisthes] name) Neopetrolisthes maculatus (H. Milne Edwards, 1837) = Porcellana latifons Stimpson, 1858 [Porcellana] {10} Lissoporcellana streptochiroides (Johnson, 1970) = Neopetrolisthes ohshimai Miyake, 1937 {10} [Porcellana (Pisidia)] Neopetrolisthes spinatus Osawa & Fujita, 2001 Madarateuchus Harvey, 1999 {4} Neopisosoma Haig, 1960 {11} = Madarateuchus Harvey, 1999 (type species = Neopisosoma Haig, 1960 (type species Neopisosoma Petrolisthes vanderhorsti Schmitt, 1924, by bicapillatum Haig, 1960, by original designation; monotypy; gender masculine) gender neuter) Madarateuchus vanderhorsti (Schmitt, 1924) Neopisosoma angustifrons (Benedict, 1901) [Petrolisthes] [Pisosoma] Megalobrachium Stimpson, 1858 Neopisosoma bicapillatum Haig, 1960 = Megalobrachium Stimpson, 1858 (type species Neopisosoma curacaoense (Schmitt, 1924) [Pisosoma] Megalobrachium granuliferum Stimpson, 1858, Neopisosoma dohenyi Haig, 1960 by original designation; gender neuter) Neopisosoma mexicanum (Streets, 1871) [Pachycheles] 111 Osawa & McLaughlin: Checklist of world Porcellanidae Neopisosoma neglectum Werding, 1986 Pachycheles panamensis Faxon, 1893 Neopisosoma orientale Werding, 1986 = Pachycheles sonorensis Glassell, 1936 Novorostrum Osawa, 1998 Pachycheles pectinicarpus Stimpson, 1858 = Novorostrum Osawa, 1998 (type species Pachycheles pilosus (H. Milne Edwards, 1837) Petrolisthes indicus De Man, 1893, by original [Porcellana] designation; gender neuter) Pachycheles pisoides (Heller, 1865) [Porcellana] Novorostrum decorocrus Osawa, 1998 = Pachycheles lifuensis Borradaile, 1900 Novorostrum indicum (De Man, 1893) [Petrolisthes] = Pachycheles fronto Melin, 1939 Novorostrum phuketense Osawa, 1998 {12} Pachycheles pubescens Holmes, 1900 Novorostrum securiger (Melin, 1939) [Petrolisthes] Pachycheles riisei (Stimpson, 1859) [Pisosoma] {13} = Pisosoma riisei Stimpson, 1858 (nomen nudum) Orthochela Glassell, 1936 = Pisosoma glabra Kingsley, 1880 = Orthochela Glassell, 1936 (type species Orthochela = Pisosoma reisei Young, 1900 (misspelling of pumila Glassell, 1936, by original designation; riisei) gender feminine) = Pisosoma risei Moreira, 1920 (misspelling of Orthochela pumila Glassell, 1936 riisei) Pachycheles Stimpson, 1858 Pachycheles rudis Stimpson 1859 = Pachycheles Stimpson, 1858 (type species = Pachycheles rudis Stimpson, 1858 (nomen Porcellana grossimana Guérin, 1835, by original nudum) designation; gender masculine) Pachycheles rugimanus A. Milne-Edwards, 1880 = Pisosoma Stimpson, 1858 (type species Porcellana Pachycheles sahariensis Monod, 1933 pisum H. Milne Edwards, 1837, by original Pachycheles sculptus (H. Milne Edwards, 1837) designation; gender neuter) [Porcellana] Pachycheles ackleianus A. Milne-Edwards, 1880 = Porcellana pisum H. Milne Edwards, 1837 = ? Porcellana parrai Guérin-Méneville, 1855 = Porcellana pulchella Haswell, 1882 Pachycheles attaragos Harvey & de Santo, 1997 = Pachycheles sculptus var. tuberculatus Borradaile, Pachycheles barbatus A. Milne-Edwards, 1878 1900 = Pachycheles ornatus Bouvier, 1906 Pachycheles serratus (Benedict, 1901) [Pisosoma] Pachycheles bellus (Osorio, 1887) [Porcellana] Pachycheles setiferous Yang, 1996 Pachycheles biocellatus (Lockington, 1878) [Petrolisthes Pachycheles setimanus (Lockington, 1878) [Petrolisthes (Pisosoma)] (Pisosoma)] = Petrolisthes (Pisosoma) gibbosicarpus Lockington, Pachycheles spinidactylus Haig, 1957 1878 Pachycheles spinipes (A. Milne-Edwards, 1873) = Pisosoma aphrodita Boone, 1932 [Porcellana] Pachycheles calculosus Haig, 1960 = Porcellana sollasi Whitelegge, 1897 Pachycheles chacei Haig, 1956 Pachycheles stevensii Stimpson, 1858 Pachycheles chubutensis Boschi, 1963 Pachycheles subsetosus Haig, 1960 Pachycheles crassus (A. Milne-Edwards, 1869) Pachycheles susanae Gore & Abele, 1974 [Porcellana (Pachycheles)] Pachycheles tomentosus Hendersson, 1893 Pachycheles crinimanus Haig, 1960 Pachycheles trichotus Haig, 1960 Pachycheles cristobalensis Gore, 1970 Pachycheles velerae Haig, 1960 Pachycheles garciaensis (Ward, 1942) [Pisosoma] Pachycheles vicarius Nobili, 1901 Pachycheles granti Haig, 1965 Parapetrolisthes Haig, 1962 {16} Pachycheles greeleyi (Rathbun, 1900) [Pisosoma] = Parapetrolisthes Haig, 1962 (type species Pachycheles grossimanus (Guérin, 1835) [Porcellana] Petrolisthes tortugensis Glassell, 1945, by original Pachycheles hertwigi Balss, 1913 designation; gender masculine) = Pachycheles balssi Miyake, 1943 Parapetrolisthes tortugensis (Glassell, 1945) Pachycheles holosericus Schmitt, 1921 [Petrolisthes] = Pachycheles holosericus Schmitt, 1918 (in Petrocheles Miers 1876 {17} Nininger, 1918) (nomen nudum) {14} = Petrocheles Miers 1876 (type species Petrolisthes Pachycheles johnsoni Haig, 1965 (Petrocheles) spinosus Miers, 1876, by subsequent Pachycheles laevidactylus Ortmann, 1892 designation by Haig, 1964; gender masculine) = Pachycheles haigae Rodrigues da Costa, 1960 Petrocheles australiensis (Miers, 1876) [Petrolisthes {15} (Petrocheles)] Pachycheles marcortezensis Glassell, 1936 Petrocheles spinosus (Miers, 1876) [Petrolisthes Pachycheles monilifer (Dana, 1852) [Porcellana] (Petrocheles)] = Porcellana rugosimanus White 1847 (nomen Petrolisthes Stimpson, 1858 {18} nudum) = Petrolisthes Stimpson, 1858 (type species Pachycheles natalensis (Krauss, 1843) [Porcellana] Porcellana violacea Guérin, 1831, by original designation; gender masculine) 112 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Petrolisthes aegyptiacus Werding & Hiller, 2007 Petrolisthes galathinus (Bosc, 1802) [Porcellana] Petrolisthes agassizii Faxon, 1893 = Porcellana sexispinosa Gibbes, 1850 Petrolisthes amoenus (Guérin Méneville, 1855) = Porcellana danae Gibbes 1854 [Porcellana] = Porcellana egregia Guérin-Méneville, 1855 = Perolisthes serratus Henderson 1888 = Petrolisthes occidentalis Stimpson 1858 (nomen Petrolisthes armatus (Gibbes, 1850) [Porcellana] nudum) = Porcellana gundlachii Guérin-Méneville, 1855 = Petrolisthes occidentalis Stimpson, 1859 = Porcellana leporina Heller, 1862 = Petrolisthes brasiliensis S. I. Smith, 1869 = Petrolisthes similis Henderson, 1888 (nomen Petrolisthes gertrudae Werding, 1996 nudum) Petrolisthes glasselli Haig, 1957 = Petrolisthes iheringi Ortmann, 1897 Petrolisthes gracilis Stimpson, 1859 = Petrolisthes armatus var. pallidus Verrill, 1908 = Petrolisthes gracilis Stimpson, 1858 (nomen Petrolisthes artifrons Haig, 1960 nudum) Petrolisthes asiaticus (Leach, 1820) [Pisidia] Petrolisthes granulosus (Guérin, 1835) [Porcellana] = Petrolisthes leporinoides Ortmann, 1892 = Porcellana striata H. Milne Edwards, 1837 = Petrolisthes yaeyamensis Miyake, 1937 Petrolisthes haigae Chace, 1962 Petrolisthes bifi dus Werding & Hiller, 2004 Petrolisthes haplodactylus Haig, 1988 Petrolisthes bispinosus Borradaile, 1900 Petrolisthes hastatus Stimpson, 1858 Petrolisthes bolivarensis Werding & Kraus, 2003 = Porcellana inermis Heller, 1862 Petrolisthes borradailei Kropp, 1984 = Petrolisthes tenkatei De Man, 1893 Petrolisthes boscii (Audouin, 1826) [Porcellana] Petrolisthes haswelli Miers, 1884 = ? Porcellana rugosa? White, 1847 Petrolisthes heterochrous Kropp, 1986 = Petrolisthes rugosus Miers, 1884 Petrolisthes hians Nobili, 1901 = Petrolisthes amakusensis Miyake & Nakasone, = Pisosoma fl agraciliata Glassell, 1937 1966 Petrolisthes hirtipes Lockington, 1878 Petrolisthes brachycarpus Sivertsen, 1933 Petrolisthes hirtispinosus Lockington, 1878 = Petrolisthes gracilis var. brachycarpus Siversten, Petrolisthes hispaniolensis Werding & Hiller, 2005 1933 Petrolisthes holotrichus Nobili, 1901 Petrolisthes cabrilloi Glassell, 1945 Petrolisthes japonicus (De Haan, 1849 [Porcellana] = Petrolisthes cabrilloa Glassell, 1945 {19} Petrolisthes jugosus Streets, 1872 Petrolisthes caribensis Werding, 1983 Petrolisthes kranjiensis Johnson, 1970 Petrolisthes carinipes (Heller, 1861) [Porcellana] {20} Petrolisthes laevigatus (Guérin, 1835) [Porcellana] Petrolisthes celebesensis Haig, 1981 = Porcellana valida Dana, 1852 Petrolisthes cinctipes (Randall, 1840) [Porcellana] Petrolisthes lamarckii (Leach, 1820) [Pisidia] {23} {21} = Porcellana dentata H. Milne Edwards, 1837 = Porcellana rupicola Stimpson, 1857 = Porcellana pulchripes White, 1847 (nomen Petrolisthes coccineus (Owen, 1839) [Porcellana] nudum) = Porcellana barbata Heller, 1862 = Porcellana speciosa Dana, 1852 = Petrolisthes nipponensis Miyake, 1937 = Porcellana bellis Heller, 1865 Petrolisthes cocoensis Haig, 1960 Petrolisthes leptocheles (Heller, 1861) [Porcellana] Petrolisthes columbiensis Werding, 1983 Petrolisthes lewisi (Glassell 1936) [Pisosoma] Petrolisthes crenulatus Lockington, 1878 = Petrolisthes lewisi austrinus Haig, 1960 {24} Petrolisthes decacanthus Ortmann, 1897 Petrolisthes limicola Haig, 1988 Petrolisthes desmarestii (Guérin, 1835) [Porcellana] Petrolisthes lindae Gore & Abele, 1974 = Porcellana acanthophora H. Milne Edwards & Petrolisthes magdalenensis Werding, 1978 Lucas, 1844 Petrolisthes manimaculis Glassell, 1945 = Porcellana dubia Kinahan, 1857 Petrolisthes marginatus Stimpson, 1859 Petrolisthes dissimulatus Gore, 1983 = Petrolisthes marginatus Stimpson, 1858 (nomen Petrolisthes donadio Hiller & Werding, 2007 nudum) Petrolisthes donanensis Osawa, 1997 ? = Porcellana Cessacii A. Milne-Edwards, 1878 Petrolisthes edwardsii (de Saussure, 1853) {25} [Porcellana] {22} ? = Petrolisthes Cecoci Balss, 1914 (misspelling of Petrolisthes eldredgei Haig & Kropp 1987 cessaci in fi gure legend) Petrolisthes elegans Haig, 1981 Petrolisthes masakii Miyake, 1943 Petrolisthes elongatus (H. Milne Edwards, 1837) Petrolisthes melini Miyake & Nakasone, 1966 {20} [Porcellana] Petrolisthes mesodactylon Kropp, 1984 {26} Petrolisthes eriomerus Stimpson, 1871 Petrolisthes militaris (Heller, 1862) [Porcellana] {27} Petrolisthes extremus Kropp & Haig, 1994 {20} = Porcellana annulipes White, 1847 (nomen Petrolisthes fi mbriatus Borradaile, 1898 nudum) Petrolisthes galapagensis Haig, 1960 = Petrolisthes annulipes Miers, 1884 113 Osawa & McLaughlin: Checklist of world Porcellanidae Petrolisthes miyakei Kropp, 1984 Petrolisthes violaceus (Guérin, 1831) [Porcellana] Petrolisthes moluccensis (De Man, 1888) [Porcellana = Porcellana macrocheles Poeppig, 1836 (Petrolisthes)] Petrolisthes virgatus Paul’son, 1875 Petrolisthes monodi Chace, 1956 = Petrolisthes trivirgatus Ortmann, 1894 = ? Porcellana speciosa Osorio, 1889 Petrolisthes zacae Haig, 1968 Petrolisthes nanshensis Yang, 1996 Pisidia Leach, 1820 Petrolisthes nigrunguiculatus Glassell, 1936 = Pisidia Leach, 1820 (type species Cancer Petrolisthes nobilii Haig, 1960 longicornis Linnaeus, 1767, by subsequent Petrolisthes novaezelandiae Filhol, 1885 designation by ICZN Opinion 701, 1964; gender = Petrolisthes stewarti Filhol, 1885 feminine) Petrolisthes obtusifrons Miyake, 1937 = Streptochirus Stimpson, 1907 (type species = Petrolisthes varicolor Osawa 1998 Porcellana serratifrons Stimpson, 1858, by Petrolisthes ornatus Paul’son, 1875 subsequent designation by Haig, 1960; gender = Porcellana (Petrolisthes) mossambica Hilgendorf, masculine) 1879 Pisidia bluteli (Risso, 1816) [Porcellana] {31} Petrolisthes ortmanni Nobili, 1901 Pisidia brasiliensis Haig, in Rodrigues da Costa, 1968 Petrolisthes perdecorus Haig, 1981 Pisidia dehaanii (Krauss, 1843) [Porcellana] Petrolisthes platymerus Haig, 1960 Pisidia delagoae (Barnard, 1955) [Porcellana] Petrolisthes politus (Gray, 1831) [Porcellana] Pisidia dispar (Stimpson, 1858 [Porcellana] {32} = Porcellana magnifi ca Gibbes, 1850 = Polyonyx carinatus Ortmann, 1892 Petrolisthes polymitus Glassell, 1937 = Porcellana rostrata Baker, 1905 Petrolisthes pubescens Stimpson, 1858 Pisidia gordoni (Johnson, 1970) [Porcellana (Pisidia)] Petrolisthes quadratus Benedict, 1901 Pisidia inaequalis (Heller, 1861 [Porcellana] Petrolisthes rathbunae Schmitt, 1921 {28} Pisidia longicornis (Linnaeus, 1767) [Cancer] {31} = Petrolisthes rathbunae Schmitt, in Hilton, 1916 = Cancer hexapus Linnaeus, 1767 (nomen nudum) = Pisidia linnaeana Leach, 1820 Petrolisthes robsonae Glassell, 1945 Pisidia longimana (Risso, 1816) [Porcellana] {31} Petrolisthes rosariensis Werding, 1982 Pisidia magdalenensis (Glassell, 1936) [Porcellana] Petrolisthes rufescens (Heller, 1861) [Porcellana] Pisidia serratifrons (Stimpson, 1858) [Porcellana] Petrolisthes sanfelipensis Glassell, 1936 = Porcellana spinulifrons Miers, 1879 = Petrolisthes felipensis Glassell, 1937 Pisidia streptocheles (Stimpson, 1858) [Porcellana] Petrolisthes sanmartini Werding & Hiller, 2002 Pisidia striata Yang & Sun, 1990 Petrolisthes scabriculus (Dana, 1852) [Porcellana] Pisidia variabilis (Yang & Sun, 1985) [Enosteoides] {27} {32} Petrolisthes schmitti Glassell, 1936 Polyonyx Stimpson, 1858 Petrolisthes squamanus Osawa, 1996 = Polyonyx Stimpson, 1858 (type species Porcellana Petrolisthes teres Melin, 1939 {29} macrocheles Gibbes, 1850, by original designation; = Petrolisthes inermis Haswell, 1882 (preoccupied gender masculine) name) Polyonyx biunguiculatus (Dana, 1852) [Porcellana] Petrolisthes tiburonensis Glassell, 1936 {33} Petrolisthes tomentosus (Dana, 1852) [Porcellana] = Polyonyx parabiunguiculatus Yang, 1996 = Porcellana penicillata Heller, 1862 Polyonyx boucheti Osawa, 2007 = Porcellana villosa Richters, 1880 Polyonyx bouvieri Saint Joseph, 1900 Petrolisthes tonsorius Haig, 1960 Polyonyx confi nis Haig, 1960 Petrolisthes tridentatus Stimpson, 1859 Polyonyx gibbesi Haig, 1956 (replacement name) = ? Porcellana affi nis Gray, 1831 = Porcellana macrocheles Gibbes, 1850 (primary = Petrolisthes tridentatus Stimpson, 1858 (nomen homonym) nudum) = ? Porcellana crepinii Müller, 1862 (nomen Petrolisthes trilobatus Osawa, 1996 nudum) Petrolisthes tuberculatus (Guérin, 1835) [Porcellana] = ? Porcellana (Polyonyx) creplinii Müller, 1870 = Porcellana lobifrons H. Milne Edwards, 1837 = ? Polyonix creplinii Ejchel, 1965 (misspelling of = Porcellana tuberculifrons Nicolet, 1849 Polyonyx) (misspelling of tubertculatus) = ? Polyonix creplini Ejchel, 1965 (misspelling of Petrolisthes tuberculosus (H. Milne Edwards, 1837) Polyonyx) [Porcellana] {30} Polyonyx haigae McNeil, 1968 = Porcellana affi nis Guérin, 1835 (preocccupied Polyonyx hendersoni Southwell, 1909 {34} name) Polyonyx loimicola Sankolli, 1965 = Porcellana tuberculifrons H. Milne Edwards & Polyonyx maccullochi Haig, 1965 Lucas, 1843 (unnecessary replacement name) Polyonyx nitidus Lockington, 1878 Petrolisthes unilobatus Henderson, 1888 114 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Polyonyx obesulus Miers, 1884 {33} = Porcellana stimpsoni A. Milne-Edwards, 1880 = Porcellana obesula White, 1847 (nomen {37} nudum) = ? Porcellana robertsoni Henderson, 1888 = Porcellana (Polyonyx) tuberculosa De Man, Porcellana sigsbeiana A. Milne-Edwards, 1880 1888 Porcellanella White, 1852 = Polyonyx parvidens Nobili, 1905 = Porcellanella White, 1852 (type species = Polyonyx paucidens Nobili 1906 (misspelling of Porcellanella triloba White, 1852, by monotypy; parvidens) gender feminine) Polyonyx pedalis Nobili, 1905 Porcellanella haigae Sankarankutty, 1963 Polyonyx plumatus Yang & Xu, 1994 Porcellanella triloba White, 1852 Polyonyx quadratus Chace, 1956 = Porcellanella picta Stimpson, 1858 Polyonyx quadriungulatus Glassell, 1935 Pseudoporcellanella Sankarankutty, 1961 Polyonyx senegalensis Chace, 1956 = Pseudoporcellanella Sankarankutty 1961 Polyonyx sinensis Stimpson, 1858 (type species Pseudoporcellanella manoliensis Polyonyx spina Osawa, 2007 Sankarankutty, 1961, by monotypy; gender Polyonyx splendidus Sankolli, 1963 {34} feminine) Polyonyx thai Werding, 2001 Pseudoporcellanella manoliensis Sankarankutty, 1961 Polyonyx transversus (Haswell, 1882) [Porcellana] Raphidopus Stimpson, 1858 Polyonyx triunguiculatus Zehntner, 1894 = Raphidopus Stimpson, 1858 (type species = Polyonyx acutifrons De Man, 1896 Raphidopus ciliatus Stimpson, 1858, by monotypy; Polyonyx tulearis Werding, 2001 gender masculine) Polyonyx utinomii Miyake, 1943 Raphidopus ciliatus Stimpson, 1858 Polyonyx vermicola Ng & Sasekumar, 1993 Raphidopus indicus Henderson, 1893 Porcellana Lamarck, 1801 Raphidopus johnsoni Ng & Nakasone, 1994 = Porcellana Lamarck, 1801 (type species Cancer Ulloaia Glassell, 1938 platy-cheles Pennant, 1777, by monotypy; gender = Ulloaia Glassell, 1938 (type species Ulloaia feminine) perpusillia Glassell, 1938, by original designation; = Platycheles Billberg, 1820 (type species Cancer gender feminine) platy-cheles Pennant, 1777, by tautonomy; gender Ulloaia perpusillia Glassell, 1938 masculine) = Enostea Gistel, 1848 (unnecessary replacement name for Porcellana; type species Cancer Incerta sedis platycheles Pennant, 1777) Porcellana africana Chace, 1956 {35} Petrolisthes costai Rodriguez da Costa, 1968 {38} = Porcellana platycheles africana Chace, 1956 Pisidia melloleitaoi Rodriguez da Costa, 1968 {38} Porcellana cancrisocialis Glassell, 1936 Porcellana mattosi Osorio, 1887 {39} Porcellana caparti Chace, 1956 {36} Porcellana corbicola Haig, 1960 Porcellana curvifrons Yang & Sun, 1990 NOTES Porcellana elegans Chace, 1956 {36} Porcellana foresti Chace, 1956 {36} {1} Aliaporcellana suluensis has variations in the Porcellana habei Miyake, 1961 morphology of the carapace, host animals, and Porcellana hancocki Glassell, 1938 coloration in life, which can be referred to species = Porcellana hancocki Glassell, 1937 (nomen specifi c characters. Thus, the two species currently nudum) considered synonyms of A. suluensis may prove to Porcellana lillyae Lemaitre & Campos, 2000 be valid taxa. Porcellana paguriconviva Glassell, 1936 Porcellana persica Haig, 1966 {2} The status of Eulenaios has not been satisfactorily Porcellana platycheles (Pennant, 1777) [Cancer] {35} resolved. Ng & Nakasone’s (1993) establishment of = Porcellana paivacarvalhoi Rodriguez da Costa, Eulenaios for Polyonyx cometes was not recognized 1968 {35} by Werding (2001). However, Osawa (2007a) has Porcellana pulchra Stimpson, 1858 suggested that the validity of Eulenaios does have Porcellana sayana (Leach, 1820) [Pisidia] support. = Porcellana sayii Gray, 1831 (misspelling of sayana) {3} Based on the author’s diagnosis, Haig (1978) = Porcellana ocellata Gibbes, 1850 suggested that Czerniavsky’s (1884) Porcellanides = Porcellana sagrai Guérin-Méneville, 1855 might be a senior synonym of Lissoporcellana because = Percellana sagrai Von Martens, 1872 (misspelling the type species, P. kriczagini, from Singapore was of Porcellana) likely to be a member of Lissoporcellana. 115 Osawa & McLaughlin: Checklist of world Porcellanidae {4} Madarateuchus was established by Harvey (1999) {11} The generic status of Neopisosoma has been as a monotypic genus for Clastotechus vanderhorsti. questioned since its establishment as was discussed Although the molecular analyses by Rodriquez et al. by Gore (1970), Werding (1986), Werding & Müller (2006) did not support the validity of the genus, it is (1990), Werding et al. (2001), and Rodríguez et al. presently considered a valid taxon. (2004, 2005). Although Rodríguez et al. (2006) found that Neopisosoma was supported as a genus distinct {5} Porcellanopsis was proposed by Rathbun (1910) from Pachycheles based on molecular analysis, a as a replacement name for Porcellanides Nobili, worldwide review of both genera will be required preoccupied by Porcellanides Czerniavsky. Although to clarify their relationships. Haig (1960) considered Porcellanopsis a synonym of Megalobrachium, Rodriguez et al. (2006) suggested, {12} Since the generic gender is neuter, the name N. based on molecular evidence, that Rathbun’s taxon phuketensis is emended to N. phuketense. might actually be a distinct taxon. {13} Osawa (1998a) treated N. securiger as a junior {6} Haig (1957b) noted that the specifi c name festae synonym of N. indicum. Additional specimens from must be spelled festai in accordance with current Taiwan and the Ogasawara and Ryukyu Islands, nomenculatorical practice. And while it is true the western Pacific, have shown that the differences the initial generic assignment, Porcellanides, is Osawa considered size related are actually species masculine (ICZN Code, 1999), the name festae is specifi c characters. Melin’s (1939) species has been a patronym, and as such does not change endings reinstated (Osawa & Chan, 2010). with generic reassignments. Although the feminine “e” rather than masculine “i” was used by the author {14} Although Pachycheles holosericus Schmitt, 1918 was without explanation, that is his original spelling and cited by name in Nininger (1918) and presumably is to be maintained. Nobili (1901) did not specify a accompanied by a fi gure, there was no description dedication to Enrico Festa, thus it is possible that the and according to Haig (1960) the text fi gure was dedication was meant for his wife. unrecognizable to species. Schmitt (1921) described this taxon as a new species. {7} Although Bulletin of Marine Science, 23(3) was supposed to have been published in September of {15} Buckup & Bond-Buckup (1999) included Pachycheles 1973, the publication was delayed and that issue did haigae as a valid species in the Brazilian fauna, not appear in print until early 1974. perhaps unaware that Harvey & de Santo (1996) had placed this taxon in synonymy with Pachycheles {8} Ng et al. (2008) cited the footnote by Evenhuis (2003: laevidactylus. 17) explaining the problem with correct citations of the author Guérin-Méneville. Guérin added the {16} Parapetrolisthes was established for Petrolisthes honorifi c suffi x “Méneville” to his surname in 1836 tortugensis on the basis of the shape of the fi ngers. after his Encyclopédie Méthodique was completed. However, the molecular analyses by Hiller et al. (2006) Ng et al. (2008) choose to cite authorship of all and Rodriguez et al. (2006) suggested that this species taxa published by this author prior to 1836 simply should be grouped with the Petrolisthes galathinus as Guérin and Guérin-Méneville from 1836 onward. complex. The validity of the genus and its postulated We have followed this same protocol. relationship with the Galatheidae were not supported by these molecular studies. Nonetheless, it is retained {9} Based on their molecular analyses, Rodriguez et al. as a valid genus until such time as revisionary studies (2006) suggested that Megalobrachium soriatum determine its proper affi liation. contained a cryptic species. {17} The developmental and morphological characters {10} Haig (1965, 1979) considered Neopetrolisthes ohshimai of larvae and adults of Petrocheles suggest a close a junior synonym of N. maculatus and pointed out relationship between this genus and some galatheids that the Indian and Pacifi c Ocean populations of N. (Hale, 1927; Wear, 1965; pers. obs. of one of the maculatus had different patterns of spots. However, present authors, MO). Molecular analyses of Hiller Osawa & Fujita (2001) noted that both spot-patterns et al. (2006) and Ahyong et al. (2009) suggested that occurred in the Ryukyu Islands, western Pacifi c. Nor Petrocheles should be excluded from the porcellanid were the two species distinguishable from one other clade or that the Galatheidae might be paraphyletic by morphological characters, thus supporting the on the basis of an internally nested Porcellanidae. status of N. ohshimai as a junior synonym. It was also found that N. maculatus had various spot sizes {18} The genus Petrolisthes is the most species-rich genus and patterns of reddish purple, red, or brown color on in the Porcellanidae. Several informal species groups the carapace and pereopods. Further ecological and have been distinguished in this genus based on the genetic studies are needed before these variations are adult and larval morphology (e.g., Haig, 1960; Osawa, fully understood. 1995; Fujita et al., 2002). Molecular phylogenies 116 THE RAFFLES BULLETIN OF ZOOLOGY 2010 by Stillman & Reeb (2001) and Rodriguez et al. {26} Certain characters of P. mesodactylon suggest a (2006) show Petrolisthes as currently recognized is possible relationship between this species and paraphyletic on the basis of nested positions of species Novorostrum securiger. It is possible that further of Allopetrolisthes, Clastotoechus, Liopetrolisthes, study will show that P. mesodactylon actually should Megalobrachium, and Parapetrolisthes. be assigned to Novorostrum. {19} Haig (1960) emended the spelling from cabrilloa {27} Two forms of both Petrolithes militaris and P. to cabrilloi to conform with the ICZN rule for scabriculus have been found in the western Pacifi c, the formation of specifi c names based on modern each differing in coloration and habitats. Further surnames. investigation is needed to determine if one or both represent more than one taxon. {20} Miyake & Nakasone (1966) determined that the specimens reported by Melin (1939) as P. carinipes {28} Although Petrolisthes rathbunae was cited by name did not agree with Heller’s (1861) taxon and proposed in Hilton (1916) and presumably accompanied by the name Petrolisthes melini for those specimens. a fi gure, there was no description and according to Similarly, Lewinsohn (1974) found differences Haig (1960), the text fi gure was unrecognizable to between Melin’s (1939) material and the type of P. species. Schmitt (1921) described this taxon as a new carinipes, but did not describe Melin’s material as species. a new species. Recent studies suggest that P. melini as defi ned by Miyake & Nakasone (1966) agrees {29} Melin (1939) proposed the replacement name more closely with P. extremus Kropp & Haig, 1994 Petrolisthes teres for Petrolisthes inermis Haswell, than with P. carinipes. If further study shows that P. 1882, preoccupied by Porcellana inermis Heller, 1862 melini and P. extremus are conspecifi c, P. melini has (= Petrolisthes hastatus Stimpson, 1858). Although priority. the types of Petrolisthes teres should have been the material described as P. inermis by Haswell (1882), {21} The correct date of publication of Randall’s paper Melin (1939) used his non-type material from the in the Journal of the Academy of Natural Sciences, Bonin (Ogasawara) Islands to describe P. teres. Re- Philadelphia is 1840 (Melville & Smith, 1987) examination of some Melin’s (1939) specimens has shown that they are actually P. japonicus (De Haan, {22} Whereas Ng et al. (2008) elected to omit the honorifi c 1849). Judging from the photographs of the syntypes “de” for all French authors, we have chosen to retain of P. teres (= P. inermis Haswell, 1882), there is a it for those authors commonly cited in Anomuran small spine on the anterior margin of the third segment literature; e.g., de Saussure, de Saint Laurent. of the antennal peduncle. Such spine is absent in P. japonicus. Among other morphologically allied {23} The taxonomic relationship of Petrolisthes lamarckii congeners, the spine is present in P. elongatus, P. with its presumed synonyms is unsatisfactory. kranjiensis, P. limicola, as well as those ‘P. inermis’ Examination by one of the current authors (MO) has reported by Miyake (1943) and ‘P. teres’ reported by shown that specimens from various western Pacifi c Johnson (1970). The exact identity of P. teres has yet localities have exhibited considerable variations in to be determined. certain signifi cant characters, in addition to variations in living color. Further study will undoubtedly show {30} Haig (1960) discussed the confusion that had existed that several distinct taxa have been confounded under among the names applied to Petrolisthes tuberculosus. the name P. lamarckii. She pointed out that although Ortmann (1897) argued that Porcellana affi nis Gray, 1831 was undefi ned, thus {24} Hendrickx & Harvey (1999) indicated that the not a homonym of P. affi nis Guérin, 1835, Gray’s subspecies austrinus and the nominal subspecies lewisi description albeit brief, was suffi cient to validate were not distinct and considered P. lewisi austinus a it. Consequently, Guérin’s (1835) P. affi nis was not synonym. available. {25} Gore (1983) placed P. cessacii in synonymy with {31} The validity of Pisidia bluteli, P. longicornis and P. P. marginatus on the basis of adult morphological longimana as distinct species has been the subject characters, but cautioned that whether or not the of debate. Holthuis (1961) and Zariquiey Álvarez eastern and western Atlantic populations of the (1968) clearly distinguished the three species. In species were reproductively isolated remained to contrast, García Raso (1987) found what he believed be determined. At that time Gore believed that the to be a morphological cline that existed between P. answer might be found in a comparison of the larval longicornis and P. longimana, and d’Udekem d’Acoz morphology of the different populations. With the (1999) discussed his belief that P. bluteli was not advances being made in molecular determinations, distinct from P. longicornis. However, Koukouras the answer well may be found in examinations of et al. (2002) found constant characters that separated the populations’ DNA. the three. 117 Osawa & McLaughlin: Checklist of world Porcellanidae {32} Pisidia dispar and P. variabilis differ considerably LITERATURE CITED from other congeners in such characters as carapace convexity, rostral structure and orbital angle. The Ahyong, S. T. & D. O’Meally, 2004. Phylogeny of the Decapoda possibility exists that these species should be placed Reptantia: Resolution using three molecular loci and morphology. Raffl es Bulletin of Zoology 52: 673–693. in a separate genus. Ahyong, S. T., K. E. Schnabel, & E.W. Maas, 2009. Anomuran {33} The taxonomy of Polyonyx biunguiculatus and P. phylogeny: new insights from molecular data. In: Martin, J. W., K. A. Crandall, & D. L. Felder (Eds.). Decapod Crustacean obesulus is still problematic (see Osawa, 2007b) and Phylogenetics. Crustacean Issues, 18. CRC Press, Boca Raton, a review of specimens hitherto reported as the two FL, New York, London. Pp. 399–414. species as well as their synonyms is needed. Audouin, V., 1826. Explication sommaire des planches de Crustacés de l’Egypte et de la Syrie, publiées par Jules-Cesar Savigny, {34} Werding (2001) suggested that P. hendersoni and P. member de l’Institu; Offrant un exposé des caracters naturels splendidus represented a distinct genus, although no des generes avec la distinction des especes. Desription de formal action was taken. l’Egypte, ou recueil des observation et des recherches que ont ete faites en Égypte pendant l’Expedition de l’armée francaise. {35} Chace’s (1956) subspecies of Porcellana platycheles, Histoire Naturelle, 1: 77–98. Paris. P. p. africanus, was considered a distinct taxon Baker, W. H., 1905. Notes on South Australian decapod Crustacea. by d’Udekem d’Acoz (1999) and elevated to full Part III. Transactions and Proceedings of the Royal Society of specifi c rank. No additional subspecies have been South Australia, 29: 252–269. described. Veloso & Melo (1993) also considered Balss, H., 1913. Ostasiatische Decapoden. I. Die Galatheiden P. paivacarvalhoi to be a junior synonym of P. und Paguriden. In: Dofl ein, F., Beiträge zur Naturgeschichte platycheles. Ostasiens. Abhandlungen der Mathematisch-Physische Klasse der Königlich Bayerischen Akademie der Wissenschaften, München, Supplement 2(9): 1–85. {36} In her revision of Porcellana, Haig (1978) established four new genera for nine species formerly assigned to Balss, H., 1914. Decapode Crustaceen von den Giinea-Inseln, the genus. Nonetheless, the retention of certain species Süd-Kamerun und dern Congogebiet. Ergebnisse der Zweiten Deutschen Zentral-Afrika-Expedition 1910-1911 unter Führung in Porcellana is questionable. Haig mentioned that P. Adolph Friedrichs, Herzog zu Mecklenburg, 1 (Zoologie): caparti and P. elegans should perhaps be reassigned 97–108. to Enosteoides and Capilliporcellana, respectively. Barnard, K. H., 1955. Additions to the fauna-list of South African She believed that P. foresti appeared to be related Crustacea and Pycnogonida. Annuals of the South African to Ancyclocheles. Until the type materials of all Museum, 43: 1–107. three taxa are reexamined, the questions of generic Benedict, J. E., 1901. The anomuran collections made by the Fish placement remain ambiguous. However, based on the Hawk Expedition to Porto Rico. Bulletin of the U. S. Fish structures of the carapaces and chelipeds, at least P. Commission, 20: 129–148. caparti and P. foresti appear clearly different from Billberg, G. J., 1820. Enumeratio insectorum in Museo Gust. Typis species of Enosteoides and Ancylocheles. Gadelianis, Holmiae. 138 pp. Boas, J. E. V., 1880. Studier over decapodernes Slægtskabsforhold. {37} Porcellana stimpsoni was briefl y described by A. Kongelige Danske Videnskabernes Selskabs Skrifter, Milne-Edwards (1880). The presumably differentiating naturvidenskabelig og mathema tisk Afdeling, (5)6: 25–210. characters of A. Milne-Edwards’ taxon were found Boone, L., 1930. Scientifi c results of the cruises of the yachts to be the result of abnormal development of the left “Eagle” and “Ara”, 1921–1928, William K. Vanderbilt, cheliped and frontal region in the type specimen. commanding. Crustacea: Ano mura, Macrura, Schizopoda, Consequently, P. stimpsoni cannot be distinguished Isopoda, Amphipoda, Mysidacea, Cirripedia, and Copepoda. from P. sayana, and is considered a junior synonym Bulletin of the Vanderbilt Marine Museum, 3: 1–221. of the latter species. (B. Werding, in litt.; see Lemaitre Boone, L., 1932. The littoral crustacean fauna of the Galapagos & Campos, 2000). Islands. Part II. Anomura. Zoologica, New York, 14: 1–62. Borradaile, L. A., 1898. On some crustaceans from the South {38} According to Veloso & de Melo (1993) and Rodriguez Pacifi c. Part II. Macrura Anomala. Proceedings of the Zoological et al. (2005), Petrolisthes costai Rodriguez da Costa, Society of London, 1898: 457–468. 1968 and Pisidia melloleitaoi Rodriguez da Costa, Borradaile, L. A., 1900. On the Stomatopoda and Macrura brought 1968 could not be confi rmed as valid species. by Dr Willey from the south seas. In: Willey, A., Zoological results based on material from New Britain, New Guinea, {39} According to Chace (1956), until suffi cient material Loyalty Islands and elsewhere, collected during the years is available, the true generic status of Porcellana 1895, 1896 and 1897. Part IV. Cambridge, University Press. mattosi Osorio, 1887, cannot be determined. Pp. 395–428. Bosc, L. A. G., 1802. Histoire naturelle des Crustacés, contenant leur description et leurs moeurs, avec fi gures dessinées d’après nature. 2: 1–296. Paris, Deterville. Boschi, E. E., 1963. Sobre dos especies de Pachycheles de la Argentina (Crustacea, Anomura). Neotropica, 9(28): 31–37. 118